Lemonadeberry Rhus integrifolia in bloom, MBSP

''Rhus integrifolia'', also known as lemonade sumac, lemonade berry, or lemonadeberry, is a

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

to small

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

. It is native to the

Transverse Transverse may refer to: *Transverse engine, an engine in which the crankshaft is oriented side-to-side relative to the wheels of the vehicle *Transverse flute, a flute that is held horizontally * Transverse force (or ''Euler force''), the tangen ...

and

Peninsular Ranges The Peninsular Ranges (also called the Lower California province) are a group of mountain ranges that stretch from Southern California to the southern tip of the Baja California peninsula; they are part of the North American Coast Ranges, which ...

and the

South Coast South Coast is a name often given to coastal areas to the south of a geographical region or major metropolitan area. Geographical Australia *South Coast (New South Wales), the coast of New South Wales, Australia, south of Sydney * South Coast (Q ...

regions of

Southern California Southern California (commonly shortened to SoCal) is a geographic and Cultural area, cultural region that generally comprises the southern portion of the U.S. state of California. It includes the Los Angeles metropolitan area, the second most po ...

. This extends from Santa Barbara County and the

Channel Islands The Channel Islands ( nrf, Îles d'la Manche; french: îles Anglo-Normandes or ''îles de la Manche'') are an archipelago in the English Channel, off the French coast of Normandy. They include two Crown Dependencies: the Bailiwick of Jersey, ...

San Diego County San Diego County (), officially the County of San Diego, is a county in the southwestern corner of the U.S. state of California. As of the 2020 census, the population was 3,298,634, making it California's second-most populous county and the fi ...

and extending into north-central Pacific coastal

Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...

and its offshore islands such as Cedros Island. It is in height, with a sprawling form, and is a member of the

chaparral Chaparral ( ) is a shrubland plant community and geographical feature found primarily in the U.S. state of California, in southern Oregon, and in the northern portion of the Baja California Peninsula in Mexico. It is shaped by a Mediterranean c ...

plant community often found in canyons and on north-facing slopes below elevations of . ''Rhus integrifolia'' often hybridizes with ''

Rhus ovata ''Rhus ovata'', commonly known as sugar bush or sugar sumac, is a shrub or small tree found growing in the canyons and slopes of the chaparral and related ecosystems in Southern California, Arizona, Baja California and Baja California Sur. It ...

''.

Description

''Rhus integrifolia'' leaves are simple (unusual in a genus where most species are trifoliate), alternating,

evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...

and leathery, ranging from two to four centimeters wide on reddish twigs; length of leaves is five to seven centimeters. Leaves are toothed with a waxy smooth appearance above and a paler tone below. The

flowers A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...

which appear from February to May are small, clustered closely together, and may be either

bisexual Bisexuality is a romantic or sexual attraction or behavior toward both males and females, or to more than one gender. It may also be defined to include romantic or sexual attraction to people regardless of their sex or gender identity, whi ...

pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...

late. These fragrant flowers exhibit radial symmetry with five green

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...

s, five white to rosy-pink

petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...

s, and five

stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

s. The small flowers are only six millimeters across. The ovary is superior and usually has a single

ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fe ...

; although in pistillate flowers, the stamens are small and infertile. The mature

fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...

of ''Rhus integrifolia'' is sticky, reddish, covered with hairs, and about seven to ten millimeters in diameter. The elliptical fruit presents tight clusters at the very ends of twigs. Young plants manifest smooth reddish

bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, ...

, while more mature individuals have cracked, even scaly, grayish bark with the smooth red bark displayed underneath.

Twig A twig is a thin, often short, branch of a tree or bush. The buds on the twig are an important diagnostic characteristic, as are the abscission scars where the leaves have fallen away. The color, texture, and patterning of the twig bark are ...

s are rather stout and flexible, and reddish bud ends are diminutive and pointed. There is often a multi-furcate branching structure from the base of the plant. A mature plant is large and thicket-like with a sprawling arrangement.

Ecology

The lemonade berry plant is found on north and east-facing slopes and canyon floors in coastal areas of southern California and northern Baja California; however, one colony has even been observed as far north as Santa Cruz County. The lemonade berry occurs in both

and

coastal sage scrub Coastal sage scrub, also known as coastal scrub, CSS, or soft chaparral, is a low scrubland plant community of the California coastal sage and chaparral subecoregion, found in coastal California and northwestern coastal Baja California. It is w ...

communities. This plant thrives on well-drained soils and endures heat and windy conditions well. The species tolerates sandy as well as medium loam soils, and it can even thrive in

nutrient A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...

deficient soil. This plant tolerates mildly

acid In computer science, ACID ( atomicity, consistency, isolation, durability) is a set of properties of database transactions intended to guarantee data validity despite errors, power failures, and other mishaps. In the context of databases, a sequ ...

to mildly

alkaline In chemistry, an alkali (; from ar, القلوي, al-qaly, lit=ashes of the saltwort) is a base (chemistry), basic, ionic compound, ionic salt (chemistry), salt of an alkali metal or an alkaline earth metal. An alkali can also be defined as ...

soils, and is common on north and east-facing slopes with where shady, moist conditions are prevalent. Propagation is by seed or by

rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...

. Bird species enjoy the fruits and act as a major disperser. Lemonade berry is not frost tolerant, but will hybridize with the frost tolerant

(sugarbush) in the foothills of California and Mexico where their ranges overlap.

Uses

Many plants within this genus are considered

toxic Toxicity is the degree to which a chemical substance or a particular mixture of substances can damage an organism. Toxicity can refer to the effect on a whole organism, such as an animal, bacterium, or plant, as well as the effect on a subst ...

, although some reports indicate the berries of this species can be used to make lemonade flavored drinks (hence its common name). Allergic reactions may also result from skin contact with

sap Sap is a fluid transported in xylem cells (vessel elements or tracheids) or phloem sieve tube elements of a plant. These cells transport water and nutrients throughout the plant. Sap is distinct from latex, resin, or cell sap; it is a separa ...

from some of the genera. Lemonade berry leaves are rich in

tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'', ...

s. Even though the species is evergreen, there is some leaf fall in autumn, at which time the fallen leaves may be used as a brown

dye A dye is a colored substance that chemically bonds to the substrate to which it is being applied. This distinguishes dyes from pigments which do not chemically bind to the material they color. Dye is generally applied in an aqueous solution an ...

mordant A mordant or dye fixative is a substance used to set (i.e. bind) dyes on fabrics by forming a coordination complex with the dye, which then attaches to the fabric (or tissue). It may be used for dyeing fabrics or for intensifying stains in ...

. An oil can be extracted from lemonade berry seeds; moreover, this oil achieves a

tallow Tallow is a rendering (industrial), rendered form of beef or mutton fat, primarily made up of triglycerides. In industry, tallow is not strictly defined as beef or mutton fat. In this context, tallow is animal fat that conforms to certain techn ...

consistency when left to stand. Thereafter the oil can be employed to manufacture

candle A candle is an ignitable wick embedded in wax, or another flammable solid substance such as tallow, that provides light, and in some cases, a fragrance. A candle can also provide heat or a method of keeping time. A person who makes candles i ...

s, which burn brightly, albeit emitting a pungent scent. The wood of mature plants is dense and hard, making it prized for wood-burning

fireplace A fireplace or hearth is a structure made of brick, stone or metal designed to contain a fire. Fireplaces are used for the relaxing ambiance they create and for heating a room. Modern fireplaces vary in heat efficiency, depending on the design. ...

kindling.

Cultivation

''Rhus integrifolia'' can also be used as a landscape

and is suitable for hedging and espalier. The plant is vulnerable to frost, but often the plant will regrow by summer, after it appears to have died from cold. The growth habit is slow to moderate and, as a garden plant, this

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

is quite resistant to

deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...

, and supplies nectar and fruit for birds and nectar for butterflies. Its cultural requirements mimic its natural environment with ample drainage and little summer water.

References

External links