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''Rhododendron maximum'' — its common names include great laurel, great rhododendron, rosebay rhododendron, American rhododendron and big rhododendron — is a species of ''
Rhododendron ''Rhododendron'' (; from Ancient Greek ''rhódon'' "rose" and ''déndron'' "tree") is a very large genus of about 1,024 species of woody plants in the heath family (Ericaceae). They can be either evergreen or deciduous. Most species are nati ...
'' native to the
Appalachians The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. They ...
of eastern
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, from
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north to coastal
Nova Scotia Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. Nova Scotia is Latin for "New Scotland". Most of the population are native Eng ...
.


Description

''Rhododendron maximum'' is an
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
growing to , rarely , tall. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are long and broad. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s are diameter, white, pink or pale purple, often with small greenish-yellow spots. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
is a dry capsule long, containing numerous small
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s. The leaves can be poisonous. Leaves are sclerophyllous, simple, alternate, and oblong (10 to 30 cm long, 5 to 8 cm wide). It retains its waxy, deep-green leaves for up to 8 years, but once shed are slow to decompose. It produces large, showy, white to purple flowers each June and July.


Range

Rosebay rhododendron is the most frequently occurring and dominant species of ''Rhododendron'' in the southern Appalachian region, and occurs occasionally on mesic hill-slopes throughout the upper Piedmont Crescent of the Southeastern United States.


Ecology

Approximately 12,000 square miles in the southern Appalachians are occupied by this species where it dominates the understory. This species has historically been confined to riparian areas and other mesic sites but takes advantage of disturbed areas where it is present to advance onto sub-mesic sites. It prefers deep well-drained acid soils high in organic matter where it produces a thick, peat-like humus. It prefers low to medium light conditions for optimum carbon gain, and has a tremendous capacity for avoiding cavitation during freeze-thaw cycles. Where extensive overstory mortality has eliminated most of the overstory, this species forms a thick and continuous subcanopy known locally as ‘laurel slicks’ or ‘laurel hells’. Rosebay rhododendron is an important structural and functional component of southern Appalachian forest ecosystems. What isn't clear is whether or not we are in a period of advancement or retreat for this species. For example, on poorly drained sites on ridge or upper slope positions, large areas of rosebay rhododendron, particularly at the high elevations, have recently died out presumably due to the ''
Phytophthora ''Phytophthora'' (from Greek (''phytón''), "plant" and (), "destruction"; "the plant-destroyer") is a genus of plant-damaging oomycetes (water molds), whose member species are capable of causing enormous economic losses on crops worldwide, a ...
'' fungus, or due to recent prolonged periods of below-average precipitation. Yet, rosebay rhododendron now occupies sites that historically were free of evergreen understory. There are still important questions to be answered regarding this species to completely understand its role in forest understories.


Reproduction

Rosebay rhododendron is clonal. It is capable, however, of reproducing both vegetatively and sexually. It reproduces vegetatively through a process called ‘
layering Layering has evolved as a common means of vegetative propagation of numerous species in natural environments. Layering is also utilized by horticulturists to propagate desirable plants. Natural layering typically occurs when a branch touches ...
’ where it produces roots from above ground woody parts when in contact with the forest floor. The fruit is produced from showy flowers from March to August. The fruit is an oblong capsule that ripens in the fall, and splits along the sides soon after ripening to release large numbers of minute seed (approx. 400 per capsule). Microsite requirements for seed germination are relatively specific (e.g., high in organic matter such as rotting logs); hence, the majority of reproduction is vegetative resulting in a clonal distribution.


Growth and management

Seeds from rosebay rhododendron are minute and it is estimated that approximately 11 million are contained in 1 kg. Commercial seed production is generally from cultivated hybrids. Seeds from wild sources are not commonly sold commercially. Rosebay rhododendron is a slow-growing shrub and has a very high sprout potential. If mechanical removal is attempted in the case of forest management, extremely high densities are attained by this species in a matter of a few years. Prescribed fire has also been used to control this species but with limited success.


Benefits

Rosebay rhododendron is a striking and aesthetically pleasing feature of mesic southern Appalachian forests. It is one of the largest and hardiest rhododendrons grown commercially. Several cultivars with white to purple flowers have been selected for the horticultural trade. Where it occurs naturally, it produces a showy, white, pink, or light purple flower primarily in June, but occurs from March into August. Rosebay rhododendron maintains deep-green foliage year round. This species affords protection to steep watersheds and shelter for wildlife. The wood is very hard and is occasionally used for specialty wood products.


Detrimental effects

For all its prized qualities as a naturally occurring component of the landscape or as plantings in residential and commercial landscaping, rosebay rhododendron can have an inhibitory effect on regeneration of other plant species. There is some evidence to suggest that due to fire suppression and the absence of other cultural activities (i.e., mountain-land grazing), this species has advanced beyond the mesic forest sites into sub-mesic understories. The significance of this movement onto previously unoccupied sites centers around the impacts of rosebay rhododendron on plant succession and resource availability. Rosebay rhododendron is associated with reduced woody and herbaceous seedling abundance throughout its range, and hence poses a serious impediment to the production of wood products. The mechanism(s) by which rosebay rhododendron reduces seedling survival has been the subject of much debate. Possible sources of inhibition include
allelopathy Allelopathy is a biological phenomenon by which an organism produces one or more biochemicals that influence the germination, growth, survival, and reproduction of other organisms. These biochemicals are known as allelochemicals and can have be ...
, competition for resources including light, physical and chemical attributes of the forest floor and soil, and interactions between some or all sources.


Alternate common names

''R. maximum'' has also been called: *Great rhododendron *Late rhododendron *Summer rhododendron *Great laurel *Bigleaf laurel *Deertongue laurel *Rose tree *Rose bay *Bayis


Symbolism

''Rhododendron maximum'' is the state flower of the U.S. state of
West Virginia West Virginia is a state in the Appalachian, Mid-Atlantic and Southeastern regions of the United States.The Census Bureau and the Association of American Geographers classify West Virginia as part of the Southern United States while the B ...
.


See also

*
Catawbiense hybrid A catawbiense hybrid is a hybrid of the plant genus '' Rhododendron'', derived from crosses of American species '' Rhododendron catawbiense'' and ''Rhododendron maximum'' with European and Asian species '' R. arboreum'', '' R. caucasicum'' and ...
– R. catawbiense & R. maximum hybrids *
Central and southern Appalachian montane oak forest The Central and southern Appalachian montane oak forest is a forest system found in the Appalachian Mountains of West Virginia, Virginia, Kentucky, Tennessee, North Carolina, South Carolina, and Georgia. These forests occur on exposed sites such a ...
*
List of Rhododendron species This List of ''Rhododendron'' species includes species of the genus ''Rhododendron'', which is in the plant family Ericaceae. Depending on the source, there are anywhere from 800 to over 1,100 wild species. The vast majority of ''Rhododendron'' sp ...


References

*


External links


USDA Plants Profile of ''Rhododendron maximum'' (great rhododendron)Interactive Distribution Map for ''Rhododendron maximum''Plants for a Future: ''Rhododendron maximum''
{{Authority control
maximum In mathematical analysis, the maxima and minima (the respective plurals of maximum and minimum) of a function, known collectively as extrema (the plural of extremum), are the largest and smallest value of the function, either within a given r ...
Flora of the Appalachian Mountains Flora of the Northeastern United States Flora of the Southeastern United States Flora of Nova Scotia Plants described in 1753 Taxa named by Carl Linnaeus Symbols of West Virginia