Blyth's hornbill (''Rhyticeros plicatus''), also known as the Papuan hornbill, is a large

hornbill Hornbills (Bucerotidae) are a family (biology), family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a Casque (an ...

inhabiting the

forest canopy In biology, the canopy is the aboveground portion of a plant cropping or crop, formed by the collection of individual plant crowns. In forest ecology, canopy also refers to the upper layer or habitat zone, formed by mature tree crowns an ...

Wallacea Wallacea is a biogeographical designation for a group of mainly Indonesian islands separated by deep-water straits from the Asian and Australian continental shelves. Wallacea includes Sulawesi, the largest island in the group, as well as Lo ...

and

Melanesia Melanesia (, ) is a subregion of Oceania in the southwestern Pacific Ocean. It extends from Indonesia's New Guinea in the west to Fiji in the east, and includes the Arafura Sea. The region includes the four independent countries of Fiji, Va ...

. Its local name in

Tok Pisin Tok Pisin (,Laurie Bauer, 2007, ''The Linguistics Student’s Handbook'', Edinburgh ; Tok Pisin ), often referred to by English speakers as "New Guinea Pidgin" or simply Pidgin, is a creole language spoken throughout Papua New Guinea. It is an ...

is ''kokomo''. Previously, this hornbill was placed in the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

Aceros The rufous-necked hornbill (''Aceros nipalensis'') is a species of hornbill in Bhutan, northeastern India, especially in Arunachal Pradesh, Indian Subcontinent and Southeast Asia. It is locally extinct in Nepal due to hunting and significant loss ...

''. It has often been lumped with the

plain-pouched hornbill The plain-pouched hornbill (''Rhyticeros subruficollis'') is a species of hornbill in the family Bucerotidae. It is found in forests of the Dawna Range and the Tenasserim Hills of southern Myanmar, adjacent parts of western Thailand and northern ...

(''R. subruficollis''), and sometimes considered to include the

Narcondam hornbill The Narcondam hornbill (''Rhyticeros narcondami'') is a species of hornbill in the family Bucerotidae. It is endemic to the Indian island of Narcondam in the Andamans. Males and females have a distinct plumage. The Narcondam hornbill has the sma ...

(''R. narcondami'') and the

wreathed hornbill The wreathed hornbill (''Rhyticeros undulatus'') is an Old World tropical bird of the hornbill family Bucerotidae, also called bar-pouched wreathed hornbill due to its distinctive blue-black band on its lower throat sac. It is named after its cha ...

(''R. undulatus'') as

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

. The common name commemorates

Edward Blyth Edward Blyth (23 December 1810 – 27 December 1873) was an English zoologist who worked for most of his life in India as a curator of zoology at the museum of the Asiatic Society of India in Calcutta. Blyth was born in London in 1810. In 1841 ...

(1810–1873), English zoologist and Curator of the Museum of the Asiatic Society of Bengal.

Description

Up to in length, the adult male has mainly black

plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...

with a golden or orange-buff head, white throat and a white tail. Its irises are reddish brown, and the eye is surrounded by naked pale blue skin. The female is a smaller, mainly black bird with a white throat and tail. Both sexes have a very large horn-coloured bill and

casque Casque is a French word for helmet. It can refer to: * Casque (anatomy), an enlargement on the beaks of some species of birds, including many hornbills *Hornbill ivory, the casque of the helmeted hornbill, collected as a decorative material * S. C ...

. Young birds of both sexes resemble the male. Adults have up to eight folds on the pale casque, depending on age, while young birds have none.Coates (1985): p.442 In flight the sound of its wings is loud and distinctive, a rushing noise that has been compared to the sound of steam escaping from a steam locomotive. It has a range of far-reaching,

guttural Guttural speech sounds are those with a primary place of articulation near the back of the oral cavity, especially where it's difficult to distinguish a sound's place of articulation and its phonation. In popular usage it is an imprecise term for s ...

grunting and laughing calls.

Subspecies

Various subspecies have been described across its range: * ''R. p. plicatus'' (Forster, 1781) – South Moluccas * ''R. p. ruficollis'' (Vieillot, 1816) – North Moluccas and West Papua, eastwards to the Southern Highlands and

Simbu Province Chimbu, more frequently spelled Simbu, is a province in the Highlands Region of Papua New Guinea. The province has an area of 6,112 km2 and a population of 376,473 (2011 census). The capital of the province is Kundiawa. Mount Wilhelm, the tal ...

Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...

* ''R. p. jungei'' Mayr, 1937 – Eastern

New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...

, west as far as the

Fly River The Fly River is the third longest river in the island of New Guinea, after the Sepik River and Mamberamo River, with a total length of and the largest by volume of discharge in Oceania, the largest in the world without a single dam in its catc ...

region * ''R. p. dampieri'' Mayr, 1934 –

Bismarck Archipelago The Bismarck Archipelago (, ) is a group of islands off the northeastern coast of New Guinea in the western Pacific Ocean and is part of the Islands Region of Papua New Guinea. Its area is about 50,000 square km. History The first inhabitants o ...

* ''R. p. harterti'' Mayr, 1934 – Bougainville and

Buka Island Buka Island is the second-largest island in the Autonomous Region of Bougainville, in eastern Papua New Guinea. It is in Buka Rural LLG of North Bougainville District, with the Autonomous Region's and district's capital city of Buka, Bougain ...

s * ''R. p. mendanae'' Hartert, 1924 –

Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capita ...

from Choiseul to

Guadalcanal Guadalcanal (; indigenous name: ''Isatabu'') is the principal island in Guadalcanal Province of Solomon Islands, located in the south-western Pacific, northeast of Australia. It is the largest island in the Solomon Islands by area, and the seco ...

and

Malaita Malaita is the primary island of Malaita Province in Solomon Islands. Malaita is the most populous island of the Solomon Islands, with a population of 161,832 as of 2021, or more than a third of the entire national population. It is also the se ...

Distribution and ecology

The Papuan hornbill occurs throughout lowland forests, from sea level up to 1,200–1,500 m

ASL American Sign Language (ASL) is a natural language that serves as the predominant sign language of Deaf communities in the United States of America and most of Anglophone Canada. ASL is a complete and organized visual language that is express ...

, in the

Moluccas The Maluku Islands (; Indonesian: ''Kepulauan Maluku'') or the Moluccas () are an archipelago in the east of Indonesia. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located eas ...

, the

, and as far east as the

. It is the only hornbill

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

native to New Guinea, and one of the largest flying birds of the region. Its diet consists mainly of fruits, especially

figs The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the world ...

, occasionally supplemented with

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...

s and other small animals. The Papuan hornbill nests in a large

tree hollow A tree hollow or tree hole is a semi-enclosed cavity which has naturally formed in the trunk or branch of a tree. They are found mainly in old trees, whether living or not. Hollows form in many species of trees, and are a prominent feature of nat ...

in the

rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...

, from at least up to above the ground. The female is restricted to the nest cavity throughout the incubation and nestling period, being largely sealed inside by plastering up the entrance with a mixture of fruit pulp and rotten wood, leaving only a narrow aperture through which the male feeds her. The clutch size is about two eggs.

Status and relationship with humans

Still widespread throughout its large range, the Papuan hornbill is assessed as

least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...

on the

IUCN Red List of Threatened Species The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biologi ...

. However, it is subject to hunting pressure by some tribal groups, who use its feathers in

headdress Headgear, headwear, or headdress is the name given to any element of clothing which is worn on one's head, including hats, helmets, turbans and many other types. Headgear is worn for many purposes, including protection against the elements, de ...

es, its bill as a personal adornment, and the lower mandible as a spear point. As a consequence, it is becoming rarer in some areas of New Guinea. On the other hand, this species has withstood tens of

millennia A millennium (plural millennia or millenniums) is a period of one thousand years, sometimes called a kiloannum (ka), or kiloyear (ky). Normally, the word is used specifically for periods of a thousand years that begin at the starting point (ini ...

of human hunting pressure. So as long as sufficient

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

is preserved, it is unlikely that hunting alone will prove a significant threat unless population increases include concomitant increases in hunting pressures.Majnep & Bulmer (1977): p.129, Coates (1985): pp.442,444, Steadman (2006): pp.366-367

References

* Coates, Brian J. (1985): ''The Birds of Papua New Guinea'' (Vol.1: Non-Passerines). Dove Publications, Alderley, Queensland, Australia. * Coates, Brian J. & Bishop, K. David (1997): ''A Guide to the Birds of Wallacea, Sulawesi, the Moluccas and Lesser Sunda Islands, Indonesia''. Dove Publications, Alderley, Queensland, Australia. * Majnep, Ian Saem & Bulmer, Ralph (1977): ''Birds of my Kalam Country''. Auckland University Press, Auckland, New Zealand. * * Steadman, David William (2006): ''Extinction and Biogeography of Tropical Pacific Birds''. University of Chicago Press.

External links

BirdLife Species Factsheet
{{Taxonbar, from=Q1193728

Blyth's hornbill Blyth's hornbill (''Rhyticeros plicatus''), also known as the Papuan hornbill, is a large hornbill inhabiting the forest canopy in Wallacea and Melanesia. Its local name in Tok Pisin is ''kokomo''. Previously, this hornbill was placed in the gen ...

Birds of the Maluku Islands Birds of Melanesia