Rufous-capped Thornbill
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The rufous-capped thornbill (''Chalcostigma ruficeps'') is a species of
hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics aro ...
in the "coquettes", tribe
Lesbiini Lesbiini is one of the two tribes that make up the subfamily Lesbiinae in the hummingbird family Trochilidae. The other tribe is Heliantheini (brilliants). The informal name "coquettes" has been proposed for this group as the largest genus, ''L ...
of subfamily
Lesbiinae Lesbiinae is one of the six subfamily, subfamilies that make up the hummingbird family (biology), family Trochilidae. The subfamily is divided into two Tribe (biology), tribes: Heliantheini ("brilliants") containing 14 genera and Lesbiini ("coqu ...
. It is found in
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
, and
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
.HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved 27 May 2021


Taxonomy and systematics

The taxonomy of genus ''Chalcostigma'' and of the rufous-capped thornbill in particular are unsettled. The South American Classification Committee (SACC) of the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its m ...
is considering a proposal to merge the genus into '' Oxypogon''. The species was for a time placed in genus ''Metallura'' with the Perijá metaltail (''M. iracunda'') and also has been assigned its own genus, ''Selatopogon''.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 31 January 2022. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved February 1, 2022Heindl, M. and P. F. D. Boesman (2020). Rufous-capped Thornbill (''Chalcostigma ruficeps''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.ructho1.01 retrieved February 23, 2022 The rufous-capped thornbill is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
.


Description

The rufous-capped thornbill is about long and weighs . Both sexes have a very short black bill. The male's crown is mahogany red and the rest of the upperparts bottle green. It has a narrow emerald green
gorget A gorget , from the French ' meaning throat, was a band of linen wrapped around a woman's neck and head in the medieval period or the lower part of a simple chaperon hood. The term later described a steel or leather collar to protect the thro ...
with a border of various shades of yellow. Its underparts are rufous with green mottling that is strongest on the flanks. The tail is dull olive green. The female's upperparts including the crown are bottle green. Its throat and breast are ochre-orange with olive green dots. Its outer tail feathers have pale tips.


Distribution and habitat

The rufous-capped thornbill is found on the east slope of the Andes from southeastern Ecuador's
Zamora-Chinchipe Province Zamora Chinchipe (), Province of Zamora Chinchipe is a province of the Republic of Ecuador, located at the southeastern end of the Amazon Basin, which shares borders with the Ecuadorian provinces of Azuay and Morona Santiago to the north, Loja a ...
through Peru into Bolivia as far as
Cochabamba Department Cochabamba ( ay, Quchapampa Jach'a Suyu, es, Departamento de Cochabamba , qu, Quchapampa Suyu), from Quechua ''qucha'' or ''qhucha'', meaning "lake", ''pampa'' meaning "plain", is one of the nine departments of Bolivia. It is known to be the ...
. It has also been recorded as a vagrant in Colombia.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 31 January 2022. Species Lists of Birds for South American Countries and Territories. https://www.museum.lsu.edu/~Remsen/SACCCountryLists.htm retrieved February 1, 2022 It inhabits a variety of
montane Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
landscapes including humid
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has re-grown after a timber harvest or clearing for agriculture, until a long enough period has passed so that the effects of the disturbance are no longer evident. ...
, humid
montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
, and
cloudforest A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest (TMCF), is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud c ...
. In elevation it mostly ranges between but occurs as high as ; it is most common around .


Behavior


Movement

The rufous-capped thornbill possibly makes seasonal elevational movements.


Feeding

The rufous-capped thornbill feeds on nectar from a variety of flowers, mostly in the family
Melastomataceae Melastomataceae is a family of dicotyledonous flowering plants found mostly in the tropics (two-thirds of the genera are from the New World tropics) comprising c. 175 genera and c. 5115 known species. Melastomes are annual or perennial herbs, sh ...
. It usually nectars by clinging to the flower and rarely by hovering. It sometimes "robs" nectar by piercing the base of a flower or by feeding at holes made by '' Diglossa'' flowerpiercers. It forages from low to medium level, and males defend feeding territories. It also feeds on insects but its usual method of capture has not been determined.


Breeding

The rufous-capped thornbill's breeding season spans from December to March in Bolivia and perhaps July to October in Peru. The female alone incubates the two white eggs. Nothing else has been documented about its breeding
phenology Phenology is the study of periodic events in biological life cycles and how these are influenced by seasonality, seasonal and interannual variations in climate, as well as environmental factor, habitat factors (such as elevation). Examples includ ...
.


Vocalization

The rufous-capped thornbill's vocalizations are not well known, but do include "a short dry double rattle 'trr-trr'" and "a rising/falling twitter 't-tr-tr-tsee-tseee-tititrrr'."


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed the rufous-capped thornbill as being of Least Concern. It has a large range but its population size is unknown and believed to be decreasing. It is generally uncommon but appears to tolerate some human-caused environmental changes.


References

{{Taxonbar, from=Q1263925 rufous-capped thornbill Birds of the Peruvian Andes Birds of the Bolivian Andes rufous-capped thornbill Taxonomy articles created by Polbot