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Rye (''Secale cereale'') is a grass grown extensively as a
grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and legum ...
, a
cover crop In agriculture, cover crops are plants that are planted to cover the soil rather than for the purpose of being harvested. Cover crops manage soil erosion, soil fertility, soil quality, water, weeds, pests, diseases, biodiversity and wildlife i ...
and a forage crop. It is a member of the wheat tribe ( Triticeae) and is closely related to both wheat (''Triticum'') and barley (genus ''Hordeum''). Rye grain is used for
flour Flour is a powder made by grinding raw grains, roots, beans, nuts, or seeds. Flours are used to make many different foods. Cereal flour, particularly wheat flour, is the main ingredient of bread, which is a staple food for many culture ...
,
bread Bread is a staple food prepared from a dough of flour (usually wheat) and water, usually by baking. Throughout recorded history and around the world, it has been an important part of many cultures' diet. It is one of the oldest human-made f ...
, beer, crispbread, some whiskeys, some vodkas, and animal
fodder Fodder (), also called provender (), is any agriculture, agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, domestic rabbit, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food g ...
. It can also be eaten whole, either as boiled rye berries or by being rolled, similar to rolled oats. Rye is a
cereal A cereal is any Poaceae, grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, Cereal germ, germ, and bran. Cereal Grain, grain crops are grown in greater quantit ...
grain and should not be confused with ryegrass, which is used for lawns, pasture, and as hay for livestock.


Distribution and habitat

Rye is one of a number of species that grow wild in the Levant, central and eastern Turkey and in adjacent areas. Evidence uncovered at the Epipalaeolithic site of Tell Abu Hureyra in the
Euphrates The Euphrates () is the longest and one of the most historically important rivers of Western Asia. Tigris–Euphrates river system, Together with the Tigris, it is one of the two defining rivers of Mesopotamia ( ''the land between the rivers'') ...
valley of northern
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
suggests that rye was among the first
cereal A cereal is any Poaceae, grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, Cereal germ, germ, and bran. Cereal Grain, grain crops are grown in greater quantit ...
crops to be systematically cultivated, around 13,000 years ago. However, that claim remains controversial; critics point to inconsistencies in the radiocarbon dates, and identifications based solely on grain, rather than on
chaff Chaff (; ) is the dry, scaly protective casing of the seeds of cereal grains or similar fine, dry, scaly plant material (such as scaly parts of flowers or finely chopped straw). Chaff is indigestible by humans, but livestock can eat it. In agri ...
. Domesticated rye occurs in small quantities at a number of Neolithic sites in Asia Minor ( Anatolia, now Turkey), such as the Pre-Pottery Neolithic B
Can Hasan III Can may refer to: Containers * Aluminum can * Drink can * Oil can * Steel and tin cans * Trash can * Petrol can * Metal can (disambiguation) Music * Can (band), West Germany, 1968 ** ''Can'' (album), 1979 * Can (South Korean band) Other * ...
near Çatalhöyük, but is otherwise absent from the archaeological record until the Bronze Age of central Europe, c. 1800–1500 BCE. It is possible that rye traveled west from Asia Minor as a minor admixture in wheat (possibly as a result of
Vavilovian mimicry In plant biology, Vavilovian mimicry (also crop mimicry or weed mimicry) is a form of mimicry in plants where a weed evolves to share one or more characteristics with a domesticated plant through generations of artificial selection. It is name ...
), and was only later cultivated in its own right. Archeological evidence of this grain has been found in Roman contexts along the Rhine and the Danube and in Ireland and Britain. page 26. Roman naturalist Pliny the Elder was dismissive of a grain that may have been rye, writing that it "is a very poor food and only serves to avert starvation". He said it was mixed with
spelt Spelt (''Triticum spelta''), also known as dinkel wheat or hulled wheat, is a species of wheat that has been cultivated since approximately 5000 BC. Spelt was an important staple food in parts of Europe from the Bronze Age to medieval times. No ...
"to mitigate its bitter taste, and even then is most unpleasant to the stomach". Winter rye is any breed of rye planted in the autumn to provide ground cover for the winter. It grows during warmer days of the winter when sunlight temporarily warms the plant above freezing, even while there is general snow cover. It can be used to prevent the growth of winter-hardy weeds, and can either be harvested as a bonus crop or tilled directly into the ground in spring to provide more organic matter for the next summer's crop. It is sometimes used in winter gardens and is a common nurse crop. Rye grows better than any other cereal in heavy clay and light sandy and infertile or drought-affected soils. It can tolerate pH between 4.5 and 8.0, but soils having pH 5.0 to 7.0 are best suited for rye cultivation. Rye grows best in fertile, well-drained loam or clay-loam soils.


Frost resistance

''S. cereale'' can thrive in subzero environments. The leaves of winter rye produce various antifreeze polypeptides (different from the antifreeze polypeptides produced by some fish and insects).


Ecology

The nematode '' Ditylenchus dipsaci'', leaf beetle, fruit fly, gout fly, cereal chafer, dart moth, cereal bug, Hessian fly, and
rustic shoulder knot ''Apamea sordens'', the rustic shoulder-knot or bordered apamea, is a moth of the family Noctuidae. The species was first described by Johann Siegfried Hufnagel in 1766. It is distributed throughout Europe, east across the Palearctic to Central ...
are among insects which can seriously affect rye health.


Diseases

Rye is highly susceptible to the
ergot Ergot ( ) or ergot fungi refers to a group of fungi of the genus ''Claviceps''. The most prominent member of this group is ''Claviceps purpurea'' ("rye ergot fungus"). This fungus grows on rye and related plants, and produces alkaloids that ca ...
fungus. Consumption of ergot-infected rye by humans and animals results in a serious medical condition known as
ergotism Ergotism (pron. ) is the effect of long-term ergot poisoning, traditionally due to the ingestion of the alkaloids produced by the ''Claviceps purpurea'' fungus—from the Latin "club" or clavus "nail" and for "head", i.e. the purple club-head ...
. Ergotism can cause both physical and mental harm, including convulsions, miscarriage, necrosis of digits, hallucinations and death. Historically, damp northern countries that have depended on rye as a staple crop were subject to periodic epidemics of this condition. Such epidemics have been found to correlate with periods of frequent witch trials, such as the Salem witch trials in Massachusetts in 1692. Modern grain-cleaning and milling methods have practically eliminated the disease, but contaminated flour may end up in bread and other food products if the ergot is not removed before milling. After an absence of 60 years, stem rust (''Puccinia graminis'' f. sp. ''tritici'') has returned to Europe.Multiple sources: Areas affected include Germany, Russia ( Western Siberia), Spain, and
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
. Races PTGBH, PTGFH,
RKBBK Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, durum wh ...
, SKGBC,
SKGBH Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, durum whe ...
,
SKGBR Stem rust, also known as cereal rust, black rust, red rust or red dust, is caused by the fungus ''Puccinia graminis'', which causes significant disease in cereal crops. Crop species that are affected by the disease include bread wheat, durum whe ...
, TKDLK, TKGBK, TKGBR, and TKGDH are now present on rye in Spain.


Invasiveness

Rye is a common, unwanted invader of winter wheat fields. If allowed to grow and mature, it may cause substantially reduced prices (docking) for harvested wheat.


Cultivation

Since the Middle Ages, people have cultivated rye widely in
Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known as ...
and Eastern Europe. It serves as the main
bread Bread is a staple food prepared from a dough of flour (usually wheat) and water, usually by baking. Throughout recorded history and around the world, it has been an important part of many cultures' diet. It is one of the oldest human-made f ...
cereal in most areas east of the France–Germany border and north of Hungary. In Southern Europe, it was cultivated on marginal lands. Rye grows well in much poorer soils than those necessary for most cereal grains. Thus, it is an especially valuable crop in regions where the soil has sand or peat. Rye plants withstand cold better than other small grains do. Rye will survive with snow cover that would otherwise result in winter-kill for winter wheat. Most farmers grow winter ryes, which are planted and begin to grow in autumn. In spring, the plants develop and produce their crop. Autumn-planted rye shows fast growth; by the
summer solstice The summer solstice, also called the estival solstice or midsummer, occurs when one of Earth's poles has its maximum tilt toward the Sun. It happens twice yearly, once in each hemisphere ( Northern and Southern). For that hemisphere, the summer ...
, plants reach their maximum height of about a , while spring-planted wheat has only recently germinated. Vigorous growth suppresses even the most noxious weed competitors and rye can be grown without application of herbicides. Physical properties of rye affect attributes of the final food product such as seed size and surface area, and porosity. The surface area of the seed directly correlates to the drying and heat transfer time. Smaller seeds have increased heat transfer, which leads to lower drying time. Seeds with lower amounts of porosity also have lower tendencies to lose water during the process of drying.


Diversity and uses

Along with ''S. cereale'''s relationship and impact on the environment, it is also a valuable species because of its expansive diversity and uses. In northern Portugal, fourteen different populations of ''S. cereale'' were analyzed in order to better understand their differences. It was discovered that the storage proteins are very diverse and possess a lot of overall genetic variation as well, which is useful information to know because scientists can use its diversity in breeding to produce the most efficient cultivar of ''S. cereale''. Moreover, the beneficial characteristics of ''S. cereale'' can also be used to improve certain characteristics of other useful plants, like wheat. The pollination abilities of wheat were vastly improved when there was cross-pollination with ''S. cereale''. The addition of the rye chromosome 4R increased the size of the wheat anther along with increasing the number of pollen grains present. Along with improved wheat, the optimal characteristics of ''S. cereale'' can also be combined with another perennial rye, specifically ''S. montanum Guss'', in order to produce ''S. cereanum'', which has the beneficial characteristics of each. The hybrid rye (''S. cereanum'') can be grown in all environments, even with less than favorable soil and protects some soils from erosion. In addition, the plant mixture has improved forage and is known to contain digestible fiber and protein. Information about the diversity, the genome and ''S. cereanum'''s ability to cross fertilize with other species is useful information for scientists to know as they attempt to come up with various plant species that will be able to feed humanity in the future without leaving a negative footprint on the environment. The chromosome is the source of many
crop disease resistance gene Plant disease resistance protects plants from pathogens in two ways: by pre-formed structures and chemicals, and by infection-induced responses of the immune system. Relative to a susceptible plant, disease resistance is the reduction of pathoge ...
s. is a variety which has donated 1R-originating resistance to wheat, as have several other varieties including Insave, Amigo, and Imperial.


Harvesting

The
harvest Harvesting is the process of gathering a ripe crop from the fields. Reaping is the cutting of grain or pulse for harvest, typically using a scythe, sickle, or reaper. On smaller farms with minimal mechanization, harvesting is the most labor-i ...
ing of rye is similar to that of wheat. It is usually done with
combine harvester The modern combine harvester, or simply combine, is a versatile machine designed to efficiently harvest a variety of grain crops. The name derives from its combining four separate harvesting operations—reaping, threshing, gathering, and winnow ...
s, which cut the plants, thresh and
winnow Winnowing is a process by which chaff is separated from grain. It can also be used to remove pests from stored grain. Winnowing usually follows threshing in grain preparation. In its simplest form, it involves throwing the mixture into the ...
the grain, and release the
straw Straw is an agricultural byproduct consisting of the dry stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the yield of cereal crops such as barley, oats, rice, rye and wheat. It has a number ...
to the field where it is later pressed into bales or left as soil amendment. The resultant grain is stored in local
silo A silo (from the Greek σιρός – ''siros'', "pit for holding grain") is a structure for storing bulk materials. Silos are used in agriculture to store fermented feed known as silage, not to be confused with a grain bin, which is used t ...
s or transported to regional
grain elevator A grain elevator is a facility designed to stockpile or store grain. In the grain trade, the term "grain elevator" also describes a tower containing a bucket elevator or a pneumatic conveyor, which scoops up grain from a lower level and deposits ...
s and combined with other lots for storage and distant shipment. Before the era of
mechanised agriculture Mechanised agriculture or agricultural mechanization is the use of machinery and equipment, ranging from simple and basic hand tools to more sophisticated, motorized equipment and machinery, to perform agricultural operations. In modern times, po ...
, rye harvesting was a
manual Manual may refer to: Instructions * User guide * Owner's manual * Instruction manual (gaming) * Online help Other uses * Manual (music), a keyboard, as for an organ * Manual (band) * Manual transmission * Manual, a bicycle technique similar to ...
task performed with scythes or sickles. The cut rye was often shocked for drying or storage, and the threshing was done by manually beating the seed heads against a floor or other object.


Production and consumption statistics

Rye is grown primarily in Eastern, Central and Northern Europe. The main rye belt stretches from northern Germany through Poland, Ukraine, Belarus,
Lithuania Lithuania (; lt, Lietuva ), officially the Republic of Lithuania ( lt, Lietuvos Respublika, links=no ), is a country in the Baltic region of Europe. It is one of three Baltic states and lies on the eastern shore of the Baltic Sea. Lithuania ...
and
Latvia Latvia ( or ; lv, Latvija ; ltg, Latveja; liv, Leţmō), officially the Republic of Latvia ( lv, Latvijas Republika, links=no, ltg, Latvejas Republika, links=no, liv, Leţmō Vabāmō, links=no), is a country in the Baltic region of ...
into central and northern Russia. Rye is also grown in North America ( Canada and the United States), in South America ( Argentina, Brazil and Chile), in Oceania (
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
and New Zealand), in Turkey, in Kazakhstan and in northern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. Production levels of rye have fallen since 1992 in most of the producing nations, . For instance, production of rye in Russia fell from in 1992 to in 2020. Corresponding figures for other countries are as follows: Polandfalling from in 1992 to in 2020; Belarusfalling from in 1992 to in 2020; Chinafalling from in 1992 to in 2020. However, production levels rose in Germany from in 1992 to in 2020. World trade of rye is low compared with other grains such as wheat. The total export of rye for 2016 was $186M compared with $30.1B for wheat. Poland consumes the most rye per person at per capita (2009). Nordic and Baltic countries are also very high. The EU in general is around per capita. The entire world only consumes per capita.


Uses

Rye grain is refined into a
flour Flour is a powder made by grinding raw grains, roots, beans, nuts, or seeds. Flours are used to make many different foods. Cereal flour, particularly wheat flour, is the main ingredient of bread, which is a staple food for many culture ...
high in gliadin but low in glutenin. It therefore has a lower gluten content than wheat flour. It also contains a higher proportion of soluble fiber. Alkylresorcinols are phenolic lipids present in high amounts in the bran layer (e.g. pericarp,
testa Testa may refer to: * Testa (botany), a term to describe the seed coat * Testa (surname) * Testa, Hurwitz & Thibeault, a former Boston law firm * 11667 Testa, a main-belt asteroid discovered in 1997 * Testa (ceramics), fired clay material, espe ...
and
aleurone Aleurone (from Greek ''aleuron'', flour) is a protein found in protein granules of maturing seeds and tubers. The term also describes one of the two major cell types of the endosperm, the aleurone layer. The aleurone layer is the outermost layer ...
layers) of wheat and rye (0.1–0.3% of dry weight). Rye bread, including pumpernickel, is made using rye flour and is a widely eaten food in Northern and Eastern Europe. Rye is also used to make crisp bread. Rye grain is used to make alcoholic drinks, such as rye whiskey and rye beer. Other uses of rye grain include
kvass Kvass is a fermented cereal-based Alcohol by volume, low alcoholic beverage with a slightly cloudy appearance, light-brown colour and sweet-sour taste. It may be flavoured with berries, fruits, herbs or honey. Kvass stems from the northeastern ...
and an
herbal medicine Herbal medicine (also herbalism) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. With worldwide research into pharmacology, some herbal medicines have been translated into modern remed ...
known as rye extract. Rye
straw Straw is an agricultural byproduct consisting of the dry stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the yield of cereal crops such as barley, oats, rice, rye and wheat. It has a number ...
is used as livestock bedding, as a
cover crop In agriculture, cover crops are plants that are planted to cover the soil rather than for the purpose of being harvested. Cover crops manage soil erosion, soil fertility, soil quality, water, weeds, pests, diseases, biodiversity and wildlife i ...
and
green manure In agriculture, a green manure is a crop specifically produced to be incorporated into the soil while still green. Typically, the green manure's biomass is incorporated with a plow or disk, as is often done with (brown) manure. The primary goal ...
for
soil amendment A soil conditioner is a product which is added to soil to improve the soil’s physical qualities, usually its fertility (ability to provide nutrition for plants) and sometimes its mechanics. In general usage, the term "soil conditioner" is often ...
, and to make crafts such as corn dollies. Rye flour is used in the original way to make
Falun red Falu red or falun red ( ; sv, falu rödfärg, ) is a permeable red paint commonly used on wooden cottages and barns in Sweden, Finland, and Norway. History Following hundreds of years of mining in Falun, large piles of tailings, residual prod ...
paint (in addition to linseed oil and iron oxide) in Sweden. Rye grain (aka "Rye Berries") is a popular medium to use as a grain spawn when cultivating some varieties of edible mushrooms. The grain is cleaned, hydrated, and sterilized and then injected with mushroom spores and the mycelium grow using the grain to obtain water and nutrients.


Food


Nutrition

A reference serving of rye provides of food energy and is a rich source (20% or more of the Daily Value V of essential nutrients, including protein,
dietary fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...
, the B vitamins, niacin (27% DV) and vitamin B6 (23% DV), and several dietary minerals (table). Highest nutrient contents are for manganese (143% DV) and phosphorus (47% DV).


Health effects

According to
Health Canada Health Canada (HC; french: Santé Canada, SC)Health Canada is the applied title under the Federal Identity Program; the legal title is Department of Health (). is the Structure of the Canadian federal government#Departments, with subsidiary unit ...
and the U.S. Food and Drug Administration, consuming at least per day of rye beta-glucan or per serving of soluble fiber can lower levels of blood cholesterol, a risk factor for cardiovascular diseases. Eating whole-grain rye, as well as other high-fibre grains, improves regulation of blood sugar (i.e., reduces blood glucose response to a meal). Consuming
breakfast cereal Cereal, formally termed breakfast cereal (and further categorized as cold cereal or warm cereal), is a traditional breakfast food made from processed cereal grains. It is traditionally eaten as part of breakfast, or a snack food, primarily in ...
s containing rye over weeks to months also improved cholesterol levels and glucose regulation.


Health concerns

Like wheat, barley, and their hybrids and derivatives, rye contains gluten, which makes it an unsuitable grain for consumption by people with gluten-related disorders, such as celiac disease, non-celiac gluten sensitivity, and wheat allergy, among others. Nevertheless, some wheat allergy patients can tolerate rye or barley.
Ergotism Ergotism (pron. ) is the effect of long-term ergot poisoning, traditionally due to the ingestion of the alkaloids produced by the ''Claviceps purpurea'' fungus—from the Latin "club" or clavus "nail" and for "head", i.e. the purple club-head ...
is an illness that can result from eating rye and other grains infected by ergot fungi (which produce ergoline toxins in infected products). Although it is no longer a common illness because of modern
food safety Food safety (or food hygiene) is used as a scientific method/discipline describing handling, preparation, and storage of food in ways that prevent food-borne illness. The occurrence of two or more cases of a similar illness resulting from t ...
efforts, it was common before the 20th century, and it can still happen today if food safety vigilance breaks down.


Hybridization

Rye has long been considered an inferior grain to wheat in quality and digestibility, but has far larger kernels and is more hardy. In the 19th century, efforts were made to create a hybrid with the best qualities of both, known initially as triticosecale, but eventually becoming known as triticale. Initially fraught with fertility and germination problems, triticale is becoming more common worldwide in the 21st century, with millions of acres being produced.


Gallery


References


Further reading

* Schlegel provides
2011 updated version
online.


External links

* * * {{Authority control Cereals Plants described in 1753 Pooideae Staple foods