The New Zealand king shag (''Leucocarbo carunculatus''), also known as the rough-faced shag, king shag or kawau, is a rare bird

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to New Zealand. Some taxonomic authorities, including the

International Ornithologists' Union The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...

, place this species in the genus ''

Leucocarbo ''Leucocarbo'' is a genus of birds in the family Phalacrocoracidae with the members commonly known as blue-eyed shags. This is a group of closely related cormorant taxa. Many have a blue, purple or red ring around the eye (not a blue iris); other ...

''. Others place it in the genus ''

Phalacrocorax ''Phalacrocorax'' is a genus of fish-eating birds in the cormorant family Phalacrocoracidae. Members of this genus are also known as the Old World cormorants. Taxonomy The genus ''Phalacrocorax'' was introduced by the French zoologist Mathurin ...

''.

Taxonomy

The New Zealand king shag was formally described in 1789 by the German naturalist

Johann Friedrich Gmelin , fields = , workplaces = University of GöttingenUniversity of Tübingen , alma_mater = University of Tübingen , doctoral_advisor = Philipp Friedrich GmelinFerdinand Christoph Oetinger , academic_advisors = , doctora ...

in his revised and expanded edition of

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...

's ''

Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...

''. He placed it in the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

'' Pelecanus'' and coined the

binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

''Pelecanus carunculatus''. Gmelin based his description on the "carunculated shag" that had been described in 1785 by the English ornithologist John Latham in his book ''A General Synopsis of Birds ''. Latham had based his description on a specimen in the

Leverian Museum The Leverian collection was a natural history and ethnographic collection assembled by Ashton Lever. It was noted for the content it acquired from the voyages of Captain James Cook. For three decades it was displayed in London, being broken up ...

. The New Zealand king shag is now one of 16 species placed in the genus ''

'' that was introduced in 1856 by the French naturalist

Charles Lucien Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857), was a French naturalist and ornithologist. Lucien and his wife had twelve children, including Cardinal Lucien Bonaparte. Life and career ...

. The name ''Leucocarbo'' combines the

Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...

''leukos'' meaning "white" with the genus name ''Carbo'' introduced by

Bernard Germain de Lacépède Bernard-Germain-Étienne de La Ville-sur-Illon, comte de Lacépède or La Cépède (; 26 December 17566 October 1825) was a French naturalist and an active freemason. He is known for his contribution to the Comte de Buffon's great work, the ...

in 1799. The specific epithet is from

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

''caruncula '' meaning "small piece of flesh". The species is

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...

: no

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

are recognised.

Description

It is a large (76 cm long, 2.5 kg in weight) black and white

cormorant Phalacrocoracidae is a family of approximately 40 species of aquatic birds commonly known as cormorants and shags. Several different classifications of the family have been proposed, but in 2021 the IOC adopted a consensus taxonomy of seven ge ...

with pink feet. White patches on the wings appear as bars when the wings are folded. Yellow-orange swellings ( caruncles) are found above the base of the bill. The grey

gular pouch Gular skin (throat skin), in ornithology, is an area of featherless skin on birds that joins the lower mandible of the beak (or ''bill'') to the bird's neck. Other vertebrate taxa may have a comparable anatomical structure that is referred to as e ...

is reddish in the breeding season. A blue eye-ring indicates its kinship with the other

blue-eyed shag ''Leucocarbo'' is a genus of birds in the family Phalacrocoracidae with the members commonly known as blue-eyed shags. This is a group of closely related cormorant taxa. Many have a blue, purple or red ring around the eye (not a blue iris); othe ...

Distribution and habitat

New Zealand king shags can be seen from the

Cook Strait ferries Cook or The Cook may refer to: Food preparation * Cooking, the preparation of food * Cook (domestic worker), a household staff member who prepares food * Cook (professional), an individual who prepares food for consumption in the food industry * ...

in Queen Charlotte Sound opposite the beginning of the Tory Channel. They live in the coastal waters of the

Marlborough Sounds The Marlborough Sounds are an extensive network of sea-drowned valleys at the northern end of the South Island of New Zealand. The Marlborough Sounds were created by a combination of land subsidence and rising sea levels. According to Māori m ...

where they are known to breed only on rocky islets at four small sites.

References

External links

Image and Classification at Animal Diversity Web
{{Taxonbar, from=Q25626

New Zealand king shag The New Zealand king shag (''Leucocarbo carunculatus''), also known as the rough-faced shag, king shag or kawau, is a rare bird endemic to New Zealand. Some taxonomic authorities, including the International Ornithologists' Union, place this spe ...

Endemic birds of New Zealand Vulnerable fauna of Oceania Marlborough Sounds