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The roadrunners (genus ''Geococcyx''), also known as chaparral birds or chaparral cocks, are two species of fast-running ground
cuckoo Cuckoos are birds in the Cuculidae family, the sole taxon in the order Cuculiformes . The cuckoo family includes the common or European cuckoo, roadrunners, koels, malkohas, couas, coucals and anis. The coucals and anis are sometimes separ ...
s with long tails and crests. They are found in the
southwestern The points of the compass are a set of horizontal, radially arrayed compass directions (or azimuths) used in navigation and cartography. A compass rose is primarily composed of four cardinal directions—north, east, south, and west—each sepa ...
and south-central United States and
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
, usually in the desert. Although capable of flight, roadrunners generally run away from predators. On the ground, some have been clocked at while a few have also been clocked up to .


Species

The subfamily
Neomorphinae The Neomorphinae are a subfamily of the cuckoo family, Cuculidae. Members of this subfamily are known as New World ground cuckoos, since most are largely terrestrial and native to the Americas. Only ''Dromococcyx'' and ''Tapera'' are more arbor ...
, the New World ground cuckoos, includes 11 species of birds, while the genus ''Geococcyx'' has just two:


Morphology

The roadrunner generally ranges in size from from tail to
beak The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food ...
. The average weight is about . The roadrunner is a large, slender, black-brown and white-streaked ground bird with a distinctive head
crest Crest or CREST may refer to: Buildings *The Crest (Huntington, New York), a historic house in Suffolk County, New York *"The Crest", an alternate name for 63 Wall Street, in Manhattan, New York *Crest Castle (Château Du Crest), Jussy, Switzerla ...
. It has long legs, strong feet, and an oversized dark
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
. The tail is broad with white tips on the three outer tail feathers. The bird has a bare patch of skin behind each eye; this patch is shaded blue anterior to red posterior. The lesser roadrunner is slightly smaller, not as streaky, and has a smaller bill. Both the lesser roadrunner and the greater roadrunner leave behind very distinct "X" track marks appearing as if they are travelling in both directions. Roadrunners and other members of the cuckoo family have
zygodactyl In biology, dactyly is the arrangement of digits (fingers and toes) on the hands, feet, or sometimes wings of a tetrapod animal. It comes from the Greek word δακτυλος (''dáktylos'') = "finger". Sometimes the ending "-dactylia" is used ...
feet. The roadrunner can run at speeds of up to and generally prefer sprinting to flying, though it will fly to escape
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
s. During flight, the short, rounded wings reveal a white crescent in the
primary feathers Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
.


Vocalization

The roadrunner has a slow and descending dove-like "coo". It also makes a rapid, vocalized clattering sound with its beak.


Geographic range

Roadrunners inhabit the
Southwestern United States The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural region of the United States that generally includes Arizona, New Mexico, and adjacent portions of California, Colorado, Ne ...
, to parts of Missouri, Arkansas, and Louisiana, as well as
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
and
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
. They live in
arid A region is arid when it severely lacks available water, to the extent of hindering or preventing the growth and development of plant and animal life. Regions with arid climates tend to lack vegetation and are called xeric or desertic. Most ar ...
lowland or mountainous
shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It m ...
or
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
. They are non-migratory, staying in their breeding area year-round. The greater roadrunner is not currently considered threatened in the US, but is habitat-limited.


Food and foraging habits

The roadrunner is an opportunistic
omnivore An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutr ...
. Its diet normally consists of
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s (such as
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are among what is possibly the most ancient living group of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grasshopp ...
s,
crickets Crickets are orthopteran insects which are related to bush crickets, and, more distantly, to grasshoppers. In older literature, such as Imms,Imms AD, rev. Richards OW & Davies RG (1970) ''A General Textbook of Entomology'' 9th Ed. Methuen 8 ...
,
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s, and
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s), small
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s (such as
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s and
snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...
s, including
rattlesnake Rattlesnakes are venomous snakes that form the genera ''Crotalus'' and ''Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting small anim ...
s),
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
s and other small
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s,
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s (including
tarantula Tarantulas comprise a group of large and often hairy spiders of the family Theraphosidae. , 1,040 species have been identified, with 156 genera. The term "tarantula" is usually used to describe members of the family Theraphosidae, although ...
s),
scorpion Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always end ...
s,
centipede Centipedes (from New Latin , "hundred", and Latin , " foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an ...
s,
snail A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gastro ...
s, small birds (and
nestling Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight s ...
s), eggs, and fruits and seeds like those from
prickly pear cactus ''Opuntia'', commonly called prickly pear or pear cactus, is a genus of flowering plants in the cactus family Cactaceae. Prickly pears are also known as ''tuna'' (fruit), ''sabra'', ''nopal'' (paddle, plural ''nopales'') from the Nahuatl word f ...
es and
sumac Sumac ( or ), also spelled sumach, is any of about 35 species of flowering plants in the genus ''Rhus'' and related genera in the cashew family (Anacardiaceae). Sumacs grow in subtropical and temperate regions throughout the world, including Eas ...
s. The lesser roadrunner eats mainly insects. The roadrunner forages on the ground and, when hunting, usually runs after prey from under cover. It may leap to catch insects, and commonly batters certain prey against the ground. Because of its quickness, the roadrunner is one of the few animals that preys upon rattlesnakes; it is also the only real predator of tarantula hawk wasps.


Behavior and breeding

The roadrunner usually lives alone or in pairs. Breeding pairs are monogamous and mate for life, and pairs may hold a territory all year. During the
courtship display A courtship display is a set of display behaviors in which an animal, usually a male, attempts to attract a mate; the mate exercises choice, so sexual selection acts on the display. These behaviors often include ritualized movement ("dances"), ...
, the male bows, alternately lifting and dropping his wings and spreading his tail. He parades in front of the female with his head high and his tail and wings drooped, and may bring an offering of food. The
reproductive season Seasonal breeders are animal species that successfully mate only during certain times of the year. These times of year allow for the optimization of survival of young due to factors such as ambient temperature, food and water availability, and cha ...
is spring to mid-summer (depending on geographic location and species). The roadrunner's
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
is often composed of sticks, and may sometimes contain leaves, feathers, snakeskins, or dung. It is commonly placed above ground level"Usually 1-3 meters above ground; infrequently higher than 3 meters (Hughes 1996)." in a low tree, bush, or
cactus A cactus (, or less commonly, cactus) is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1750 known species of the order Caryophyllales. The word ''cactus'' derives, through Latin, from the Ancient Greek ...
. Roadrunner
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
are generally white. The greater roadrunner generally lays 2–6 eggs per
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...
, but the lesser roadrunner's clutches are typically smaller. Hatching is asynchronous. Both sexes incubate the nest (with males incubating the nest at night) and feed the hatchlings. For the first one to two weeks after the young hatch, one parent remains at the nest. The young leave the nest at two to three weeks old, foraging with parents for a few days after.


Thermoregulation

During the cold desert night, the roadrunner lowers its body temperature slightly, going into a slight
torpor Torpor is a state of decreased physiological activity in an animal, usually marked by a reduced body temperature and metabolic rate. Torpor enables animals to survive periods of reduced food availability. The term "torpor" can refer to the time ...
to conserve energy. To warm itself during the day, the roadrunner exposes dark patches of skin on its back to the sun.


Indigenous lore

The
Hopi The Hopi are a Native American ethnic group who primarily live on the Hopi Reservation in northeastern Arizona, United States. As of the 2010 census, there are 19,338 Hopi in the country. The Hopi Tribe is a sovereign nation within the Unite ...
and other
Pueblo In the Southwestern United States, Pueblo (capitalized) refers to the Native tribes of Puebloans having fixed-location communities with permanent buildings which also are called pueblos (lowercased). The Spanish explorers of northern New Spain ...
tribes believed that roadrunners were medicine birds and could protect against evil spirits. Their unusual X-shaped footprints are used as sacred symbols to ward off evil in many Pueblo tribes—partially because they invoke the protective power of the roadrunners themselves, and partially because the X shape of the tracks conceals which direction the bird is headed (thus throwing malignant spirits off track.) Stylized roadrunner tracks have been found in the rock art of ancestral Southwestern tribes like the
Anasazi The Ancestral Puebloans, also known as the Anasazi, were an ancient Native American culture that spanned the present-day Four Corners region of the United States, comprising southeastern Utah, northeastern Arizona, northwestern New Mexico, a ...
and Mogollon cultures, as well. Roadrunner feathers were traditionally used to decorate Pueblo cradleboards as spiritual protection for the baby. In Mexican Indian and American Indian tribes, such as the
Pima Pima or PIMA may refer to: People * Pima people, the Akimel O'odham, Indigenous peoples in Arizona (U.S.) and Sonora (Mexico) Places * Pima, Arizona, a town in Graham County * Pima County, Arizona * Pima Canyon, in the Santa Catalina Mountains ...
, it is considered good luck to see a roadrunner. In some Mexican tribes, the bird was considered sacred and never killed, but most Mexican Indians used the meat of the roadrunner as a folk remedy to cure illness or to boost stamina and strength. Indigenous peoples of Central America have developed numerous beliefs about the roadrunner. The Ch’orti’, who call it ''t’unk’u’x'' or ''mu’'', have taboos against harming the bird. The Ch'ol
Maya Maya may refer to: Civilizations * Maya peoples, of southern Mexico and northern Central America ** Maya civilization, the historical civilization of the Maya peoples ** Maya language, the languages of the Maya peoples * Maya (Ethiopia), a populat ...
believe roadrunners to have special powers. It is known to them as ''ajkumtz’u’'', derived from the bird's call that is said to make the listener feel tired. The word for roadrunner in the
O'odham language The O'odham peoples, including the Tohono O'odham, the Pima or Akimel O'odham, and the Hia C-ed O'odham, are indigenous Uto-Aztecan peoples of the Sonoran desert in southern and central Arizona and northern Sonora, united by a common heritage ...
is , which is the name of a transit center in
Tucson, Arizona , "(at the) base of the black ill , nicknames = "The Old Pueblo", "Optics Valley", "America's biggest small town" , image_map = , mapsize = 260px , map_caption = Interactive map ...
.


In media

The roadrunner was made popular by the
Warner Bros. Warner Bros. Entertainment Inc. (commonly known as Warner Bros. or abbreviated as WB) is an American film and entertainment studio headquartered at the Warner Bros. Studios complex in Burbank, California, and a subsidiary of Warner Bros. Di ...
cartoon characters
Wile E. Coyote and the Road Runner Wile E. Coyote and the Road Runner are a duo of cartoon characters from the ''Looney Tunes'' and ''Merrie Melodies'' series of animated cartoons, first appearing in 1949 in the theatrical cartoon short ''Fast and Furry-ous''. In each episode, t ...
, created in 1949, and the subject of a long-running series of theatrical cartoon shorts. In each episode, the cunning, insidious, and constantly hungry Wile E. Coyote repeatedly attempts to catch and subsequently eat the Road Runner, but is never successful. The cartoons led to a misconception that roadrunners say "meep, meep" because the roadrunner in this cartoon series made that sound instead of the aforementioned sound of a real roadrunner. In some shorts, the Road Runner makes a noise while sticking his tongue out at Wile E. Coyote, which resembles its actual call. The cartoons rely on a misconception that a roadrunner is much faster than a coyote. In fact, a coyote's fastest sprinting speed is , which is greater than a roadrunner at .


Citations


General references

* * * * * * * *


Further reading

*


External links


Distribution map of the Greater RoadrunnerGreater Roadrunner soundsVideo clips of roadrunners in action
{{Taxonbar, from=Q1452250 * Articles containing video clips Birds of Mexico