Rya Formation
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The Rya Formation (Swedish: Ryaformationen) is a
geologic formation A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics ( lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock exp ...
in Skåne County, southern Sweden. It is Early to early Middle Jurassic ( early Sinemurian to late Aalenian) in age. The Rya Formation comprises siltstones, claystones,
sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates ...
s, mudstones and rare
coal Coal is a combustible black or brownish-black sedimentary rock, formed as rock strata called coal seams. Coal is mostly carbon with variable amounts of other elements, chiefly hydrogen, sulfur, oxygen, and nitrogen. Coal is formed when ...
beds. The formation overlies the Höganäs Formation and is overlain by the Vilhelmsfält and Mariedal Formations. The formation was deposited in the
Höganäs Höganäs () (old da, Højenæs) is a locality and the seat of Höganäs Municipality, Scania County, Sweden with 14,107 inhabitants in 2010. Höganäs is nationally known for its ceramics industry, Höganäs Keramik. Höganäs Keramik is par ...
and Øresund Basins that formed in the earliest Jurassic as part of the break-up of
Pangea Pangaea or Pangea () was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from the earlier continental units of Gondwana, Euramerica and Siberia during the Carboniferous approximately 335 million y ...
. The thick formation comprises four members, from base to top the Döshult, Pankarp, Katslösa and Rydebäck Members. The
depositional environment In geology, depositional environment or sedimentary environment describes the combination of physical, chemical, and biological processes associated with the deposition of a particular type of sediment and, therefore, the rock types that will be ...
of the formation ranges from continental to open marine. The Rya Formation has provided fossils of a number of
shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachi ...
s, ammonites, bivalves and
ichnofossil A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils, ...
s. Coalified wood occurs as scattered pieces up to long and indeterminate
belemnite Belemnitida (or the belemnite) is an extinct order of squid-like cephalopods that existed from the Late Triassic to Late Cretaceous. Unlike squid, belemnites had an internal skeleton that made up the cone. The parts are, from the arms-most ...
s,
echinoid Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) of ...
s,
serpulid The Serpulidae are a family of sessile, tube-building annelid worms in the class Polychaeta. The members of this family differ from other sabellid tube worms in that they have a specialized operculum that blocks the entrance of their tubes wh ...
s,
ostracod Ostracods, or ostracodes, are a class of the Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 70,000 species (only 13,000 of which are extant) have been identified, grouped into several orders. They are small crustaceans, typi ...
s and nodosariid
foraminifera Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly ...
were also recorded in the formation. Iron
ooid Ooids are small (commonly ≤2 mm in diameter), spheroidal, "coated" (layered) sedimentary grains, usually composed of calcium carbonate, but sometimes made up of iron- or phosphate-based minerals. Ooids usually form on the sea floor, mo ...
s containing erratic boulders, called ''Geschiebe'' in German, attributed to the Rya Formation were found in
Holstein Holstein (; nds, label=Northern Low Saxon, Holsteen; da, Holsten; Latin and historical en, Holsatia, italic=yes) is the region between the rivers Elbe and Eider. It is the southern half of Schleswig-Holstein, the northernmost state of German ...
, northern Germany.


Description

The Rya Formation crops out in Colonus Shale Trough of western
Scania Scania, also known by its native name of Skåne (, ), is the southernmost of the historical provinces (''landskap'') of Sweden. Located in the south tip of the geographical region of Götaland, the province is roughly conterminous with Skåne ...
.Ahlberg et al., 2003, p.528 The formation overlies the Höganäs Formation and is overlain by the Vilhelmsfält Formation in the
Helsingborg Helsingborg (, , , ) is a city and the seat of Helsingborg Municipality, Scania (Skåne), Sweden. It is the second-largest city in Scania (after Malmö) and ninth-largest in Sweden, with a population of 113,816 (2020). Helsingborg is the cent ...
area and by the Mariedal Formation in the area of
Landskrona Landskrona (old da, Landskrone) is a town in Scania, Sweden. Located on the shores of the Öresund, it occupies a natural port, which has lent the town at first military and subsequent commercial significance. Ferries operate from Landskrona t ...
and
Kävlinge Kävlinge () is a locality and the seat of Kävlinge Municipality, Skåne County, Sweden with 32,341 inhabitants in 2021. In 1996, a train containing large amounts of ammonia derailed and around 9,000 people had to be evacuated from the area. Thi ...
.Ahlberg et al., 2003, p.530 The Rya Formation is subdivided, from base to top, into the Döshult, Pankarp, Katslösa and Rydebäck Members. The formation is found in the
Ängelholm Ängelholm is a locality and the seat of Ängelholm Municipality in Skåne, Sweden with 39,612 inhabitants in 2010. History The old settlement ''Rynestad'' was mentioned around the year 1600. The city was founded in 1516 as Engelholm by King Chri ...
, Helsingborg, Landskrona and Kävlinge areas. In southwest Skåne, the Rya Formation is missing or only poorly developed.Ahlberg et al., 2003, p.533 In the Øresund Basin between Sweden and Denmark, the formation is truncated by the Basal Middle Jurassic
unconformity An unconformity is a buried erosional or non-depositional surface separating two rock masses or strata of different ages, indicating that sediment deposition was not continuous. In general, the older layer was exposed to erosion for an interval ...
.Erlström et al., 2018, p.129 Erratic boulders, called ''Geschiebe'' in German, attributed to the Rya Formation and containing iron
ooid Ooids are small (commonly ≤2 mm in diameter), spheroidal, "coated" (layered) sedimentary grains, usually composed of calcium carbonate, but sometimes made up of iron- or phosphate-based minerals. Ooids usually form on the sea floor, mo ...
s were found in
Holstein Holstein (; nds, label=Northern Low Saxon, Holsteen; da, Holsten; Latin and historical en, Holsatia, italic=yes) is the region between the rivers Elbe and Eider. It is the southern half of Schleswig-Holstein, the northernmost state of German ...
, northern Germany.Hinz-Schallreuter & Schallreuter, 2009, p.2


Subdivision

;Döshult Member The early Sinemurian Döshult Member (Swedish ''Döshultsledet'') comprises coarse-grained cross-layered sandstones and siltstones in the lower part, and is dominated by dark clays and marls rich in marine fossils in the upper part.Andersson & Hybertsen, 2010-8-17 The member is up to thick in the Ängelholm, Helsingborg and Landskrona areas. Presently, the basal part of this member is exposed at three localities in the Helsingborg area. These contain mineralogically and texturally mature, trough cross-bedded sandstones, commonly with herringbone structures showing north and south oriented
paleocurrent A paleocurrent or paleocurrent indicator is a geological feature (typically a sedimentary structure) that helps one determine the direction of flowing water in the geologic past. This is an invaluable tool in the reconstruction of ancient deposit ...
directions. The occurrence of herringbone structures in well-sorted sand suggests high energy foreshore to subtidal marine depositional conditions for the lower part of the member. In an abandoned quarry in northwest Skåne (Gantofta brickpit in the Helsingborg area, outcrop very limited at present) the upper part of the Döshult Member commences with bioturbated marine nearshore sands, including burrows, as well as abundant marine invertebrate body fossils. This is followed by a bioturbated shelf mudstone with storm-deposited sand and silt intercalations ( tempestites). A massive red mudstone with scarce marine body fossils and burrows follows, which is interpreted as having been deposited rapidly, in a low energy but oxidizing environment. The youngest part of the succession comprises siltstones and mudstones, with carbonate-rich beds, deposited in a shallow marine setting. Coaly detritus,
muscovite Muscovite (also known as common mica, isinglass, or potash mica) is a hydrated phyllosilicate mineral of aluminium and potassium with formula K Al2(Al Si3 O10)( F,O H)2, or ( KF)2( Al2O3)3( SiO2)6( H2O). It has a highly perfect basal cleavag ...
, very small shells and shell fragments, and framboidal
pyrite The mineral pyrite (), or iron pyrite, also known as fool's gold, is an iron sulfide with the chemical formula Fe S2 (iron (II) disulfide). Pyrite is the most abundant sulfide mineral. Pyrite's metallic luster and pale brass-yellow hue giv ...
nodules (0.1–0.3 mm in diameter) are characteristic constituents of this member. ;Pankarp Member The dark mudstones of the Döshult Member are overlain by the Pankarp Member (Swedish ''Pankarpsledet'') with a sharp conglomeratic boundary.Frandsen & Surlyk, 2003, p.550 The late Sinemurian Pankarp Member has an estimated thickness of up to in the subsurface of the Ängelholm, Helsingborg and Landskrona areas. In the Kävlinge area, the thickness is about thick. In westernmost Skåne, the member has been observed in small diameter drill cores. There, the member is subdivided into a lower unit of variegated clays and shales, a middle, poorly sorted silty to sandy unit including a coal bed, and an upper monotonous mudstone unit which is silty and rich in organic matter at the base, and reddish–greenish at the top. It presents different coloration probably due to different degrees of oxidation of iron in the claystone. In the uppermost part of one core, the Pankarp Member comprises lenticular bedded heteroliths with '' Planolites'' burrows. ;Katslösa Member The late Sinemurian to early Pliensbachian Katslösa Member (Swedish ''Katslösaledet'') is mainly known from the subsurface in westernmost Skåne, and it has a thickness of in the Ängelholm, Helsingborg and Landskrona areas. In the Kävlinge area, the thickness is about . Sedimentological interpretations are mainly based on the results of petrographical studies of museum collections. The Katslösa Member yields a rich marine microfauna and macrofauna, and it is dominated by homogeneous mudstone deposited in a marine low-energy environment. Is composed mostly by marine green, brown and dark gray claystones and sandstones. Thin beds of matrix-rich quartz wackes are common. They are typically mineralogically mature but texturally highly immature with abundant angular sand grains. The matrix comprises organic matter,
micrite Micrite is a limestone constituent formed of calcareous particles ranging in diameter up to four μm formed by the recrystallization of lime mud. Flügel, Erik, ''Microfacies of Carbonate Rocks: Analysis, Interpretation and Application,'' Springe ...
, mica and clay minerals. In
thin section In optical mineralogy and petrography, a thin section (or petrographic thin section) is a thin slice of a rock or mineral sample, prepared in a laboratory, for use with a polarizing petrographic microscope, electron microscope and electron ...
, the sandstones show evidence of intense burrowing, which has obliterated depositional structures. Scattered berthierine ooids, as well as authigenic
siderite Siderite is a mineral composed of iron(II) carbonate (FeCO3). It takes its name from the Greek word σίδηρος ''sideros,'' "iron". It is a valuable iron mineral, since it is 48% iron and contains no sulfur or phosphorus. Zinc, magnesium and ...
crystals have been observed in the member.Ahlberg et al., 2003, p.534 ;Rydebäck Member The late Pliensbachian to late Aalenian Rydebäck Member (Swedish ''Rydebäcksledet'') is up to thick in the Ängelholm, Helsingborg and Landskrona areas. It is only known from subsurface material in westernmost Skåne, and sedimentological analysis is based on observations from two wells (Rydebäck–Fortuna-1 and -4). These layers were deposited in small bands during a sea regression, and consist of marine gray, black, green and reddish brown sand and siltstones. The member comprises a uniform succession of muddy
arenite Arenite ( Latin: ''arena'', "sand") is a sedimentary clastic rock with sand grain size between 0.0625 mm (0.00246 in) and 2 mm (0.08 in) and contain less than 15% matrix. The related adjective is ''arenaceous''. The equivalen ...
s with a rich marine microfauna (mostly
foraminifera Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly ...
), and represents deposition in an offshore low-energy environment. The sediments are strongly burrowed, which has caused an effective mixing of sand and mud, resulting in the forming of quartz wackes. The sand is quartz-rich, and grains are typically well rounded. Berthierine
ooid Ooids are small (commonly ≤2 mm in diameter), spheroidal, "coated" (layered) sedimentary grains, usually composed of calcium carbonate, but sometimes made up of iron- or phosphate-based minerals. Ooids usually form on the sea floor, mo ...
s are common constituents of the sediment.


Depositional environments

Tethys Sea transgression entailed formation of fossil-bearing marine deposits in Skane, also associated with an increased tectonic activity. Deposition of the Rya Formation began with nearshore coarse clastics, and continued with offshore mudstones with tempestites (the Döshult Member), followed by offshore muddy sediments with a brief non-marine interval (the Pankarp Member), and ended with deposition of open marine low-energy deposits (Katslösa and Rydebäck Members). The marine Rya Formation shows an overall fining-upwards trend, and an up-section bathymetric deepening of the depositional environment. The depositional environment in western Skåne was either physically protected from the storm energy due to basin topography, or deposition in Skåne took place below storm wavebase. Berthierine ooids occur scattered in the Katslösa Member and are increasingly abundant up-section in the Rydebäck Member. There is a possibility that iron ooid formation was promoted by precipitation of iron and silica from volcanic fluids rising up through the substrate, as has been reported from modern marine sediments offshore Indonesia. This hypothesis has emerged with the publication of age data for the volcanic rocks in Skåne, which now appear to be comparable in age to the prominent iron ooid-bearing deposits, i.e. the Rydebäck Member and the Röddinge Formation.


Age

Based on
foraminifer Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly an ...
s, ammonites and
ostracod Ostracods, or ostracodes, are a class of the Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 70,000 species (only 13,000 of which are extant) have been identified, grouped into several orders. They are small crustaceans, typi ...
s, the Döshult Member is dated to the early Sinemurian, the Pankarp Member to the late Sinemurian, the Katslösa Member to the late Sinemurian to early
Pliensbachian The Pliensbachian is an age (geology), age of the geologic timescale and stage (stratigraphy), stage in the stratigraphic column. It is part of the Early Jurassic, Early or Lower Jurassic epoch (geology), Epoch or series (stratigraphy), Series an ...
and the Rydebäck Member to the late Pliensbachian to late
Aalenian The Aalenian () is a subdivision of the Middle Jurassic Epoch/Series of the geologic timescale that extends from about 174.1 Ma to about 170.3 Ma (million years ago). It was preceded by the Toarcian and succeeded by the Bajocian. Stratigraphi ...
. The formation is time-equivalent with the Röddinge Formation of the Vomb Trough, the
Djupadal Formation The Djupadal Formation is a Formation (geology), geologic formation in Skåne County, southern Sweden. It is Early Jurassic (probably Pliensbachian-Toarcian, or Late Toarcian) in age. It is part of the Central Skåne Volcanic Province, know by the ...
in central Skane and the
Sorthat Formation The Sorthat Formation is a geologic formation on the island of Bornholm, Denmark and in the Rønne Graben in the Baltic Sea. It is of Latest Pliensbachian to Late Toarcian age. Plant fossils have been recovered from the formation, along with sev ...
of Denmark, with which it shares the '' Spheripollenites''–'' Leptolepidites'' and '' Callialasporites''–'' Perinopollenites'' Zones.Nielsen et al., 2003, p.588Nielsen et al., 2003, p.590 The formation also correlates with the Fjerritslev Formation of the
Danish Basin Danish may refer to: * Something of, from, or related to the country of Denmark People * A national or citizen of Denmark, also called a "Dane," see Demographics of Denmark * Culture of Denmark * Danish people or Danes, people with a Danish ance ...
,Frandsen & Surlyk, 2003, p.543 and the Gassum Formation of the Øresund Basin.Erlström et al., 2018, p.138 The storm-dominated, hummocky cross-stratified
Hasle Formation The Hasle Formation is a geologic formation on the island on Bornholm, Denmark. It is of early to late Pliensbachian age. Vertebrate fossils have been uncovered from this formation. The type section of the formation is found at the south of the c ...
on Bornholm is contemporaneous with the muddy Katslösa Member of the Rya Formation.


Basin history

;Basement The basins where the Rya Formation was deposited form part of the Sorgenfrei-Tornquist Zone (STZ) of the
Trans-European Suture Zone The Trans-European Suture Zone (TESZ), also known as the Tornquist Zone, is the crustal boundary between the Precambrian East European Craton and the Phanerozoic orogens of South-Western Europe. The zone runs from the North Sea to the Black Sea. ...
, the boundary between
Baltica Baltica is a paleocontinent that formed in the Paleoproterozoic and now constitutes northwestern Eurasia, or Europe north of the Trans-European Suture Zone and west of the Ural Mountains. The thick core of Baltica, the East European Craton, ...
to the northeast and Peri-Gondwana to the southwest. The orogeny was active in the
Late Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. ...
, or approximately 445 million years ago. At the Carboniferous-
Permian The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last ...
boundary around 300 Ma, the area was influenced by the Skagerrak-Centered Large Igneous Province, another large igneous province stretching across the North Sea, the eponymous
Skagerrak The Skagerrak (, , ) is a strait running between the Jutland peninsula of Denmark, the southeast coast of Norway and the west coast of Sweden, connecting the North Sea and the Kattegat sea area through the Danish Straits to the Baltic Sea. T ...
between Denmark and Sweden and to the northwest up to northern England and Scotland. ;Break-up of Pangea The basins of southern Sweden and eastern Denmark were formed during the latest Triassic and earliest Jurassic. During this time the
Central Atlantic magmatic province The Central Atlantic magmatic province (CAMP) is the Earth's largest continental large igneous province, covering an area of roughly 11 million km2. It is composed mainly of basalt that formed before Pangaea broke up in the Mesozoic Era, near the ...
(CAMP), with an estimated the largest igneous province in Earth's history, was formed to the present southwest of the Danish-Swedish realm. In the Skåne area, the
Central Skåne Volcanic Province The Central Skåne Volcanic Province was a site of volcanic activity in the Scania region of Sweden during the Mesozoic Era of the Earth's geological history. The volcanism began with a first and main phase in late Sinemurian to Toarcian times aro ...
was active during the time of deposition of the Rya Formation, commencing around the Sinemurian-Pliensbachian boundary. The earliest magmatism was partly emplaced into and across pre-existing extensional basin structures.Bryan & Ferrari, 2013, p.1058 The first and the main volcanic phase of this volcanic province occurred in the Early Jurassic (late Sinemurian to Toarcian) at 191–178 Ma.Bergelin, 2009 Analysis of the volcanic rocks produced by this Jurassic volcanism suggests a continental Strombolian-type eruptive character close to the oceans of the Early Jurassic.Augustsson, 2009 No correlative pyroclastic beds have yet been identified in sedimentary basins surrounding central Skåne.Ahlberg et al., 2003, p.539 ;Toarcian During deposition of the Rydebäck Member, the
Toarcian turnover The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period o ...
happened. This event at the
Pliensbachian The Pliensbachian is an age (geology), age of the geologic timescale and stage (stratigraphy), stage in the stratigraphic column. It is part of the Early Jurassic, Early or Lower Jurassic epoch (geology), Epoch or series (stratigraphy), Series an ...
-
Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toarcian ...
boundary characterized by widespread anoxic conditions globally, led to the extinction of various groups of flora and fauna. Taxa inhabiting the upper water column were unaffected by anoxia and included ammonites and belemnites. Epifaunal taxa adapted to low-oxygen conditions, such as the buchiids, posidoniids and inoceramids, flourished in the post-extinction environment during the survival interval.Harries & Little, 1999


Economic geology

A study on the geothermal potential of reservoirs in the Øresund Basin published in 2018 by Erlström et al. gave results of the formation together with the Gassum and Höganäs Formations, giving the following characteristics of the three Early Jurassic formations:Erlström et al., 2018, p.139 * Net sand thickness - *
Porosity Porosity or void fraction is a measure of the void (i.e. "empty") spaces in a material, and is a fraction of the volume of voids over the total volume, between 0 and 1, or as a percentage between 0% and 100%. Strictly speaking, some tests measur ...
- 18 to 34% * Permeability - 50 to 1500 mD * Cl concentration - 120 to 190 gram/liter * Productivity index - 7.0 m3/hr/bar A study published in the same year analyzing the CO2 storage potential of the Rya and Höganäs Formations concluded a storage capacity of 543 megatons of carbon dioxide.Sjöberg, 2018, p.90 The organic content of the Jurassic strata in Skåne is typically dominated by gas-prone
kerogen Kerogen is solid, insoluble organic matter in sedimentary rocks. Comprising an estimated 1016 tons of carbon, it is the most abundant source of organic compounds on earth, exceeding the total organic content of living matter 10,000-fold. It ...
(type III), which is below, or at the onset of, thermal maturity.


Paleoenvironment

The sedimentological evolution of the Jurassic in southwestern Skåne, specially the Rya Formation, has provided depth diverse data about its different environments, specially on those samples recovered on the Höllviken-2 core and sidewall cores from the FFC-1 well. The unit was linked on all its sedimentological history with the main Fennoscandinavian land. The local succession in the Höllviken Halfgraben and Barsebäck Platform has measured continue during most of the
Triassic The Triassic ( ) is a geologic period and system (stratigraphy), system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago (Year#Abbreviations yr and ya, Mya), to the beginning of the Jurassic Period 201.36 ...
and
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
times. Inte western part of the Höllviken Halfgraben during the Hettangian
Pliensbachian The Pliensbachian is an age (geology), age of the geologic timescale and stage (stratigraphy), stage in the stratigraphic column. It is part of the Early Jurassic, Early or Lower Jurassic epoch (geology), Epoch or series (stratigraphy), Series an ...
succession there is evidence of higher subsidence rate along the Öresund Fault, indicating tectonically controlled deposition and the presence of a submarine high at the west of this fault. In the east is the Skarup Platform, interpreted as a Jurassic high based on its incomplete or missing Rhaetian–Lower Jurassic strata, along redeposited spores and pollen in the Höllviken Halfgraben and increased sand output, that point to this zone as the major freshwater terrestrial source. This deposits shows several nearshore environments, from offshore marine environment probably below wave base on the sinemurian level, as proven by the presence of well cemented fine sand and 2 m thick clay dominated heterolites, that change to more marine influenced flaser bedded heterolites with some strongly bioturbated horizons and convolute beds in the Pliensbachian, and end in the Pliensbachian–Lower Toarcian with a sandy seashore setting with high energy lenticular beds. Some sections are suggested to be tidal flat zones and/or distal bar deltaic deposits. Coeval with the Rydebäck member, at Anholt ( Fjerritslev Formation) where studied several levels that point to a connected fluvial system.Seidenkrantz & Koppelhus (1993)-p201 Concretely the Upper Pliensbachian-Toarcian boundary marks the beginning of a major regression, which continued through the Toarcian and Aalenian. The discovery of the foraminifera '' Eoguttuliiia liassica'', Bony Fishes and
Scolecodont A scolecodont is the jaw of a polychaete annelid, a common type of fossil-producing segmented worm useful in invertebrate paleontology. Scolecodonts are common and diverse microfossils, which range from the Cambrian period (around half a billion ...
s, points that the Pliens-Toar boundary has a shallow water environment, possibly of reduced salinity.Seidenkrantz & Koppelhus (1993)-p210 The Pollen and spores, identical of those found on the Rydebäck member (Vilhelmsfält Bore No. 1 and Karindal bore no. 1), increase, and the dinoflagellate cysts decrease, suggesting more proximal shoreline. This layers, on both Anholt and the Rydebäck member, are considered to represent a regression from marine inner shelf environments to near lagoonal or deltaic conditions during the late Pliensbachian and early Toarcian, but with light marine influence, as show the presence of ''
Dactylioceras ''Dactylioceras'' was a widespread genus of ammonites from the Lower Jurassic period, approximately 180 million years ago ( mya). Etymology The name ''Dactylioceras'' comes from the Greek ''dactyl'', meaning “finger”, and refers to the shell ...
'' on the Rydebäck member. The sequence points concretely than in both units The Late Pliensbachian sediments at were deposited in a storm-influenced Inner Shelf setting, that changed on the Lower Toarcian into a more restricted, marginal-marine conditions with delta progradation.Nagy, J. (2003)-p28 Then in the late Toarcian increased brackish conditions, to end on a short marine encroachment during the Early Aalenian. The study of the Jurassic sediments has allowed to know that the basement rocks of southern Sweden were deeply weathered in Late
Triassic The Triassic ( ) is a geologic period and system (stratigraphy), system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago (Year#Abbreviations yr and ya, Mya), to the beginning of the Jurassic Period 201.36 ...
-
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
times, with formed saprolites on the sub-mesozoic basement and high amount of Kaolinite on the stronger weathered profiles, opposed to
smectite A smectite (from ancient Greek ''σμηκτός'' smektos 'lubricated'; ''σμηκτρίς'' smektris 'walker's earth', 'fuller's earth'; rubbing earth; earth that has the property of cleaning) is a mineral mixtures of various swelling sheet sil ...
on the less weathered ones.Ahlberg, Olsson & Šimkevičius, 2003-p15 Thus, fluvial currents released smectite-rich weathering material to the Late Triassic–Jurassic receiving basins. The members of the Rya Formation recover a transition from deltaic to parallic coast and shallow marine, dominated by kaolinite, along with peaks of illite and smectite.Ahlberg, Olsson & Šimkevičius, 2003-p17 The record of this minerals showed that at the time of deposition of the Rya Formation, mid-latitude warmth and pronounced humidity to drier pseudomediterranean climates allowed on the denuded bedrock in the
Fennoscandian Shield The Baltic Shield (or Fennoscandian Shield) is a segment of the Earth's crust belonging to the East European Craton, representing a large part of Fennoscandia, northwestern Russia and the northern Baltic Sea. It is composed mostly of Archean an ...
, composed of
Gneiss Gneiss ( ) is a common and widely distributed type of metamorphic rock. It is formed by high-temperature and high-pressure metamorphic processes acting on formations composed of igneous or sedimentary rocks. Gneiss forms at higher temperatures a ...
es or
Granite Granite () is a coarse-grained ( phaneritic) intrusive igneous rock composed mostly of quartz, alkali feldspar, and plagioclase. It forms from magma with a high content of silica and alkali metal oxides that slowly cools and solidifies under ...
s, at places intersected by
Dolerite Diabase (), also called dolerite () or microgabbro, is a mafic, holocrystalline, subvolcanic rock equivalent to volcanic basalt or plutonic gabbro. Diabase dikes and sills are typically shallow intrusive bodies and often exhibit fine-grain ...
dykes, fracturation and active erosion/weathering.Ahlberg, Olsson & Šimkevičius, 2003-p19 The coeval
Djupadal Formation The Djupadal Formation is a Formation (geology), geologic formation in Skåne County, southern Sweden. It is Early Jurassic (probably Pliensbachian-Toarcian, or Late Toarcian) in age. It is part of the Central Skåne Volcanic Province, know by the ...
volcanic ash falls at the east Skarup Platform-Cenntral Skane Volcanic Province, may have diluted the marine sediments, with raised smectite content. The pronounced mineralogical maturity of most Swedish post-Norian Mesozoic arenites confirms widespread feldspar destruction in the weathering profiles of the Paleozoic crystalline basement at the north, as a consequence of the increased humidity.Ahlberg, Olsson & Šimkevičius, 2003-p21 A fish tooth recovered from the lower Toarcian of the North West German Basin presents a radiogenic seawater value of −6 ε-units, which is quite counter-intuitive to the idea of massive unradiogenic crustal-derived inputs from Laurasia. Clastic fractions found on the same layer suggest brief radiogenic Nd influxes from the Skåne flood basalts erupted at this time. ;Pliensbachian Lower-late Pliensbachian Layers at Katslösa are similar in faunal composition. The Katslösa Member composition suggest deposition on a Low Energy Marginal marine environment with absence of changes in the salinity (As proven by the presence of Echinoderms and other low salt intolerant fauna), with active bottoms filled by traces of diverse invertebrates and abundance of Bivalves. The Upper Pliensbachian section that mark the appearance of the Rydebäck Member continues the stable salinity environment with increased presence of Echinoderms, but lack of a diverse Bivalve fauna as in the older section. This unit was deposited likely on a marine setting more proximal to the shore and low-energy, more strongly burrowed than older layers. The presence of Selachian fauna can be more likely derived from a more suitable depositional setting for this remains than a biota turnover. Palynology in this section is dominated by spores in the Karindal bore no. 1, suggesting a humid climate on nearby emerged lands. ;Toarcian Toarcian layers of the formation where influenced by the ongoing vulcanism located in the
Central Skåne Volcanic Province The Central Skåne Volcanic Province was a site of volcanic activity in the Scania region of Sweden during the Mesozoic Era of the Earth's geological history. The volcanism began with a first and main phase in late Sinemurian to Toarcian times aro ...
(
Djupadal Formation The Djupadal Formation is a Formation (geology), geologic formation in Skåne County, southern Sweden. It is Early Jurassic (probably Pliensbachian-Toarcian, or Late Toarcian) in age. It is part of the Central Skåne Volcanic Province, know by the ...
), as, marine water influence is observed in the main outcrop of the last unit, where enriched
Zeolite Zeolites are microporous, crystalline aluminosilicate materials commonly used as commercial adsorbents and catalysts. They mainly consist of silicon, aluminium, oxygen, and have the general formula ・y where is either a metal ion or H+. These p ...
by Barium is suggested to derive from oceanic water, that may have circulated as hydrothermal flows in the lapilli tuff and facilitating diagenetic changes. Is also known by palynological analysis on the Bonnarp Cone, where saltwater/brackish acritarchs like '' Leiosphaera'' and '' Leiofusa'' or Dinoflajellates like '' Nannoceratopsis'' where recovered. This section also recovers the increased influence of Terrestrial weathering measured in the layers after the Toarcian AOE, as seen in the Vilhelmsfalt borehole, where plant remains increase their presence, suggesting a regression of the coast influenced by the increased volcanic-derived materials. The Palynology in this section is dominated by the Pollen of '' Chasmatosporites'' (Cycads), with at least six species recovered, playing more than 50% of the total palynological samples, implicating a clear dominant role of the producer of this Pollen and suggesting along the increased amount of
Cheirolepidiaceae Cheirolepidiaceae is an extinct family of conifers. They first appeared in the Triassic, and were widespread during most of the Mesozoic era. They are united by the possession of a distinctive pollen type assigned to the form genus '' Classopolli ...
pollen and general decrease of spores a shif towards a more arid climate on nearby settings.


Fossil content

The formation has provided fossils of typically marine fauna. With the exception of a continental coal bed, the formation is marine in character. Shark teeth were reported from the Rydebäck Member.Rya Formation
at Fossilworks.org
Apart from a few teeth of the hybodont '' Hybodus reticulatus'', the shark fauna from the Rya Formation is exclusively neoselachian.Rees, 2000, p.411 ;Fish ;Annelida * ''
Serpula ''Serpula'' (also known as calcareous tubeworm, serpulid tubeworm, fanworm, or plume worm) is a genus of sessile, marine annelid tube worms that belongs to the family Serpulidae. Serpulid worms are very similar to tube worms of the closely re ...
terquemi''Troedsson (1951)-p146-147 * ''
Serpula ''Serpula'' (also known as calcareous tubeworm, serpulid tubeworm, fanworm, or plume worm) is a genus of sessile, marine annelid tube worms that belongs to the family Serpulidae. Serpulid worms are very similar to tube worms of the closely re ...
quinquesulcata'' * ''
Serpula ''Serpula'' (also known as calcareous tubeworm, serpulid tubeworm, fanworm, or plume worm) is a genus of sessile, marine annelid tube worms that belongs to the family Serpulidae. Serpulid worms are very similar to tube worms of the closely re ...
cf. raricostati'' * '' Eklexibella johanni'' ;Echinodermata * '' Isocrinus ranae'' sp. nov.Hunter, A.W. & Rees, J.(2010)-p10-24 * '' Pentacrinus basaltiformis'' * '' Balanocrinus subteroides''Troedsson (1951)-p15 * '' Hispidocrinus scalaris'' * ''? Acrosaleniidae Indeterminate'' ;Ammonites * '' Coroniceras rejnesi'' * '' Megarietites meridionalis'' * '' Paracoroniceras crossi'' * '' Agassiceras nodulatum'' * '' Arnioceras cf. falcaries'' * '' Arnioceras sp. indet.'' * '' Eparietites sp. indet.'' * '' Cymbites striaries'' * ''
Asteroceras obtusum ''Asteroceras'' is an extinct genus of cephalopod belonging to the Ammonite subclass. These fast-moving nektonic carnivores lived during the Triassic and Jurassic periods (from 205.6 to 189.6 Ma). Species *''Asteroceras blakei'' Spath, 1925 *'' ...
''Frandsen & Surlyk, 2003, p.545 * '' Oxynoticeras oxynotum'' * '' Oxynoticeras? sp. indet.'' * '' Promicroceras planicostatum'' * '' Promicroceras sp. juv.'' * '' Euagassiceras resupinatum'' * '' Euagassiceras spinaries'' * '' Euagassiceras lundgreni'' * '' Euagassiceras cf. lundgreni'' * '' Uptonia jamesoni''Sivhed, U. 1984-p19 * '' Polymorphites angustus'' * '' Amaltheus margaritatus'' * '' Pleuroceras spinatum'' * '' Pleuroceras hawskerense'' * '' Amauroceras ferrugineum'' * '' Echioceras raricostatum''Sivhed, U. (1984)- p.251 * '' Tragophylloceras ibex'' * '' Prodactylioceras davoei'' * '' Dactylioceras tenuicostatum'' * '' Dactylioceras cf. semicelatum'' * '' Eleganticeras elegantulum'' ;Belemnites * '' Passaloteuthis alveolata''Troedsson (1951)-p243-245 * '' Passaloteuthis apicicurvata'' * ''
Passaloteuthis ''Passaloteuthis'' is a genus of belemnite, an extinct group of cephalopods. Belemnites are typically known for having about 40 micro-hooks on each one of its appendage. However, ''Passaloteuthis'' is notable for being associated with a pair of me ...
cf. virgata'' * ?''
Passaloteuthis ''Passaloteuthis'' is a genus of belemnite, an extinct group of cephalopods. Belemnites are typically known for having about 40 micro-hooks on each one of its appendage. However, ''Passaloteuthis'' is notable for being associated with a pair of me ...
sp.'' * '' Pseudohastites charmouthensis'' * ''
Pseudohastites ''Pseudohastites'' is a genus of belemnite, an extinct group of cephalopods. See also * Belemnite * List of belemnites References Belemnites {{paleo-cephalopod-stub ...
cf. arundineus'' * ''" Belemnites" elongatus'' * ''" Belemnites" milleri'' ;Brachiopods * '' Zeilleria perforata''Troedsson (1951)-p148-150 * '' Zeilleria numismalis'' * '' Spiriferina walcotti'' * ''
Rhynchonella ''Rhynchonella'' is an extinct genus of brachiopod found in Ordovician to Eocene strata worldwide. It was a stationary epifaunal suspension feeder. Description These 1.75 to 3.75 cm long articulate brachiopods are characterized by a tria ...
deffneri'' ;Bivalves * '' Chlamys interpunctata'' * '' Chlamys textoria'' * '' Chlamys janiformis'' * '' Chlamys subulata'' * '' Chlamys tullbergi''Troedsson (1951)-p213 * '' Chlamys textaria'' * '' Chlamys interpunctata'' * '' Oxytoma sinemurensis'' * '' Oxytoma inaequivalvis''Troedsson (1951)-p200 * '' Oxytoma scanica'' * '' Astarte sp.'' * '' Astarte angelini''Troedsson (1951)-p167 * '' Astarte fructuum''Troedsson (1951)-p168 * '' Astarte scanensis'' * '' Astarte fortuna'' * '' Astarte deltoidea'' * '' Astarte oerbyensis''Troedsson (1951)-p171 * '' Astarte ryensis'' * '' Tutcheria cingulata'' * '' Tutcheria cf. rickardsoni''Troedsson (1951)-p173 * '' Pseudopis sp.'' * ' * '' Entolium sp.'' * '' Homomya sp.'' * '' Leda sp.'' * '' Liogryphaeaarcuata sp.'' * '' Nucula sp.'' * '' Nucula distinguenda''Troedsson (1951)-p154 * '' Nuculana sp.'' * '' Nuculana zieteni'' * '' Nuculana doris'' * '' Palaeoneilo sp.'' * '' Palaeoneilo galatea''Troedsson (1951)-p149 * '' Palaeoneilo bornholmiensis''Troedsson (1951)-p150 * '' Palaeoneilo oviformis''Troedsson (1951)-p151 * '' Pleuromya sp.'' * '' Rollieria sp.'' * '' Rollieria bronni'' * '' Grammatodon cypriniformis'' * '' Grammatodon sinuatus'' * '' Grammatodon subrhomboidalis'' * '' Barbatia pulla'' * '' Cardinia tollini'' * '' Cardinia expansa'' * '' Cardinia ingelensis'' * '' Cardinia kullensis'' * '' Trigonia primaeva'' * '' Trigonia modesta'' * '' Tancredia arenacea'' * '' Tancredia securiformis'' * '' Tancredia erdmanni''Troedsson (1951)-p176 * '' Tancredia johnstrupi'' * '' Tancredia lineata'' * '' Sphaeriola kurremolinae''Troedsson (1951)-p178 * '' Protocardia philippiana'' * '' Protocardia oxynoti'' * '' Protocardia truncata'' * '' Eotrapezium gerrnari''Troedsson (1951)-p181 * '' Eotrapezium pullastra'' * '' Eotrapezium heberti'' * '' Eotrapezium menkei''Troedsson (1951)-p185 * '' Anisocardia luggudensis'' * '' Platymya aquarum'' * '' Homomya ovalis'' * '' Homomya venulithus'' * '' Homomya centralis'' * '' Homomya librata''Troedsson (1951)-p192 * '' Arcomya decora'' * '' Arcomya cf. elongata''Troedsson (1951)-p193 * '' Pleuromya forchhammeri'' * '' Pleuromya corrugata'' * '' Goniomya heteropleura'' * '' Gervillia angelini'' * '' Gervillia scanica'' * '' Gervillia hagenowi'' * '' Gervillia sjögreni'' * '' Isognomon sp. ''Troedsson (1951)-p207 * '' Lima duplicata'' * '' Lima pectinoides''Troedsson (1951)-p208 * '' Plagiostoma succincta'' * '' Limea acuticostata''Troedsson (1951)-p210 * '' Limea katsloesensis'' * '' Entolium hehli''Troedsson (1951)-p216 * '' Entolium calvum'' * '' Entolium cingulatum'' * '' Entolium lundgreni'' * '' Pseudopecten aequivalvis'' * '' Dimyodon sp.''Troedsson (1951)-p220 * '' Plicatula spinosa'' * '' Plicatula orbiculoides'' * '' Terquemia arietis'' * '' Anomia pellucida'' * '' Liastrea hisingeri'' * '' Liogryphaea arcuata'' * '' Liogryphaea lata'' * '' Lioqryphaea regularis''Troedsson (1951)-p228 * '' Modiola hillana'' * '' Modiola ruuthi'' * '' Modiola cf. tenuissima''Troedsson (1951)-p232 * '' Modiola scalprum'' * '' Mytilus cf. lamellosus'' * '' Myoconcha decorata''Troedsson (1951)-p234 * '' Taeniodon nathorsti'' ;Gasteropods * '' Actaeonina nathorsti''Troedsson (1951)-p238-240 * '' Actaeonina cf. striata'' * '' Chrysostoma cf. solarium'' * '' Chemnitzia sp.'' * ''
Trochus ''Trochus'' is a genus of medium-sized to large, top-shaped sea snails with an operculum and a pearly inside to their shells, marine gastropod molluscs in the subfamily Trochinae of the family Trochidae, the top snails.Bouchet, P.; Gofas, S. ( ...
cf. imbricatus'' * ''
Trochus ''Trochus'' is a genus of medium-sized to large, top-shaped sea snails with an operculum and a pearly inside to their shells, marine gastropod molluscs in the subfamily Trochinae of the family Trochidae, the top snails.Bouchet, P.; Gofas, S. ( ...
laevis'' * '' Ptychomphalus cf. expansus'' * '' Kalchreuthia frankei'' ;Scaphopods * ''" Dentalium" hexagonale'' * '' Prodentalium bandeli'' * ''
Laevidentalium ''Laevidentalium'' is a genus of molluscs belonging to the family Laevidentaliidae. The genus has cosmopolitan distribution In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all or most of th ...
sp.'' * ''
Laevidentalium ''Laevidentalium'' is a genus of molluscs belonging to the family Laevidentaliidae. The genus has cosmopolitan distribution In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all or most of th ...
elongatum''Troedsson (1951)-p230-245 * '' Progadilina spaethi'' * '' Progadilina subtrigonalis'' * '' Baltodentalium weitschati'' ;Ostracods * '' Citharina inaequistriata'' * '' Marginulina spinata spinata'' * '' Astacolus denticulina carinata'' * '' Brizalina liassica amalthea'' * '' Ogmoconchella danica'' * '' Ogmoconchella mouhersensis'' * '' Ogmoconchella adenticulata'' * '' Pleurifera harpa'' * '' Ogmoconcha amalthei amalthei'' * '' Cristacythere crassireticulata'' * '' Nanacythere simplex'' ;Mites * '' Hydrozetes nsp.''Sivhed & Wallwork (1978)-p65-66 ;Ichnofossils * '' Chondrites sp.''Frandsen & Surlyk, 2003, p.546 * '' Rhizocorallium sp.'' * '' Diplocraterion sp.'' * '' Planolites sp.'' * '' Skolithos sp.'' * '' Teichichnus sp.''


Plant Remains

Coalified wood occurs as scattered pieces up to long and indeterminate
belemnite Belemnitida (or the belemnite) is an extinct order of squid-like cephalopods that existed from the Late Triassic to Late Cretaceous. Unlike squid, belemnites had an internal skeleton that made up the cone. The parts are, from the arms-most ...
s,
echinoid Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) of ...
s,
serpulid The Serpulidae are a family of sessile, tube-building annelid worms in the class Polychaeta. The members of this family differ from other sabellid tube worms in that they have a specialized operculum that blocks the entrance of their tubes wh ...
s,
ostracod Ostracods, or ostracodes, are a class of the Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 70,000 species (only 13,000 of which are extant) have been identified, grouped into several orders. They are small crustaceans, typi ...
s and nodosariid
foraminifera Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly ...
were also recorded in the formation.Frandsen & Surlyk, 2003, p.547 Ammonites where found on layers with plant fossils. Fragments of ''
Dactylioceras ''Dactylioceras'' was a widespread genus of ammonites from the Lower Jurassic period, approximately 180 million years ago ( mya). Etymology The name ''Dactylioceras'' comes from the Greek ''dactyl'', meaning “finger”, and refers to the shell ...
'' were found at a depth of 170 m in the Vilhelmsfalt borehole, together with plant remains and beautifully preserved
Pelecypod Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, biva ...
s.Reyment (1959)-p152 The plant material, probably derived from watercourses emptying in the neighborhood, consists mostly of small fragments that seem to have been intimately sedimented with the muddy material that flocculated on meeting the seawater. Other specimen, '' Arnioceras'' sp. indet. occurs on what appears to be a transitional environment, probably a back-beach with coalified fragments of plants, some of the large. Probably due to the shell being washed to land due to a storm. ;Palynology ;Plant Macrofossils


See also

*
List of fossiliferous stratigraphic units in Sweden See also * Lists of fossiliferous stratigraphic units in Europe These lists of fossiliferous stratigraphic units in Europe enumerate the rock layers which preserve the fossilized remains of ancient life in Europe by the modern countries wher ...
*
Kristianstad Basin The Kristianstad Basin (Swedish: ''Kristianstadsbassängen'') is a Cretaceous-age structural basin and geological formation in northeastern Skåne, the southernmost province of Sweden. The basin extends from Hanöbukten, a bay in the Baltic Sea, ...
* Toarcian formations *
Djupadal Formation The Djupadal Formation is a Formation (geology), geologic formation in Skåne County, southern Sweden. It is Early Jurassic (probably Pliensbachian-Toarcian, or Late Toarcian) in age. It is part of the Central Skåne Volcanic Province, know by the ...
, Central Skane * Marne di Monte Serrone, Italy * Calcare di Sogno, Italy * Mizur Formation, North Caucasus * Sachrang Formation, Austria *
Saubach Formation The Saubach Formation is a geological formation in Austria and Germany, dating to about 180–174 mya (unit), million years ago.Böhm, F. (2003). Lithostratigraphy of the Adnet Group (Lower to Middle Jurassic, Salzburg, Austria) (pp. 231–268). V ...
, Austria *
Posidonia Shale The Posidonia Shale (german: Posidonienschiefer, also called Schistes Bitumineux in Luxembourg) geologically known as the Sachrang Formation, is an Early Jurassic (Toarcian) geological formation of southwestern and northeast Germany, northern Swit ...
,
Lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These f ...
in Germany *
Ciechocinek Formation The Ciechocinek Formation (also known as the Gryfice Formation at Suliszewo), formerly known in Germany as the Green Series (German: ''Grüne Serie'') is a Jurassic (lower Toarcian) geologic formation that extends across the Baltic coast, from ...
, Germany and Poland * Krempachy Marl Formation, Poland and Slovakia * Lava Formation, Lithuania


References


Bibliography

* * * * * * * * * * * E- * * * * * * * * * * * * * * * * * * * * * * * * * *


External links

{{Mesozoic Sweden, state=expanded Geologic formations of Denmark Geologic formations of Sweden Jurassic System of Europe Early Jurassic Europe Middle Jurassic Europe Jurassic Denmark Jurassic Sweden Sinemurian Stage Pliensbachian Stage Toarcian Stage Aalenian Stage Siltstone formations Shale formations Sandstone formations Mudstone formations Coal formations Coal in Sweden Open marine deposits Shallow marine deposits Paleontology in Sweden Formations Denmark–Sweden border