Ramshorn snail
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The term ''ramshorn snail'' or ''ram's horn snail'' is used in two different ways. In the
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
trade it is used to describe various kinds of
freshwater snail Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs ...
s whose shells are planispiral, meaning that the shell is a flat coil. Such shells resemble a coil of rope, or (as the name suggests) a
ram Ram, ram, or RAM may refer to: Animals * A male sheep * Ram cichlid, a freshwater tropical fish People * Ram (given name) * Ram (surname) * Ram (director) (Ramsubramaniam), an Indian Tamil film director * RAM (musician) (born 1974), Dutch * ...
's horn. In a more general natural history context, the term "ramshorn snails" is used more precisely to mean those aquatic
pulmonate Pulmonata or pulmonates, is an informal group (previously an order, and before that a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includ ...
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s in the family
Planorbidae Planorbidae, common name the ramshorn snails or ram's horn snails, is a family of air-breathing freshwater snails, aquatic pulmonate gastropod molluscs. Unlike most molluscs, the blood of ram's horn snails contains iron-based hemoglobin instea ...
that have planispiral coiled shells. Ramshorn snails have been bred for the aquarium trade and various color forms have been selected. The two species commonly found in aquariums are ''
Planorbella duryi ''Planorbella duryi'', common name the Seminole rams-horn, is a species of air-breathing freshwater snail, a pulmonate gastropod mollusk in the family Planorbidae, the ram's horn snails. The species is endemic to Florida and is found frequently ...
'' and ''
Planorbarius corneus ''Planorbarius corneus'', common name the great ramshorn, is a relatively large species of air-breathing freshwater snail, an aquatic pulmonate gastropod mollusk in the family Planorbidae, the ram's horn snails, or planorbids, which all have ...
'', both in the family ''
Planorbidae Planorbidae, common name the ramshorn snails or ram's horn snails, is a family of air-breathing freshwater snails, aquatic pulmonate gastropod molluscs. Unlike most molluscs, the blood of ram's horn snails contains iron-based hemoglobin instea ...
''. One species (Columbia ramshorn, Marisa cornuarietis) is from a totally different family, the ''
Ampullariidae Ampullariidae, commonly known as the apple snails, is a family of large freshwater snails, aquatic gastropod mollusks with a gill and an operculum. These snails simultaneously have a gill and a lung as functional respiratory structures ...
''. Ramshorn snails can sometimes become a nuisance in an aquarium, because some of them breed so profusely.


Description

Most of these snails are of the family
Planorbidae Planorbidae, common name the ramshorn snails or ram's horn snails, is a family of air-breathing freshwater snails, aquatic pulmonate gastropod molluscs. Unlike most molluscs, the blood of ram's horn snails contains iron-based hemoglobin instea ...
, and they include the species ''
Planorbarius corneus ''Planorbarius corneus'', common name the great ramshorn, is a relatively large species of air-breathing freshwater snail, an aquatic pulmonate gastropod mollusk in the family Planorbidae, the ram's horn snails, or planorbids, which all have ...
''. There are two different coloured skin forms: black and red; the latter lack the dark skin pigment
melanin Melanin (; from el, μέλας, melas, black, dark) is a broad term for a group of natural pigments found in most organisms. Eumelanin is produced through a multistage chemical process known as melanogenesis, where the oxidation of the amino ...
and consequently have a bright reddish skin, which is the colour of their blood. Their
blood Blood is a body fluid in the circulatory system of humans and other vertebrates that delivers necessary substances such as nutrients and oxygen to the cells, and transports metabolic waste products away from those same cells. Blood in the c ...
contains red
hemoglobin Hemoglobin (haemoglobin BrE) (from the Greek word αἷμα, ''haîma'' 'blood' + Latin ''globus'' 'ball, sphere' + ''-in'') (), abbreviated Hb or Hgb, is the iron-containing oxygen-transport metalloprotein present in red blood cells (erythrocyte ...
, unlike other snails'
blood Blood is a body fluid in the circulatory system of humans and other vertebrates that delivers necessary substances such as nutrients and oxygen to the cells, and transports metabolic waste products away from those same cells. Blood in the c ...
, which contains greenish
hemocyanin Hemocyanins (also spelled haemocyanins and abbreviated Hc) are proteins that transport oxygen throughout the bodies of some invertebrate animals. These metalloproteins contain two copper atoms that reversibly bind a single oxygen molecule (O2) ...
. These ramshorn snails breathe air. Although most of them are extremely small, some may reach a size of two and a half centimeters (one inch). The shells range from translucent through various shades of brown to a dark, nearly black color. The dark color appears to originate from dietary materials not generally available in the home aquarium, although many varieties from ponds are this dark shade. There are two common skin colourations: black and red. The red colouration is the colour of their blood (containing hemoglobin), unlike other snails with green blood (due to hemocyanin). An absence of melanin reveals the red colour. Rare varieties have been selectively bred in pale pink and blue. These traits are recessive and if crossbred the resultant offspring are likely to be wild type (brown). Snails of this family are spiralled sinistrally, with the opening hole slanted downward toward the right. Large folds of skin may protrude out of the more open left side. Like all air-breathing water snails, the animal has no operculum, and has only one pair of
tentacle In zoology, a tentacle is a flexible, mobile, and elongated organ present in some species of animals, most of them invertebrates. In animal anatomy, tentacles usually occur in one or more pairs. Anatomically, the tentacles of animals work main ...
s with the eye spots at the base of the tentacles. Ramshorn snails have a lifespan on one year.


Breeding

Ramshorn snails are
hermaphroditic In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have separ ...
; Red Ramshorn Snail Factoids
/ref> two organisms of any sex have the ability to breed and produce offspring. Ramshorn snails lay eggs in globules, which tend to be brownish in color. The globules contain about a dozen or so eggs, though it can vary. The globules are translucent, so it is possible to visually see the new snails develop in size. The newborn snails are clearish white.


Interaction with environment

Ramshorn snails generally will eat only the most delicate plants, preferring
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...
, uneaten fish food, and dead
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
. Some varieties do particularly enjoy eating the leaves of stem plants such as cabomba and anacharis. Some aquarium species will eat ramshorn snails. More voracious eaters include puffers,
loach Loaches are fish of the superfamily Cobitoidea. They are freshwater, benthic (bottom-dwelling) fish found in rivers and creeks throughout Eurasia and northern Africa. Loaches are among the most diverse groups of fish; the 1249 known species of ...
es (such as the
clown loach The clown loach (''Chromobotia macracanthus''), or tiger botia, is a tropical freshwater fish belonging to the botiid loach family. It is the sole member of the genus ''Chromobotia''. It originates in inland waters in Indonesia on the islands ...
or any other member of the genus Botia),
crayfish Crayfish are freshwater crustaceans belonging to the clade Astacidea, which also contains lobsters. In some locations, they are also known as crawfish, craydids, crawdaddies, crawdads, freshwater lobsters, mountain lobsters, rock lobsters, mu ...
, and most
gourami Gouramis, or gouramies , are a group of freshwater anabantiform fishes that comprise the family Osphronemidae. The fish are native to Asia—from the Indian Subcontinent to Southeast Asia and northeasterly towards Korea. The name "gourami", of ...
s— though many other fish will also consume snail meat. Apple snails and assassin snails will also prey upon ramshorn snails. Good fish roommates for snails include, but are not limited to,
danios ''Danio'' is a genus of small freshwater fish in the family Cyprinidae found in South and Southeast Asia, commonly kept in aquaria. They are generally characterised by a pattern of horizontal stripes, rows of spots or vertical bars. Some species ...
,
guppies The guppy (), also known as millionfish and rainbow fish, is one of the world's most widely distributed tropical fish and one of the most popular List of freshwater aquarium fish species, freshwater aquarium fish species. It is a member of the ...
,
White Cloud Mountain minnow The White Cloud Mountain minnow (''Tanichthys albonubes'') is a hardy species of freshwater fish and coldwater fish often kept in an aquarium. The species is a member of the carp family (family Cyprinidae) of the order Cypriniformes, native to C ...
s,
neon tetra The neon tetra (''Paracheirodon innesi'') is a Freshwater fish of the characin family (family Characidae) of order Characiformes. The type species of its genus, it is native to blackwater and clearwater streams in the Amazon basin of South Am ...
s, and
cory catfish ''Corydoras'' is a genus of freshwater catfish in the family Callichthyidae and subfamily Corydoradinae. The species usually have more restricted areas of endemism than other callichthyids, but the area of distribution of the entire genus almost ...
. All of these are non-aggressive fish that cohabit easily with snails. One should also be aware that pond-reared red ramshorn snails are able to carry various parasitic flukes, which can be transmitted to fish, or humans. Most of these flukes require intermediate
host A host is a person responsible for guests at an event or for providing hospitality during it. Host may also refer to: Places * Host, Pennsylvania, a village in Berks County People *Jim Host (born 1937), American businessman * Michel Host ...
s, so that leaving the snails in a fish-free aquarium for a month or so will eliminate any parasites. If the population is kept to a manageable size, ramshorn snails can be good tank cleaners. They eat algae and dead or dying plants generally, so they can be useful. However, if they breed too prolifically they can become a nuisance. In warm climates (such as those in mainland Australia or the southern United States) they much prefer ponds, especially outdoor ponds. Algae, dead leaves that sink to the bottom, mulm and dead animals can be a problem, as they foul the water. Ramshorn snails eat all of these things.


Role as aquarium pest

Most ramshorn snails are considered minor aquarium pests. They may arrive in a tank as
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
bundles hidden in newly acquired
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
s. Although their red color may make them somewhat interesting aquarium subjects, their
hermaphroditic In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have separ ...
ability to breed prolifically from any two specimens can make them troublesome. Absolute eradication is difficult, but their numbers can usually be kept to a moderate level. Common methods to reduce population include treating plants to prevent introduction, various manual methods of control, introducing the snail-eating animals listed earlier, and poisoning the snails. Soaking the plants in various
chemical A chemical substance is a form of matter having constant chemical composition and characteristic properties. Some references add that chemical substance cannot be separated into its constituent elements by physical separation methods, i.e., wi ...
s may kill off the snails and their eggs. A 10-minute bath in a solution of 20 parts water to 1 part chlorine
bleach Bleach is the generic name for any chemical product that is used industrially or domestically to remove color (whitening) from a fabric or fiber or to clean or to remove stains in a process called bleaching. It often refers specifically, to ...
has been suggested for hardier plants, followed by soaking in water containing a dechlorinating agent. A more gentle treatment calls for 5-10 tablespoons of
alum An alum () is a type of chemical compound, usually a hydrated double salt, double sulfate salt (chemistry), salt of aluminium with the general chemical formula, formula , where is a valence (chemistry), monovalent cation such as potassium or a ...
to 1 gallon of water for 2–3 days. A safer alternative may be placing the plants in a
quarantine A quarantine is a restriction on the movement of people, animals and goods which is intended to prevent the spread of disease or pests. It is often used in connection to disease and illness, preventing the movement of those who may have been ...
tank, and adding snail poisons to that tank rather than the main show tank. Manual methods include baiting the snails with lettuce (run it under hot water first and leave overnight),
cucumber Cucumber (''Cucumis sativus'') is a widely-cultivated Vine#Horticultural climbing plants, creeping vine plant in the Cucurbitaceae family that bears usually cylindrical Fruit, fruits, which are used as culinary vegetables.
slices, or food pellets. These may be left out in the open, and removed with their snails, or kept in some container, such as a film canister weighed down with a pebble, and containing holes drilled in it. Crushing the snails by hand as they appear can also effectively limit their population; most ramshorn shells are fragile enough that this is quite easily done. Introducing animals to control a snail population can require some thought. Other aquarium fish may not be compatible, and some larger adult snails may be too big to be eaten by smaller snail eating species. It may occasionally be necessary to crush a few snails manually so that the fish realize the snail can be eaten. Snail eating species also do not usually discriminate between different types of snails, although this is usually not much of a concern. Snail poisons are generally considered to be a last resort, as most of them are
copper Copper is a chemical element with the symbol Cu (from la, cuprum) and atomic number 29. It is a soft, malleable, and ductile metal with very high thermal and electrical conductivity. A freshly exposed surface of pure copper has a pinkis ...
-based and are potentially toxic to plants and fish and particularly dangerous to other invertebrates. Even new safer chemicals that do not harm the other aquarium inhabitants may cause damage if large numbers of dead snails are allowed to decompose. For this reason, it is best to reduce the snail population by other means as much as possible before resorting to poisons, and to do frequent water changes afterwards. Some also recommend adding
ammonium The ammonium cation is a positively-charged polyatomic ion with the chemical formula or . It is formed by the protonation of ammonia (). Ammonium is also a general name for positively charged or protonated substituted amines and quaternary a ...
protection to the tank. Zeolite chips, and various liquid products such as amquel may help in this area.


"Giant ramshorn" snails

Totally unrelated to these ramshorn snails is the species ''
Marisa cornuarietis ''Marisa cornuarietis'', common name the Colombian ramshorn apple snail, is a species of large freshwater snail with an operculum, an aquatic gastropod mollusc in the family Ampullariidae, the apple snail family. These snails are popular ...
'', which is often sold at pet stores under the name "Columbian ramshorn" or "giant ramshorn" snail. This species is actually a kind of
apple snail Ampullariidae, commonly known as the apple snails, is a family of large freshwater snails, aquatic gastropod mollusks with a gill and an operculum. These snails simultaneously have a gill and a lung as functional respiratory structures, which ...
, albeit one with a planispiral shell. They are distinguished by having an operculum by which they can close themselves into their shell, two pairs of tentacles, separate genders, and a siphon on the left side. Its shell is yellowish, with brown stripes running the length of the shell. These apple snails lay gelatinous masses of eggs on submerged portions of plants. They can grow to up to four centimeters in size. They generally will not become a pest, although they can consume large amounts of plant matter. They are very large compared to other Ramshorn snails.


References


External links


Ramshorn Snails
on The Aquarium Wiki {{DEFAULTSORT:Ramshorn Snail Planorbidae Mollusc common names