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The red squirrel (''Sciurus vulgaris'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of
tree squirrel Tree squirrels are the members of the squirrel family (Sciuridae) commonly just referred to as "squirrels." They include more than 100 arboreal species native to all continents except Antarctica and Oceania. They do not form a single natural, o ...
in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''
Sciurus The genus ''Sciurus'' contains most of the common, bushy-tailed squirrels in North America, Europe, temperate Asia, Central America and South America. Species The number of species in the genus is subject to change. In 2005, Thorington & H ...
'' common throughout Europe and Asia. The red squirrel is an
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose nu ...
, primarily
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
. In Great Britain, Ireland, and in Italy numbers have decreased drastically in recent years. This decline is associated with the introduction by humans of the eastern grey squirrel (''Sciurus carolinensis'') from
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
. However, the population in
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to ...
is stabilising due to conservation efforts, awareness and the increasing population of the
pine marten The European pine marten (''Martes martes''), also known as the pine marten, is a mustelid native to and widespread in most of Europe, Asia Minor, the Caucasus and parts of Iran, Iraq and Syria. It is listed as Least Concern on the IUCN Red List. ...
, a European predator that selectively controls grey squirrels.


Description

The red squirrel has a typical head-and-body length of , a tail length of , and a mass of . Males and females are the same size. The red squirrel is somewhat smaller than the eastern grey squirrel which has a head-and-body length of and weighs between . The long tail helps the squirrel to balance and steer when jumping from tree to tree and running along branches and may keep the animal warm during sleep. The red squirrel, like most tree squirrels, has sharp curved
claw A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or tarsus ...
s to help it to climb and descend broad tree trunks, thin branches, and even house walls. Its strong hind legs let it leap gaps between trees. The red squirrel also can swim. The coat of the red squirrel varies in colour with time of year and location. There are several coat colour morphs ranging from black to red. Red coats are most common in Great Britain; in other parts of Europe and Asia different coat colours coexist within populations, much like hair colour in some human populations. The underside of the squirrel is always white-cream in colour. The red squirrel sheds its coat twice a year, switching from a thinner summer coat to a thicker, darker winter coat with noticeably larger ear-tufts (a prominent distinguishing feature of this species) between August and November. A lighter, redder overall coat colour, along with the ear-tufts (in adults) and smaller size, distinguish the Eurasian red squirrel from the American eastern grey squirrel. The red colour is for camouflage when seen against the bark of pine trees.


Distribution and habitat

Red squirrels occupy
boreal Boreal may refer to: Climatology and geography *Boreal (age), the first climatic phase of the Blytt-Sernander sequence of northern Europe, during the Holocene epoch *Boreal climate, a climate characterized by long winters and short, cool to mild ...
,
coniferous woods A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
in northern Europe and Siberia, preferring
Scots pine ''Pinus sylvestris'', the Scots pine (UK), Scotch pine (US) or Baltic pine, is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-green leaves and orang ...
,
Norway spruce ''Picea abies'', the Norway spruce or European spruce, is a species of spruce native to Northern, Central and Eastern Europe. It has branchlets that typically hang downwards, and the largest cones of any spruce, 9–17 cm long. It is very close ...
and Siberian pine. In western and southern Europe they are found in broad-leaved woods where the mixture of tree and shrub species provides a better year-round source of food. In most of the British Isles and in Italy, broad-leaved woodlands are now less suitable due to the better competitive feeding strategy of introduced grey squirrels.


Ecology and behaviour

The red squirrel is found in both coniferous forest and temperate broadleaf woodlands. The squirrel makes a drey (nest) out of twigs in a branch-fork, forming a domed structure about 25 to 30 cm in diameter. This is lined with
moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta ('' sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and ...
, leaves, grass and
bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, e ...
. Tree hollows and
woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions ...
holes are also used. The red squirrel is a solitary animal and is shy and reluctant to share food with others. However, outside the
breeding season Seasonal breeders are animal species that successfully mate only during certain times of the year. These times of year allow for the optimization of survival of young due to factors such as ambient temperature, food and water availability, and ch ...
and particularly in winter, several red squirrels may share a drey to keep warm. Social organization is based on dominance hierarchies within and between sexes; although males are not necessarily dominant to females, the dominant animals tend to be larger and older than subordinate animals, and dominant males tend to have larger home ranges than subordinate males or females. The red squirrel eats mostly the
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s of trees, neatly stripping conifer cones to get at the seeds within,
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately fr ...
, nuts (especially
hazelnut The hazelnut is the fruit of the hazel tree and therefore includes any of the nuts deriving from species of the genus '' Corylus'', especially the nuts of the species ''Corylus avellana''. They are also known as cobnuts or filberts according ...
s but also beech, chestnuts and acorns),
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, ras ...
,
vegetable Vegetables are parts of plants that are consumed by humans or other animals as food. The original meaning is still commonly used and is applied to plants collectively to refer to all edible plant matter, including the edible flower, flowers, ...
s, garden
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s, tree sap and young
shoot In botany, a plant shoot consists of any plant stem together with its appendages, leaves and lateral buds, flowering stems, and flower buds. The new growth from seed germination that grows upward is a shoot where leaves will develop. In the sp ...
s.Forestry Commission – Red Squirrels
. Forestry.gov.uk. Retrieved on 25 July 2013.
More rarely, red squirrels may also eat bird eggs or nestlings. A Swedish study shows that out of 600 stomach contents of red squirrels examined, only 4 contained remnants of birds or eggs. Excess food is put into caches, either buried or in nooks or holes in trees, and eaten when food is scarce. Although the red squirrel remembers where it created caches at a better-than-chance level, its
spatial memory In cognitive psychology and neuroscience, spatial memory is a form of memory responsible for the recording and recovery of information needed to plan a course to a location and to recall the location of an object or the occurrence of an event. Sp ...
is substantially less accurate and durable than that of grey squirrels. Between 60% and 80% of its active period may be spent foraging and feeding. The active period for the red squirrel is in the morning and in the late afternoon and evening. It often rests in its nest in the middle of the day, avoiding the heat and the high visibility to
birds of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predat ...
that are dangers during these hours. During the winter, this mid-day rest is often much briefer, or absent entirely, although harsh weather may cause the animal to stay in its nest for days at a time. No territories are claimed between the red squirrels, and the feeding areas of individuals overlap considerably.


Reproduction

Mating can occur in late winter during February and March and in summer between June and July. Up to two litters a year per female are possible. Each litter averages three young, called kits.
Gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pr ...
is about 38 to 39 days. The young are looked after by the mother alone and are born helpless, blind, and deaf. They weigh between 10 and 15  g. Their body is covered by hair at 21 days, their eyes and ears open after three to four weeks, and they develop all their teeth by 42 days. Juvenile red squirrels can eat solids around 40 days following birth and from that point can leave the nest on their own to find food; however, they still suckle from their mother until
weaning Weaning is the process of gradually introducing an infant human or another mammal to what will be its adult diet while withdrawing the supply of its mother's milk. The process takes place only in mammals, as only mammals produce milk. The infan ...
occurs at 8 to 10 weeks. During mating, males detect females that are in
oestrus The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrou ...
by an odour that they produce, and although there is no courtship, the male will chase the female for up to an hour prior to mating. Usually, several males will chase a single female until the dominant male, usually the largest in the group, mates with the female. Males and females will mate several times with many partners. Females must reach a minimum body mass before they enter oestrus, and heavy females on average produce more young. If food is scarce breeding may be delayed. Typically a female will produce her first litter in her second year.


Life expectancy

Red squirrels that survive their first winter have a life expectancy of 3 years. Individuals may reach 7 years of age, and 10 in captivity. Survival is positively related to the availability of autumn-winter tree seeds; on average, 75–85% of juveniles die during their first winter, and mortality is approximately 50% for winters following the first.


Enemies and threats

Arboreal predators include small mammals such as the
pine marten The European pine marten (''Martes martes''), also known as the pine marten, is a mustelid native to and widespread in most of Europe, Asia Minor, the Caucasus and parts of Iran, Iraq and Syria. It is listed as Least Concern on the IUCN Red List. ...
,
wildcat The wildcat is a species complex comprising two small wild cat species: the European wildcat (''Felis silvestris'') and the African wildcat (''F. lybica''). The European wildcat inhabits forests in Europe, Anatolia and the Caucasus, while th ...
s and the
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
, which preys on nestlings; birds, including owls and raptors such as the goshawk and buzzards, may also take the red squirrel. The
red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the Order (biology), order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe ...
, cats and dogs can prey upon the red squirrel when it is on the ground. Humans influence the population size and mortality of the red squirrel by destroying or altering habitats, by causing road casualties, and by introducing non-native populations of the North American eastern grey squirrels. The eastern grey squirrel and the red squirrel are not directly antagonistic, and violent conflict between these species is not a factor in the decline in red squirrel populations. However, the eastern grey squirrel appears to be able to decrease the red squirrel population due to several reasons: *The eastern grey squirrel carries a disease, the
squirrel parapoxvirus ''Squirrelpox virus (SQPV)'' is a virus that causes the fatal disease squirrelpox in United Kingdom red squirrels. The virus is often carried by grey squirrels from North America, which rarely die from the disease. Elsewhere in the Red Squirrel' ...
, that does not appear to affect their own health but will often kill the red squirrel. It was revealed in 2008 that the numbers of red squirrels at Formby (England) had declined by 80% as a result of this disease, though the population is now recovering. *The eastern grey squirrel can better digest
acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera '' Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and b ...
s, while the red squirrel cannot access the proteins and fats in acorns as easily. *When the red squirrel is put under pressure, it will not breed as often. In the UK, due to the above circumstances, the population has today fallen to 160,000 red squirrels or fewer (120,000 of these are in Scotland). Outside the UK and Ireland, the impact of competition from the eastern grey squirrel has been observed in
Piedmont it, Piemontese , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = , demographics1_footnotes = , demographics1_title1 = , demographics1_info1 = , demographics1_title2 ...
, Italy, where two pairs escaped from captivity in 1948. A significant drop in red squirrel populations in the area has been observed since 1970, and it is feared that the eastern grey squirrel may expand into the rest of Europe.


Conservation

The red squirrel is protected in most of Europe, as it is listed in Appendix III of the Bern Convention; it is listed as being of
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
. However, in some areas it is abundant and is hunted for its fur. Although not thought to be under any threat worldwide, the red squirrel has nevertheless drastically reduced in number in the United Kingdom; especially after the grey squirrels were introduced from North America in the 1870s. Fewer than 140,000 individuals are thought to be left in 2013; approximately 85% of which are in Scotland, with the
Isle of Wight The Isle of Wight ( ) is a county in the English Channel, off the coast of Hampshire, from which it is separated by the Solent. It is the largest and second-most populous island of England. Referred to as 'The Island' by residents, the Is ...
being the largest haven in England. A local charity, the Wight Squirrel Project, supports red squirrel conservation on the island, and islanders are actively recommended to report any invasive greys. The population decrease in Britain is often ascribed to the introduction of the eastern grey squirrel from North America, but the loss and fragmentation of its native woodland habitat has also played a significant role. In contrast, the red squirrel may present a threat if introduced to regions outside its native range. It is classed as a "prohibited new organism" under New Zealand's
Hazardous Substances and New Organisms Act 1996 The Hazardous Substances and New Organisms Act (HSNO) is an Act of Parliament passed in New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Isl ...
preventing it from being imported into the country.


Projects

In January 1998, eradication of the non-native North American grey squirrel began on the North Wales island of
Anglesey Anglesey (; cy, (Ynys) Môn ) is an island off the north-west coast of Wales. It forms a principal area known as the Isle of Anglesey, that includes Holy Island across the narrow Cymyran Strait and some islets and skerries. Anglesey island ...
. This facilitated the natural recovery of the small remnant red squirrel population. It was followed by the successful reintroduction of the red squirrel into the pine stands of Newborough Forest. Subsequent reintroductions into broadleaved woodland followed and today the island has the single largest red squirrel population in Wales.
Brownsea Island Brownsea Island is the largest of the islands in Poole Harbour in the county of Dorset, England. The island is owned by the National Trust with the northern half managed by the Dorset Wildlife Trust. Much of the island is open to the public and ...
in
Poole Harbour Poole Harbour is a large natural harbour in Dorset, southern England, with the town of Poole on its shores. The harbour is a drowned valley ( ria) formed at the end of the last ice age and is the estuary of several rivers, the largest bei ...
is also populated exclusively by red rather than grey squirrels (approximately 200 individuals). Mainland initiatives in southern Scotland and the north of England also rely upon grey squirrel control as the cornerstone of red squirrel conservation strategy. A local programme known as the "North East Scotland Biodiversity Partnership", an element of the national Biodiversity Action Plan was established in 1996. This programme is administered by the Grampian Squirrel Society, with an aim of protecting the red squirrel; the programme centres on the Banchory and
Cults In modern English, ''cult'' is usually a pejorative term for a social group that is defined by its unusual religious, spiritual, or philosophical beliefs and rituals, or its common interest in a particular personality, object, or goal. This ...
areas. In 2008, the Scottish Wildlife Trust announced a four-year project which commenced in the spring of 2009 called "Saving Scotland's Red Squirrels". Other notable projects include red squirrel projects in the Greenfield Forest, including the buffer zones of Mallerstang,
Garsdale Garsdale is a dale or valley in the south east of Cumbria, England, historically a part of the West Riding of Yorkshire. It is now within the South Lakeland local government district, but is still a "Yorkshire Dale" for planning purposes. In ...
and Widdale; the Northumberland
Kielder Forest Kielder Forest is a large forestry plantation in Northumberland, England, surrounding Kielder village and the Kielder Water reservoir. It is the largest man-made woodland in England with three-quarters of its covered by forest. The majority o ...
Project; and within the National Trust reserve in Formby. These projects were originally part of the Save Our Squirrels campaign that aimed to protect red squirrels in the north of England, but now form part of a five-year Government-led partnership conservation project called "Red Squirrels Northern England" to undertake grey squirrel control in areas important for red squirrels. On the Isle of Wight, local volunteers are encouraged to record data on the existing red squirrel population, and to monitor it for the presence of invasive greys; as the red squirrel is still dominant on the island, these volunteers are also requested to cull any greys they find. In order to protect existing populations, increasing amounts of legislation have been issued to prevent the further release and expansion of grey squirrel populations. Under the
Wildlife and Countryside Act 1981 The Wildlife and Countryside Act 1981 is an Act of Parliament in the United Kingdom implemented to comply with European Council Directive 79/409/EEC on the conservation of wild birds. In short, the act gives protection to native species (especia ...
, it is an offense to release captured grey squirrels, indicating that any captured individuals must be culled. Additional rules covered under the WCA's Schedules 5 and 6 include limitations on the keeping of red squirrels in captivity, and also prohibits the culling of red squirrels. Research undertaken in 2007 in the UK credits the
pine marten The European pine marten (''Martes martes''), also known as the pine marten, is a mustelid native to and widespread in most of Europe, Asia Minor, the Caucasus and parts of Iran, Iraq and Syria. It is listed as Least Concern on the IUCN Red List. ...
with reducing the population of the invasive eastern grey squirrel. Where the range of the expanding pine marten population meets that of the eastern grey squirrel, the population of these squirrels retreats. It is theorised that, because the grey squirrel spends more time on the ground than the red, they are far more likely to come in contact with this predator. During October 2012, four male and one female red squirrel, on permanent loan from the British Wildlife Centre, were transported to Tresco in the
Isles of Scilly The Isles of Scilly (; kw, Syllan, ', or ) is an archipelago off the southwestern tip of Cornwall, England. One of the islands, St Agnes, is the most southerly point in Britain, being over further south than the most southerly point of th ...
by helicopter, and released into Abbey Wood, near the Abbey Gardens. Only two survived and a further 20 were transported and released in October 2013. Although the red squirrel is not indigenous to the Isles of Scilly, those who supported this work intend to use Tresco as a ″safe haven″ for the endangered mammal, as the islands are free of predators such as foxes, and of the
squirrel pox ''Squirrelpox virus (SQPV)'' is a virus that causes the fatal disease squirrelpox in United Kingdom red squirrels. The virus is often carried by grey squirrels from North America, which rarely die from the disease. Elsewhere in the Red Squirre ...
-carrying grey squirrel.


Historical, cultural and financial significance

Squirrel Nutkin is a character, always illustrated as a red squirrel, in English author
Beatrix Potter Helen Beatrix Potter (, 28 July 186622 December 1943) was an English writer, illustrator, natural scientist, and conservationist. She is best known for her children's books featuring animals, such as '' The Tale of Peter Rabbit'', which was ...
's books for children. "Ekorren" (The Squirrel) is a well-known and appreciated children's song in Sweden. Text and lyrics by Alice Tegnér in 1892. Charles Dennim, protagonist of
Geoffrey Household Geoffrey Edward West Household (30 November 1900 – 4 October 1988) was a prolific British novelist who specialized in thrillers. He is best known for his novel '' Rogue Male'' (1939). Personal life He was born in Bristol; his father Horace ...
's novel ''Watcher in the Shadows,'' is a zoologist who studies and writes about red squirrels. In
Norse mythology Norse, Nordic, or Scandinavian mythology is the body of myths belonging to the North Germanic peoples, stemming from Old Norse religion and continuing after the Christianization of Scandinavia, and into the Nordic folklore of the modern per ...
,
Ratatoskr In Norse mythology, Ratatoskr (Old Norse, generally considered to mean "drill-tooth"Orchard (1997:129), Simek (2007:261), and Byock (2005:173). or "bore-tooth"Lindow (2001:259).) is a squirrel who runs up and down the world tree Yggdrasil to carry ...
is a red squirrel who runs up and down with messages in the world tree, Yggdrasil, and spreads gossip. In particular, he carried messages between the unnamed eagle at the top of Yggdrasill and the wyrm
Níðhöggr In Norse mythology, Níðhöggr (''Malice Striker'', in Old Norse traditionally also spelled Níðhǫggr , often anglicized NidhoggWhile the suffix of the name, ''-höggr'', clearly means "striker" the prefix is not as clear. In particular, the ...
beneath its roots. The red squirrel used to be widely hunted for its pelt. In Finland, squirrel pelts were used as currency in ancient times, before the introduction of coinage. The expression "squirrel pelt" is still widely understood there to be a reference to money. It has been suggested that the trade in red squirrel fur, highly prized in the medieval period and intensively traded, may have been responsible for the
leprosy Leprosy, also known as Hansen's disease (HD), is a long-term infection by the bacteria '' Mycobacterium leprae'' or '' Mycobacterium lepromatosis''. Infection can lead to damage of the nerves, respiratory tract, skin, and eyes. This nerve d ...
epidemic in medieval Europe. Within Great Britain, widespread leprosy is found early in East Anglia, to which many of the squirrel furs were traded, and the strain is the same as that found in modern red squirrels on
Brownsea Island Brownsea Island is the largest of the islands in Poole Harbour in the county of Dorset, England. The island is owned by the National Trust with the northern half managed by the Dorset Wildlife Trust. Much of the island is open to the public and ...
. The red squirrel is the national mammal of Denmark. Red squirrels are a common feature in English
heraldry Heraldry is a discipline relating to the design, display and study of armorial bearings (known as armory), as well as related disciplines, such as vexillology, together with the study of ceremony, rank and pedigree. Armory, the best-known bran ...
, where they are always depicted sitting up and often in the act of cracking a nut.


Taxonomy

There have been over 40 described
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
of the red squirrel, but the taxonomic status of some of these is uncertain. A study published in 1971 recognises 16 subspecies and has served as a basis for subsequent taxonomic work. Although the validity of some subspecies is labelled with uncertainty because of the large variation in red squirrels even within a single region, others are relatively distinctive and one of these, ''S. v. meridionalis'' of South Italy, was elevated to species status as the
Calabrian black squirrel The Calabrian black squirrel (''Sciurus meridionalis'') is a species of tree squirrel in the genus ''Sciurus'', endemic to the forests of the regions of Calabria and Basilicata, in the south of the Italian Peninsula. Taxonomy It has long been ...
in 2017. At present, there are 23 recognized subspecies of the red squirrel. Genetic studies indicate that another, ''S. v. hoffmanni'' of Sierra Espuña in southeast Spain (below included in ''S. v. alpinus''), deserves recognition as distinct. *''S. v. alpinus''. Desmarest, 1822. (synonyms: ''S. v. baeticus'', ''hoffmanni'', ''infuscatus'', ''italicus'', ''numantius'' and ''segurae'') *''S. v. altaicus''. Serebrennikov, 1928. *''S. v. anadyrensis''. Ognev, 1929. *''S. v. arcticus''. Trouessart, 1906. (synonym: ''S. v. jacutensis'') *''S. v. balcanicus''. Heinrich, 1936. (synonyms: ''S. v. istrandjae'' and ''rhodopensis'') *''S. v. chiliensis''. Sowerby, 1921. *''S. v. cinerea''. Hermann, 1804. *''S. v. dulkeiti''. Ognev, 1929. *''S. v. exalbidus''. Pallas, 1778. (synonyms: ''S. v. argenteus'' and ''kalbinensis'') *''S. v. fedjushini''. Ognev, 1935. *''S. v. formosovi''. Ognev, 1935. *''S. v. fuscoater''. Altum, 1876. (synonyms: ''S. v. brunnea'', ''gotthardi'', ''graeca'', ''nigrescens'', ''russus'' and ''rutilans'') *''S. v. fusconigricans''. Dvigubsky, 1804 *''S. v. leucourus''. Kerr, 1792. *''S. v. lilaeus''. Miller, 1907. (synonyms: ''S. v. ameliae'' and ''croaticus'') *''S. v. mantchuricus''.
Thomas Thomas may refer to: People * List of people with given name Thomas * Thomas (name) * Thomas (surname) * Saint Thomas (disambiguation) * Thomas Aquinas (1225–1274) Italian Dominican friar, philosopher, and Doctor of the Church * Thomas the A ...
, 1909. (synonyms: ''S. v. coreae'' and ''coreanus'') *''S. v. martensi''. Matschie, 1901. (synonym: ''S. v. jenissejensis'') *''S. v. ognevi''. Migulin, 1928. (synonyms: ''S. v. bashkiricus'', ''golzmajeri'' and ''uralensis'') *''S. v. orientis''.
Thomas Thomas may refer to: People * List of people with given name Thomas * Thomas (name) * Thomas (surname) * Saint Thomas (disambiguation) * Thomas Aquinas (1225–1274) Italian Dominican friar, philosopher, and Doctor of the Church * Thomas the A ...
, 1906. Ezo Red Squirrel (
Hokkaidō is Japan, Japan's Japanese archipelago, second largest island and comprises the largest and northernmost Prefectures of Japan, prefecture, making up its own List of regions of Japan, region. The Tsugaru Strait separates Hokkaidō from Honshu; th ...
). *''S. v. rupestris''.
Thomas Thomas may refer to: People * List of people with given name Thomas * Thomas (name) * Thomas (surname) * Saint Thomas (disambiguation) * Thomas Aquinas (1225–1274) Italian Dominican friar, philosopher, and Doctor of the Church * Thomas the A ...
, 1907 *''S. v. ukrainicus''. Migulin, 1928. (synonym: ''S. v. kessleri'') *''S. v. varius''. Gmelin, 1789. *''S. v. vulgaris''.
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
,
1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of ''Systema Naturae'', the starting point of modern zoologi ...
. (synonyms: ''S. v. albonotatus'', ''albus'', ''carpathicus'', ''europaeus'', ''niger'', ''rufus'' and ''typicus'')


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A partnership to save the red squirrel, between the Scottish Wildlife Trust, et al.Red squirrels make a comeback as culling of grey rivals brings resultsAcute Fatal Toxoplasmosis in Three Eurasian Red Squirrels
* * * {{Authority control Mammals described in 1758 Rodents of Europe Mammals of Azerbaijan Mammals of Mongolia Mammals of Korea Mammals of China Mammals of Saint Kitts and Nevis Mammals of the Caribbean Sciurus Articles containing video clips Taxa named by Carl Linnaeus