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''Riccia'' is a genus of
liverwort The Marchantiophyta () are a division of non-vascular land plants commonly referred to as hepatics or liverworts. Like mosses and hornworts, they have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of ...
s in the order
Marchantiales Marchantiales is an order of thallose liverworts (also known as "complex thalloid liverworts") that includes species like ''Marchantia polymorpha'', a widespread plant often found beside rivers, and ''Lunularia cruciata'', a common and often trou ...
. These plants are small and
thalloid Thallus (plural: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. Many of these organisms were ...
, that is not differentiated into root, stem and leaf. Depending on species, the thallus may be strap-shaped and about 0.5 to 4 mm wide with dichotomous branches or may form rosettes or hemirosettes up to 3 cm in diameter, that may be gregarious and form intricate mats. The thallus is
dorsiventral A dorsiventral (Lat. ''dorsum'', "the back", ''venter'', "the belly") organ is one that has two surfaces differing from each other in appearance and structure, as an ordinary leaf. This term has also been used as a synonym for dorsoventral organs, ...
ly differentiated. Its upper (dorsal) surface is green and
chlorophyll Chlorophyll (also chlorophyl) is any of several related green pigments found in cyanobacteria and in the chloroplasts of algae and plants. Its name is derived from the Greek words , ("pale green") and , ("leaf"). Chlorophyll allow plants to ...
-bearing, with a mid-dorsal longitudinal sulcus (furrow or groove). Air pores occasionally break through the dorsal surface, giving the thallus a dimpled appearance. In exceptional members such as ''Riccia caroliniana'' of Northern Australia and ''Riccia sahyadrica'' of Western Ghats, the photosynthetic region is confined to the lower half of the thallus. The lower (ventral) surface has a mid-ventral ridge bearing multicellular scales that originate as a single row but normally separate into two rows as the thallus widens. The scales are multicellular and
hyaline A hyaline substance is one with a glassy appearance. The word is derived from el, ὑάλινος, translit=hyálinos, lit=transparent, and el, ὕαλος, translit=hýalos, lit=crystal, glass, label=none. Histopathology Hyaline cartilage is ...
(glassy) in appearance, or violet due to the pigment
anthocyanin Anthocyanins (), also called anthocyans, are water-soluble vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart gave the name Anthokyan to a chemical com ...
. Rhizoids are nearly lacking in aquatic forms, but there are usually numerous unicellular rhizoids of two types on the ventral surface. One type is called smooth and the other type is the pegged or tuberculated rhizoids; these help in anchorage and absorption. The inner surface of the smooth rhizoids is smooth while that of the tuberculate rhizoid will have internal
cell wall A cell wall is a structural layer surrounding some types of cells, just outside the cell membrane. It can be tough, flexible, and sometimes rigid. It provides the cell with both structural support and protection, and also acts as a filtering mec ...
projections. Plants are usually monoicous, and sexual reproduction is by antheridia and
archegonia An archegonium (pl: archegonia), from the ancient Greek ''ἀρχή'' ("beginning") and ''γόνος'' ("offspring"), is a multicellular structure or organ of the gametophyte phase of certain plants, producing and containing the ovum or female ga ...
. Asexual reproduction occurs by
spores In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, ...
, by fragmentation of the rosettes, and by formation of apical
tubers Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing s ...
. Spores are large (45 to 200 µ) and formed in tetrads. One of the more than 100 species in this genus is the "slender riccia" ('' Riccia fluitans''), which grows on damp soil or, less commonly, floating in ponds,MacVicar, Symers M. 1971. The student's handbook of British hepatics, p. 12-13. Hitchen Whelden & Wesley, Ltd.; Authorized reprint of the 1926 edition published by Sumfield & Day, Ltd. and is sometimes used in
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
s. The
sporophyte A sporophyte () is the diploid multicellular stage in the life cycle of a plant or alga which produces asexual spores. This stage alternates with a multicellular haploid gametophyte phase. Life cycle The sporophyte develops from the zygote ...
of Riccia is the simplest amongst
bryophyte The Bryophyta s.l. are a proposed taxonomic division containing three groups of non-vascular land plants (embryophytes): the liverworts, hornworts and mosses. Bryophyta s.s. consists of the mosses only. They are characteristically limited in s ...
s. It consist of only a capsule, missing both foot and seta, and does not perform photosynthesis.Text Book Of Botany Diversity Of Microbes And Cryptogams
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References

{{Taxonbar, from=Q1856753 Ricciaceae Marchantiales genera