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The red deer (''Cervus elaphus'') is one of the largest
deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the re ...
species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region,
Anatolia Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The ...
,
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
, and parts of western Asia. It also inhabits the Atlas Mountains of Northern Africa; its early ancestors are thought to have crossed over to
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria t ...
, then to
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
,
Libya Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to the east, Suda ...
and
Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...
via the Strait of Gibraltar, becoming the only species of true deer (
Cervidae Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reind ...
) to inhabit Africa. Red deer have been introduced to other areas, including Australia,
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
,
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
,
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
,
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
and
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. In many parts of the world, the meat ( venison) from red deer is used as a food source. Red deer are
ruminant Ruminants (suborder Ruminantia) are hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbial actions. The ...
s, characterized by a four-chambered stomach. Genetic evidence indicates that the red deer, as traditionally defined, is a
species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
, rather than a single species, though exactly how many species the group includes remains disputed. The closely related and slightly larger American elk, or wapiti, native to North America and northeastern Asia, had been regarded as a subspecies of red deer, but recently it has been established as a distinct species. The ancestor of all red deer (and wapiti) probably originated in central Asia and resembled
sika deer The sika deer (''Cervus nippon''), also known as the Northern spotted deer or the Japanese deer, is a species of deer native to much of East Asia and introduced to other parts of the world. Previously found from northern Vietnam in the south to ...
. Additional genetic research has suggested that the closest living species to the red deer is the
fallow deer ''Dama'' is a genus of deer in the subfamily Cervinae, commonly referred to as fallow deer. Name The name fallow is derived from the deer's pale brown colour. The Latin word ''dāma'' or ''damma'', used for roe deer, gazelles, and antelopes ...
, ''Dama dama''. Although at one time red deer were rare in parts of Europe, they were never close to extinction. Reintroduction and conservation efforts, such as in the United Kingdom and Portugal, have resulted in an increase of red deer populations, while other areas, such as North Africa, have continued to show a population decline.


Description

The red deer is the fourth-largest extant
deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the re ...
species, behind the moose, elk, and sambar deer. It is a ruminant, eating its food in two stages and having an even number of toes on each hoof, like camels,
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
s, and
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult ma ...
. European red deer have a relatively long tail compared to their Asian and North American relatives. Subtle differences in appearance are noted between the various subspecies of red deer, primarily in size and antlers, with the smallest being the
Corsican red deer The Corsican red deer (''Cervus elaphus corsicanus''), also known simply as the Corsican deer or Sardinian deer, is a population of red deer (''Cervus elaphus'') found on the Mediterranean islands of Sardinia (Italy) and Corsica (France). Chara ...
found on the islands of Corsica and
Sardinia Sardinia ( ; it, Sardegna, label=Italian, Corsican and Tabarchino ; sc, Sardigna , sdc, Sardhigna; french: Sardaigne; sdn, Saldigna; ca, Sardenya, label=Algherese and Catalan) is the second-largest island in the Mediterranean Sea, after ...
and the largest being the Caspian red deer (or maral) of Asia Minor and the Caucasus Region to the west of the
Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad steppe of Central Asia ...
. The deer of central and western Europe vary greatly in size, with some of the largest deer found in the Carpathian Mountains in Central Europe. Western European red deer, historically, grew to large size given ample food supply (including people's crops), and descendants of introduced populations living in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
and
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
have grown quite large in both body and antler size. Large red deer stags, like the Caspian red deer or those of the Carpathian Mountains, may rival North American elk in size. Female red deer are much smaller than their male counterparts.


Size

The male (stag) red deer is typically long from the nose to the base of the tail and typically weighs ; the female (hind) is long and often weighs . The tail adds another and shoulder height is about . In
Scotland Scotland (, ) is a Countries of the United Kingdom, country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a Anglo-Scottish border, border with England to the southeast ...
, stags average in head-and-body length and high at the shoulder and females average long and tall. Based on body mass, they are likely the fourth largest extant deer species on average, behind the moose, the elk and the sambar deer. Size varies in different subspecies with the largest, the huge but small-antlered deer of the Carpathian Mountains (''C. e. elaphus''), weighing up to . At the other end of the scale, the Corsican red deer (''C. e. corsicanus'') weighs about , although red deer in poor habitats can weigh as little as .


Neck mane

The males of many subspecies also grow a short neck mane during the autumn. The male deer of the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
and
Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and the ...
tend to have the thickest and most noticeable manes. Male Caspian red deer (''C. e. maral'') and Spanish red deer (''C. e. hispanicus'') do not carry neck manes. Male deer of all subspecies, however, tend to have stronger and thicker neck muscles than female deer, which may give them an appearance of having neck manes. Red deer hinds (females) do not have neck manes.


Antlers

Only the stags have antlers, which start growing in the spring and are shed each year, usually at the end of winter. Antlers typically measure in total length and weigh , although large ones can grow to and weigh . Antlers, which are made of bone, can grow at a rate of a day. While an antler is growing, it is covered with highly vascular skin called velvet, which supplies oxygen and nutrients to the growing bone. The antlers are
testosterone Testosterone is the primary sex hormone and anabolic steroid in males. In humans, testosterone plays a key role in the development of male reproductive tissues such as testes and prostate, as well as promoting secondary sexual characteristi ...
-driven and as the stag's testosterone levels drop in the autumn, the velvet is shed and the antlers stop growing. With the approach of autumn, the antlers begin to calcify and the stags' testosterone production builds for the approaching rut (mating season). European red deer antlers are distinctive in being rather straight and
rugose Rugose means "wrinkled". It may refer to: * Rugosa, an extinct order of coral, whose rugose shape earned it the name * Rugose, adjectival form of rugae Species with "rugose" in their names * ''Idiosoma nigrum'', more commonly, a black rugose trap ...
, with the fourth and fifth tines forming a "crown" or "cup" in larger males. Any tines in excess of the fourth and fifth tines grow radially from the cup, which are generally absent in the antlers of smaller red deer, such as Corsican red deer. Western European red deer antlers feature "bez" (second) tines that are either absent or smaller than the brow tines. However, bez tines occur frequently in Norwegian red deer. Antlers of Caspian red deer carry large bez tines and form less-developed cups than western European red deer, their antlers are thus more like the "throw back" top tines of the North American elk (''C. canadensis''), known as maraloid characteristics. A stag can (exceptionally) have antlers with no tines, and is then known as a switch. Similarly, a stag that does not grow antlers is a hummel.


Coat

European red deer tend to be reddish-brown in their summer coats, and some individuals may have a few spots on the backs of their summer coats. During the autumn, all red deer subspecies grow thicker coats of hair, which helps to insulate them during the winter. Autumn is also when some of the stags grow their neck manes. The autumn/winter coats of most subspecies are most distinct. The Caspian red deer's winter coat is greyer and has a larger and more distinguished light rump-patch (like wapiti and some central Asian red deer) compared to the Western European red deer, which has more of a greyish-brown coat with a darker yellowish rump patch in the winter. By the time summer begins, the heavy winter coat has been shed; the animals are known to rub against trees and other objects to help remove hair from their bodies. Red deer have different colouration based on the seasons and types of habitats, with grey or lighter colouration prevalent in the winter and more reddish and darker coat colouration in the summer.


Distribution

The ''
Cervus ''Cervus'' is a genus of deer that primarily are native to Eurasia, although one species occurs in northern Africa and another in North America. In addition to the species presently placed in this genus, it has included a whole range of othe ...
''
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
ancestors of red deer first appear in fossil records 12 million years ago during the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
in
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago ...
.


Europe and North Africa

The European red deer is found in southwestern Asia (Asia Minor and Caucasus regions), North Africa, and Europe. The red deer is the largest nondomesticated land mammal still existing in
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
. The Barbary stag (which resembles the western European red deer) is the only member of the deer family represented in Africa, with the population centred in the northwestern region of the continent in the Atlas Mountains. As of the mid-1990s,
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria t ...
,
Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...
, and
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
were the only African countries known to have red deer. In the Netherlands, a large herd (about 3000 animals counted in late 2012) lives in the Oostvaardersplassen, a nature reserve. Ireland has its own unique subspecies. In France, the population is thriving, having multiplied five-fold in the last half-century, increasing from 30,000 in 1970 to around 160,000 in 2014. The deer has particularly expanded its footprint into forests at higher altitudes than before. In the UK, indigenous populations occur in
Scotland Scotland (, ) is a Countries of the United Kingdom, country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a Anglo-Scottish border, border with England to the southeast ...
, the Lake District, and the south west of England (principally on
Exmoor Exmoor is loosely defined as an area of hilly open moorland in west Somerset and north Devon in South West England. It is named after the River Exe, the source of which is situated in the centre of the area, two miles north-west of Simonsbath ...
). Not all of these are of entirely pure bloodlines, as some of these populations have been supplemented with deliberate releases of deer from parks, such as
Warnham Warnham is a village and civil parish in the Horsham district of West Sussex, England. The village is centred north-northwest of Horsham, from London, to the west of the A24 road. Other named settlements within the parish include the hamlets o ...
or
Woburn Abbey Woburn Abbey (), occupying the east of the village of Woburn, Bedfordshire, England, is a country house, the family seat of the Duke of Bedford. Although it is still a family home to the current duke, it is open on specified days to visitors, ...
, in an attempt to increase antler sizes and body weights. The
University of Edinburgh The University of Edinburgh ( sco, University o Edinburgh, gd, Oilthigh Dhùn Èideann; abbreviated as ''Edin.'' in post-nominals) is a public research university based in Edinburgh, Scotland. Granted a royal charter by King James VI in 15 ...
found that, in Scotland, extensive hybridisation with the closely related sika deer has occurred. Several other populations have originated either with "carted" deer kept for stag hunts being left out at the end of the hunt, escapes from deer farms, or deliberate releases. Carted deer were kept by stag hunts with no wild red deer in the locality and were normally recaptured after the hunt and used again; although the hunts are called "stag hunts", the Norwich Staghounds only hunted hinds (female red deer), and in 1950, at least eight hinds (some of which may have been pregnant) were known to be at large near Kimberley and
West Harling Harling is a civil parish in the English county of Norfolk. It covers an area of and had a population of 2,201 in 932 households at the 2001 census,Thetford Forest in
Norfolk Norfolk () is a ceremonial and non-metropolitan county in East Anglia in England. It borders Lincolnshire to the north-west, Cambridgeshire to the west and south-west, and Suffolk to the south. Its northern and eastern boundaries are the No ...
. Further substantial red deer herds originated from escapes or deliberate releases in the
New Forest The New Forest is one of the largest remaining tracts of unenclosed pasture land, heathland and forest in Southern England, covering southwest Hampshire and southeast Wiltshire. It was proclaimed a royal forest by William the Conqueror, fea ...
, the Peak District, Suffolk,
Lancashire Lancashire ( , ; abbreviated Lancs) is the name of a historic county, ceremonial county, and non-metropolitan county in North West England. The boundaries of these three areas differ significantly. The non-metropolitan county of Lancash ...
, Brecon Beacons, and
North Yorkshire North Yorkshire is the largest ceremonial counties of England, ceremonial county (lieutenancy area) in England, covering an area of . Around 40% of the county is covered by National parks of the United Kingdom, national parks, including most of ...
, as well as many other smaller populations scattered throughout
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
and
Wales Wales ( cy, Cymru ) is a country that is part of the United Kingdom. It is bordered by England to the east, the Irish Sea to the north and west, the Celtic Sea to the south west and the Bristol Channel to the south. It had a population in ...
, and they are all generally increasing in numbers and range. A census of deer populations in 2007 and again in 2011 coordinated by the British Deer Society records the red deer as having continued to expand their range in England and Wales since 2000, with expansion most notable in
the Midlands The Midlands (also referred to as Central England) are a part of England that broadly correspond to the Kingdom of Mercia of the Early Middle Ages, bordered by Wales, Northern England and Southern England. The Midlands were important in the In ...
and East Anglia.


Iran

Caspian red deer are found in the Hyrcanian Forests.


New Zealand

In
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, red deer were introduced by acclimatisation societies along with other deer and game species. The first red deer to reach New Zealand were a pair sent by Lord Petre in 1851 from his herd at Thorndon Park, Essex, to the South Island, but the hind was shot before they had a chance to breed. Lord Petre sent another stag and two hinds in 1861, and these were liberated near
Nelson Nelson may refer to: Arts and entertainment * ''Nelson'' (1918 film), a historical film directed by Maurice Elvey * ''Nelson'' (1926 film), a historical film directed by Walter Summers * ''Nelson'' (opera), an opera by Lennox Berkeley to a lib ...
, from where they quickly spread. The first deer to reach the North Island were a gift to Sir
Frederick Weld Sir Frederick Aloysius Weld (9 May 1823 – 20 July 1891), was a New Zealand politician and a governor of various British colonies. He was the List of Prime Ministers of New Zealand, sixth Prime Minister of New Zealand, premier of New Zealand, ...
from Windsor Great Park and were released near
Wellington Wellington ( mi, Te Whanganui-a-Tara or ) is the capital city of New Zealand. It is located at the south-western tip of the North Island, between Cook Strait and the Remutaka Range. Wellington is the second-largest city in New Zealand by metr ...
; these were followed by further releases up to 1914. Between 1851 and 1926, 220 separate liberations of red deer involved over 800 deer. In 1927, the State Forest Service introduced a bounty for red deer shot on their land, and in 1931, government control operations were commenced. Between 1931 and March 1975, 1,124,297 deer were killed on official operations. The introduced red deer have adapted well and are widely hunted on both islands; many of the 220 introductions used deer originating from Scotland ( Invermark) or one of the major deer parks in England, principally Warnham, Woburn Abbey or Windsor Great Park. Some hybridisation happened with the closely related American elk (''Cervus canadensis nelsoni'') introduced in Fiordland in 1921. New Zealand red deer produce very large antlers and are regarded as amongst the best in the world by hunters. Along with the other introduced deer species, they are, however, officially regarded as a noxious pest and are still heavily culled using professional hunters working with helicopters, or even poisoned.


Australia

The first red deer to reach Australia were probably the six that Prince Albert sent in 1860 from Windsor Great Park to Thomas Chirnside, who was starting a herd at
Werribee Park Werribee Park is the estate of a historical building in Werribee, Victoria, Australia. It includes Werribee Park Mansion, the Victoria State Rose Garden, formal gardens, the Werribee Park National Equestrian Centre, the Werribee Open Range Zoo, ...
, south west of Melbourne in Victoria. Further introductions were made in
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
,
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , establishe ...
,
South Australia South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...
, and
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
. Today, red deer in Australia range from Queensland south through New South Wales into Victoria and across to South Australia, with the numbers increasing. The Queensland, Victorian and most New South Wales strains can still be traced to the early releases, but South Australia's population, along with all others, is now largely recent farm escapees. This is having adverse effects on the integrity of wild herds, as now more and larger herds are being grown due to the superior genetics that have been attained by selective breeding. Wild red deer are a feral pest species in Australia, do considerable harm to the natural environment, and are a significant road traffic hazard.


Argentina and Chile

In Argentina and Chile, the red deer has had a potentially adverse impact on native animal species, such as the South Andean deer or ''huemul''; the International Union for Conservation of Nature and Natural Resources has labelled the animal as one of the world's 100 worst invaders.


Migration

Red deer in Europe generally spend their winters at lower altitudes in more wooded terrain. During the summer, they migrate to higher elevations where food supplies are greater and better for the calving season.


Taxonomy and evolution

Until recently, biologists considered the red deer and elk or wapiti ('' C. canadensis'') the same species, forming a continuous distribution throughout temperate Eurasia and North America. This belief was based largely on the fully fertile hybrids that can be produced under captive conditions. Genetic evidence clearly shows the wapiti and red deer form two separate species. Another member of the red deer group which may represent a separate species is '' C. corsicanus''. If so, ''C. corsicanus'' includes the subspecies '' C. e. barbarus'' (perhaps a synonym of ''C. e. corsicanus''), and is restricted to
Maghreb The Maghreb (; ar, الْمَغْرِب, al-Maghrib, lit=the west), also known as the Arab Maghreb ( ar, المغرب العربي) and Northwest Africa, is the western part of North Africa and the Arab world. The region includes Algeria, ...
in North Africa, Corsica, and
Sardinia Sardinia ( ; it, Sardegna, label=Italian, Corsican and Tabarchino ; sc, Sardigna , sdc, Sardhigna; french: Sardaigne; sdn, Saldigna; ca, Sardenya, label=Algherese and Catalan) is the second-largest island in the Mediterranean Sea, after ...
. The
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of nat ...
originally listed nine subspecies of red deer (''Cervus elaphus''): three as endangered, one as vulnerable, one as
near threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify f ...
, and four without enough data to give a category ( Data Deficient). The species as a whole, however, is listed as least concern. However, this was based on the traditional classification of red deer as one species (''Cervus elaphus''), including the wapiti. The common red deer is also known as simply red deer. Selected members of the red deer
species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
are listed in the table below. Of the ones listed, ''C. e. hippelaphus'' and ''C. e. scoticus'' may be junior synonyms. ''Cervus elaphus'' appeared in Europe by the beginning of the
Middle Pleistocene The Chibanian, widely known by its previous designation of Middle Pleistocene, is an age in the international geologic timescale or a stage in chronostratigraphy, being a division of the Pleistocene Epoch within the ongoing Quaternary Period. Th ...
around 800,000 years ago. These earliest forms belonged to the
palaeosubspecies A chronospecies is a species derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is p ...
'' Cervus elaphus acoronatus.'' Other palaeosubspecies are known, including those belonging to ''C. elaphus rianensis'' from the Middle Pleistocene of Italy, ''C. elaphus siciliae'' from the late Middle and Late Pleistocene of Sicily.


Behaviour

Mature red deer (''C. elaphus'') usually stay in single-sex groups for most of the year. During the mating season, called the rut, mature stags compete for the attentions of the hinds and will then try to defend the hinds they attract. Rival stags challenge opponents by belling and walking in parallel. This allows combatants to assess each other's antlers, body size and fighting prowess. If neither stag backs down, a clash of antlers can occur, and stags sometimes sustain serious injuries. Red deer are among the mammals exhibiting homosexual behavior.Bagemihl, Bruce (1999), Biological Exuberance: Animal Homosexuality and Natural Diversity. St. Martin's Press , page 378 Dominant stags follow groups of hinds during the rut, from August into early winter. The stags may have as many as 20 hinds to keep from other, less attractive males. Only mature stags hold harems (groups of hinds), and breeding success peaks at about eight years of age. Stags two to four years old rarely hold harems and spend most of the rut on the periphery of larger harems, as do stags over 11 years old. Young and old stags that do acquire a harem hold it later in the breeding season than those stags in their prime. Harem-holding stags rarely feed and lose up to 20% of their body weight. Stags that enter the rut in poor condition are less likely to make it through to the peak conception period. Male European red deer have a distinctive roar during the rut, which is an adaptation to forested environments, in contrast to male American elk stags which "bugle" during the rut in adaptation to open environments. The male deer roars to keep his harem of females together. The females are initially attracted to those males that both roar most often and have the loudest roar call. Males also use the roar call when competing with other males for females during the rut, and along with other forms of posturing and antler fights, is a method used by the males to establish dominance. Roaring is most common during the early dawn and late evening, which is also when the crepuscular deer are most active in general.


Breeding, gestation and lifespan

Female red deer reach
sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans it might be considered synonymous with adulthood, but here puberty is the name for the process of biological sexual maturation, while adulthood is based on cultural definit ...
at 2 years of age. Red deer mating patterns usually involve a dozen or more
mating In biology, mating is the pairing of either opposite- sex or hermaphroditic organisms for the purposes of sexual reproduction. ''Fertilization'' is the fusion of two gametes. ''Copulation'' is the union of the sex organs of two sexually reprod ...
attempts before the first successful one. There may be several more matings before the stag will seek out another mate in his harem. Females in their second autumn can produce one or very rarely two offspring per year. The gestation period is 240 to 262 days, and the offspring weigh about . After two weeks, calves are able to join the herd and are fully weaned after two months. The offspring will remain with their mothers for almost one full year, leaving around the time the next season's offspring are produced. The gestation period is the same for all subspecies. All red deer calves are born spotted, as is common with many deer species, and lose their spots by the end of summer. However, as in many species of Old World deer, some adults do retain a few spots on the backs of their summer coats. Red deer live over 20 years in captivity and in the wild they live 10 to 13 years, though some subspecies with less predation pressure average 15 years.


Protection from predators

Male red deer retain their antlers for more than half the year, and are less gregarious and less likely to group with other males when they have antlers. The antlers provide self-defence, as does a strong front-leg kicking action performed by both sexes when attacked. Once the antlers are shed, stags tend to form bachelor groups which allow them to cooperatively work together. Herds tend to have one or more members watching for potential danger, while the remaining members eat and rest. After the rut, females form large herds of up to 50 individuals. The newborn calves are kept close to the hinds by a series of vocalizations between the two, and larger nurseries have an ongoing and constant chatter during the daytime hours. When approached by predators, the largest and most robust females may make a stand, using their front legs to kick at their attackers. Guttural grunts and posturing is used with all but the most determined of predators with great effectiveness. Aside from humans and domestic dogs, the grey wolf is probably the most dangerous predator European red deer encounter. Occasionally, the brown bear will prey on European red deer.


Red deer in folklore and art

Red deer are widely depicted in cave art found throughout European caves, with some of the artwork dating from as early as 40,000 years ago, during the
Upper Paleolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
.
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive region, geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a ...
n cave art from the
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several p ...
of 7,000 years ago has abundant depictions of red deer, including what can be described as spiritual artwork, indicating the importance of this mammal to the peoples of that region (Note: these animals were most likely wapiti (''C. canadensis'') in Siberia, not red deer). Red deer are also often depicted on
Pictish Pictish is the extinct Brittonic language spoken by the Picts, the people of eastern and northern Scotland from Late Antiquity to the Early Middle Ages. Virtually no direct attestations of Pictish remain, short of a limited number of geographica ...
stones (''circa'' 550–850 AD), from the early medieval period in Scotland, usually as prey animals for human or animal predators. In medieval hunting, the red deer was the most prestigious quarry, especially the mature stag, which in England was called a
hart Hart often refers to: * Hart (deer) Hart may also refer to: Organizations * Hart Racing Engines, a former Formula One engine manufacturer * Hart Skis, US ski manufacturer * Hart Stores, a Canadian chain of department stores * Hart's Reptile Wo ...
.


Red deer products

Red deer are held in captivity for a variety of reasons. The meat of the deer, called venison, was until recently restricted in the United Kingdom to those with connections to the aristocratic or
poaching Poaching has been defined as the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set a ...
communities, and a licence was needed to sell it legally, but it is now widely available in supermarkets, especially in the autumn. The
Queen Queen or QUEEN may refer to: Monarchy * Queen regnant, a female monarch of a Kingdom ** List of queens regnant * Queen consort, the wife of a reigning king * Queen dowager, the widow of a king * Queen mother, a queen dowager who is the mother ...
followed the custom of offering large pieces of venison to members of the Cabinet of the United Kingdom and others. Some estates in the
Scottish Highlands The Highlands ( sco, the Hielands; gd, a’ Ghàidhealtachd , 'the place of the Gaels') is a historical region of Scotland. Culturally, the Highlands and the Lowlands diverged from the Late Middle Ages into the modern period, when Lowland S ...
still sell deer-stalking accompanied by a gillie in the traditional way, on unfenced land, while others operate more like farms for venison. Venison is widely considered to be both flavourful and nutritious. It is higher in
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, res ...
and lower in
fat In nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such compounds, most commonly those that occur in living beings or in food. The term often refers specifically to triglycerides (triple est ...
than either beef or
chicken The chicken (''Gallus gallus domesticus'') is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster or cock is a term for an adu ...
. The red deer can produce of antler velvet annually. On ranches in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, China,
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive region, geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a ...
, and elsewhere, this velvet is collected and sold to markets in East Asia, where it is used for holistic medicines, with
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korean Peninsula and sharing a land border with North Korea. Its western border is formed by the Yellow Sea, while its eas ...
being the primary consumer. In
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eig ...
, a medication produced from antler velvet is sold under the brand name Pantokrin (russian: Пантокри́н; la, Pantocrinum). The antlers themselves are also believed by East Asians to have medicinal purposes and are often ground up and used in small quantities. Historically, related deer species such as Central Asian red deer, wapiti, Thorold's deer, and
sika deer The sika deer (''Cervus nippon''), also known as the Northern spotted deer or the Japanese deer, is a species of deer native to much of East Asia and introduced to other parts of the world. Previously found from northern Vietnam in the south to ...
have been reared on deer farms in Central and Eastern Asia by
Han Chinese The Han Chinese () or Han people (), are an East Asian ethnic group native to China. They constitute the world's largest ethnic group, making up about 18% of the global population and consisting of various subgroups speaking distinctiv ...
,
Turkic peoples The Turkic peoples are a collection of diverse ethnic groups of West, Central, East, and North Asia as well as parts of Europe, who speak Turkic languages.. "Turkic peoples, any of various peoples whose members speak languages belonging t ...
, Tungusic peoples,
Mongolians The Mongols ( mn, Монголчууд, , , ; ; russian: Монголы) are an East Asian ethnic group native to Mongolia, Inner Mongolia in China and the Buryatia Republic of the Russian Federation. The Mongols are the principal member of ...
, and Koreans. In modern times, western countries such as New Zealand and United States have taken to farming European red deer for similar purposes. Deer hair products are also used in the fly fishing industry, being used to tie flies. Deer antlers are also used for decorative purposes and have been used for artwork, furniture and other novelty items. Deer antlers were and still are the source material for horn furniture. Already in the 15th century trophies of case were used for clothes hook, storage racks and chandeliers, the so-called '' Lusterweibchen''. In the 19th century the European nobility discovered red deer antlers as perfect decorations for their manors and hunting castles. This fashion trend splashes over to upper- and middle-class households in the mid of the 19th century. At the increasingly popular World Expositions, producers of horn furniture, mainly in Germany, Austria and the United States, such as and Friedrich Wenzel, showed their horn furniture and a kind of series manufacturing began. In recent times deer antler home decors can be found in home styling magazines.Newmann, Bruce M. (1985). ''Fantasy Furniture''. New York: Rizzoli.


Gallery

File:Deer Stag AdF.jpg, Mature Spanish red deer bellowing during the rut File:Cervus.elaphus.track.on.snow.jpg, Red deer tracks in Commanster, in the Ardennes of
Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to th ...
File:Red deer (Cervus elaphus) young stag.jpg, Young stag in
Great Glen The Great Glen ( gd, An Gleann Mòr ), also known as Glen Albyn (from the Gaelic "Glen of Scotland" ) or Glen More (from the Gaelic ), is a glen in Scotland running for from Inverness on the edge of Moray Firth, in an approximately straight ...
in
Scotland Scotland (, ) is a Countries of the United Kingdom, country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a Anglo-Scottish border, border with England to the southeast ...
File:Red deer stag.jpg, A soft covering known as velvet helps to protect newly forming antlers in the spring. Glen Torridon, Scotland File:Cervus elaphus - Salburúa.jpg, One male and three females in the Salburua wetlands of Basque Country in
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
File:Red deer (Cervus elaphus) hind with juvenile.jpg, Hind with juvenile in Great Glen in Scotland File:Hirved1.jpg, Young stags fleeing on the island of
Saaremaa Saaremaa is the largest island in Estonia, measuring . The main island of Saare County, it is located in the Baltic Sea, south of Hiiumaa island and west of Muhu island, and belongs to the West Estonian Archipelago. The capital of the isla ...
in
Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, a ...
File:Red deer portrait.jpg, Red deer in the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...


See also

* Deer of Great Britain * Deer farm * Medieval deer park * Venison


References


Further reading

*Cameron, Donald; Fortescue, Hugh Fortescue 3rd, earl; Shand, Alexander Innes (1896
''Red Deer: Natural history''
London, New York tc.: Longmans, Green and co. *Clarke, J. (1866),
The naturalist: A treatise on the growth of the horns of the red deer
', Barnstaple, A.P. Wood *Heptner, V. G.; Nasimovich, A. A.; Bannikov, A. G.; Hoffman, R. S. (1988
''Mammals of the Soviet Union''
Volume I, Washington, D.C. : Smithsonian Institution Libraries and National Science Foundation *Jeffries, Richard (1884)
''Red Deer''
London Longmans, Green#B. *


External links


World of Deer Museum New ZealandNew Zealand Red Stag Hunting Informationred deer antler furnishings
{{DEFAULTSORT:deer, red red deer red deer Mammals of Europe Mammals of North Africa Mammals of Western Asia National symbols of Ireland red deer red deer