Remingtonocetidae is a diverse

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

of early aquatic

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

s of the order

Cetacea Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel them ...

. The family is named after paleocetologist

Remington Kellogg Arthur Remington Kellogg (5 October 1892 – 8 May 1969) was an American naturalist and a director of the United States National Museum. His work focused on marine mammals. Early life and career Kellogg was born in Davenport, Iowa, and quick ...

Description

Remingtonocetids have long and narrow skulls with the external

nare A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turb ...

openings located on the front of the skull. Their

frontal shield A frontal shield, also known as a facial shield or frontal plate, is a feature of the anatomy of several bird species. Located just above the upper mandible, and protruding along the forehead, it is composed of two main parts: a hard, proteinaceo ...

s are narrow and their orbits small. Their mouth has a convex

palate The palate () is the roof of the mouth in humans and other mammals. It separates the oral cavity from the nasal cavity. A similar structure is found in crocodilians, but in most other tetrapods, the oral and nasal cavities are not truly sepa ...

and an incompletely fused

mandibular symphysis In human anatomy, the facial skeleton of the skull the external surface of the mandible is marked in the median line by a faint ridge, indicating the mandibular symphysis (Latin: ''symphysis menti'') or line of junction where the two lateral halves ...

. The dental formula is . The anterior teeth are flattened mediolaterally, making them appear shark-like. In the

postcrania Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated sk ...

l skeleton, the

cervical vertebrae In tetrapods, cervical vertebrae (singular: vertebra) are the vertebrae of the neck, immediately below the skull. Truncal vertebrae (divided into thoracic and lumbar vertebrae in mammals) lie caudal (toward the tail) of cervical vertebrae. In ...

are relatively long and the

sacrum The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part ...

is composed of four vertebrae of which at least three are fused. The

acetabular notch The acetabular notch is a deep notch in the acetabulum of the hip bone. The acetabular notch is continuous with a circular non-articular depression, the acetabular fossa, at the bottom of the cavity: this depression is perforated by numerous aper ...

is narrow or closed and on the

femoral head The femoral head (femur head or head of the femur) is the highest part of the thigh bone (femur). It is supported by the femoral neck. Structure The head is globular and forms rather more than a hemisphere, is directed upward, medialward, and a l ...

the

fovea Fovea () (Latin for "pit"; plural foveae ) is a term in anatomy. It refers to a pit or depression in a structure. Human anatomy *Fovea centralis of the retina * Fovea buccalis or Dimple * Fovea of the femoral head * Trochlear fovea of the fr ...

is absent. Kutchicetus minimus

Cranial fossils are common but dental remains are rare. The postcrania morphology is based entirely on a single specimen of ''

Kutchicetus ''Kutchicetus'' is an extinct genus of early whale of the family Remingtonocetidae that lived during Early-Middle Eocene (Lutetian and Ypresian) in what is now the coastal border of Pakistan and India (, paleocoordinates ). It is closely relate ...

'' which was small and had a long and muscular back and tail. Perhaps remintonocetids swam like the South American

giant otter The giant otter or giant river otter (''Pteronura brasiliensis'') is a South American carnivorous mammal. It is the longest member of the weasel family, Mustelidae, a globally successful group of predators, reaching up to . Atypical of musteli ...

which swims with its long flat tail. With long and low bodies, relatively short limbs, their elongated

rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ships * Ros ...

, remingtonocetids looked like mammalian crocodiles, more so than ''

Ambulocetus ''Ambulocetus'' (Latin ''ambulare'' "to walk" + ''cetus'' "whale") is a genus of early amphibious cetacean from the Kuldana Formation in Pakistan, roughly 48 or 47 million years ago during the Early Eocene (Lutetian). It contains one species, ' ...

''. They could both walk on land and swim in the water and most likely lived in a near-shore habitat. At least one genus, ''

Dalanistes ''Dalanistes'' is an extinct genus of remingtonocetid early whale known from the late early Eocene (Lutetian, ) of Kutch, India and Punjab and Balochistan, Pakistan. ''Dalanistes'' is closely related to '' Remingtonocetus'' (the type genus ...

'', had a marine diet. Remingtonocetids are often found in association with catfish and crocodilians, as well as

protocetid Protocetidae, the protocetids, form a diverse and heterogeneous group of extinct cetaceans known from Asia, Europe, Africa, South America, and North America. Description There were many genera, and some of these are very well known (e.g., ''R ...

whales and

sirenia The Sirenia (), commonly referred to as sea-cows or sirenians, are an order of fully aquatic, herbivorous mammals that inhabit swamps, rivers, estuaries, marine wetlands, and coastal marine waters. The Sirenia currently comprise two distinct f ...

ns. They were probably independent of freshwater.

Distribution

Remingtonocetidae was long considered endemic to the northern coastline of the ancient

Tethys Ocean The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...

(in present day Pakistan and India) during the

Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...

, but the discovery of ''

Rayanistes Rayanistes is a genus of remingtonocetid whale from the Middle Eocene deposits in Egypt. Significance and biology ''Rayanistes'' was capable of extensive power strokes during pelvic paddling, based on the robust hindlimb and innominate. The mo ...

'' in Egypt indicates that remingtonocetids had a broader distribution than previously thought.

Taxonomy

Remingtonocetidae was established by . It was considered

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...

by . It was assigned to

Odontoceti The toothed whales (also called odontocetes, systematic name Odontoceti) are a parvorder of cetaceans that includes dolphins, porpoises, and all other whales possessing teeth, such as the beaked whales and sperm whales. Seventy-three species of ...

by ; to Remingtonocetoidea by and ; to

Archaeoceti Archaeoceti ("ancient whales"), or Zeuglodontes in older literature, is a paraphyletic group of primitive cetaceans that lived from the Early Eocene to the late Oligocene (). Representing the earliest cetacean radiation, they include the initia ...

by ; to Archaeoceti by , , , , , , , and and to Cetacea by , and .. Retrieved February 2013 The name of the family was derived from the

type genus In biological taxonomy, the type genus is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearing type of a nominal f ...

''Remingtonocetus'', which was named after paleocetologist

. In 2009, paleontologists Thewissen & Bajpai proposed the subfamily

Andrewsiphiinae The Andrewsiphiinae is an extinct subfamily of early whales of the family Remingtonocetidae Remingtonocetidae is a diverse family of early aquatic mammals of the order Cetacea. The family is named after paleocetologist Remington Kellogg. ...

for the genera ''

Andrewsiphius ''Andrewsiphius'' is an extinct remingtonocetid early whale known from the Eocene (Lutetian, ) of Gujarat and Kutch, India and Balochistan, Pakistan. Discovery and naming The first specimen was collected by who described it as mandibular f ...

'' and ''

''.

Genera

*''

Remingtonocetus ''Remingtonocetus'' is an extinct genus of early cetacean freshwater aquatic mammals of the family Remingtonocetidae endemic to the coastline of the ancient Tethys Ocean during the Eocene. It was named after naturalist Remington Kellogg. History ...

'' (type) *''

'' *''

Attockicetus ''Attockicetus'' is an extinct genus of remingtonocetid early whale known from the Middle Eocene (Lutetian) Kuldana Formation in the Kala Chitta Hills, in the Attock District of Punjab, Pakistan. ''Attockicetus'' is described based on frag ...

'' , the oldest genus *''

'' *''

'' Bebej, Zalmout, El-Aziz, Antar, and Gingerich, 2016

Notes

References