Riojasauridae is an extinct

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

sauropodomorph Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lon ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

s from the

Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...

Period (late

Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by t ...

Norian The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic defi ...

Ages). It contains the

genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...

'' Riojasaurus'' and '' Eucnemesaurus''. The Riojasauridae is considered a stem taxon, and is defined as "the most inclusive clade containing ''Riojasaurus incertus'' but not ''

Plateosaurus engelhardti ''Plateosaurus'' (probably meaning "broad lizard", often mistranslated as "flat lizard") is a genus of plateosaurid dinosaur that lived during the Late Triassic period, around 214 to 204 million years ago, in what is now Central and Northern Euro ...

'', ''

Massospondylus carinatus ''Massospondylus'' ( ; from Greek, (massōn, "longer") and (spondylos, "vertebra")) is a genus of sauropodomorph dinosaur from the Early Jurassic. (Hettangian to Pliensbachian ages, ca. 200–183 million years ago). It was described by Si ...

'', or '' Anchisaurus polyzelus''".

Geologic formation A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics ( lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock exp ...

s containing riojasaurid fossils include the Lower

Elliot Formation The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, south ...

Orange Free State The Orange Free State ( nl, Oranje Vrijstaat; af, Oranje-Vrystaat;) was an independent Boer sovereign republic under British suzerainty in Southern Africa during the second half of the 19th century, which ceased to exist after it was defeat ...

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

(where fossils of ''Eucnemesaurus'' have been found), and the

Los Colorados Formation The Los Colorados Formation is a sedimentary rock formation of the Ischigualasto-Villa Unión Basin, found in the provinces of San Juan and La Rioja in Argentina. The formation dates back to the Norian age of the Late Triassic. The up to thick f ...

, in La Rioja Province,

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

(where fossils of ''Riojasaurus'' have been recovered).

Evolutionary relationships

The Riojasauridae are considered to be sauropodomorphs, but not sauropods themselves. This means that they were generally much smaller than the Sauropods of the

Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...

and

Cretaceous Period The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of t ...

s, and members of this family may not have been obligate

quadruped Quadrupedalism is a form of locomotion where four limbs are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four limbs is said to be a quadruped (from Latin ''quattuor' ...

s, the way more derived Sauropods were. The relationships of this family within other sauropodomorphs has been considered by studies in 2007 and 2020 with slightly different results. The

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

below shows basal sauropodomorph relationships simplified after Yates, 2007. In this analysis, Riojasauridae was recovered as the earliest-diverging lineage of the clade

Massopoda Massopoda is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to Late Cretaceous epochs. It was named by paleontologist Adam M. Yates of the University of the Witwatersrand in 2007. Massopoda is a stem-based taxon, define ...

, and represents a sister group to the node formed by

Massospondylidae Massospondylidae is a family (biology), family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and AntarcticaHellert, Spencer M. "A New Basal Sauropodomorph from The Early Jurassic Hanson Formation of Antarc ...

, the genus ''

Jingshanosaurus ''Jingshanosaurus'' (meaning "Jingshan lizard") is a genus of sauropodomorph dinosaurs from the early Jurassic period 201.3 million years ago that went extinct 199.3 million years ago in the Hettangian Age. Its maximum weight was around 4.3 t ...

'' and the Anchisauria. The following cladogram is from a phylogenetic analysis by Peyre de Fabrègues ''et al.'', 2020. Here, Riojasauridae was placed in a more derived position, lying closer to

Sauropoda Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...

than to

, and appearing as the out group to the clade formed by the genus ''Seitaad'' and the Sauropodiformes. The Massospondylidae and the taxa considered to be related to the family by Yates, are shown to be represent more primitive lineages.

References

{{Taxonbar, from=Q3234495 Massopoda Tetrapod unranked clades