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''Chenopodium spinescens'' (common names: spiny saltbush, berry saltbush, thorny saltbush, creeping saltbush, hedge saltbush) is a species of plant in the family
Amaranthaceae Amaranthaceae is a family of flowering plants commonly known as the amaranth family, in reference to its type genus ''Amaranthus''. It includes the former goosefoot family Chenopodiaceae and contains about 165 genera and 2,040 species, making it ...
and is endemic to all mainland states and territories of Australia where it is known as ''Rhagodia spinescens''.


Description

''Chenopodium spinescens'' is a multi-stemmed dense shrub to high, with branches often ending in spines. The leaves are alternate or opposite, and mealy, looking grey green to almost whitish from their covering of saucer-shaped/spherical hairs. The leaf shape is ovate to triangular, and are long and wide. The leaf petiole is about half the length of the lamina. Male and female flowers are usually on different plants. The flowers have five perianth segments that are tiny and all alike, and five stamens. The ripe fruit are red and succulent. The inflorescence is a panicle or spike, with the flowers' perianth being densely mealy on the outside. The male flowers are globular and 0.5–1 mm in diameter, with their filaments united into a smooth saucer-shaped disc. The female flowers are larger (about 1 mm) and globular to top-shaped. The fruit is a depressed globular pink or red succulent berry, and the seed is approximately in diameter with a reticulate coat. In the
Northern Territory The Northern Territory (commonly abbreviated as NT; formally the Northern Territory of Australia) is an Australian territory in the central and central northern regions of Australia. The Northern Territory shares its borders with Western Aust ...
, it flowers and fruits from February to December, while in
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
, it flowers from January to April.


Differences from closely related species

The
Northern Territory Government The Government of the Northern Territory of Australia, also referred to as the Northern Territory Government, is the Australian territorial democratic administrative authority of the Northern Territory. The Government of Northern Territory wa ...
website accepts the names ''R. spinescens'' (for ''C. spinescens''), ''R. parabolica'' (for '' C. parabolicum'') and ''R. eremaea'' (for '' C. eremaea''), and describes the differences between them as follows:
''Rhagodia spinescens'' differs from '' R. parabolica'', in having smaller leaves (mostly 5-17 mm long, compared to mostly 15-30 mm long), a narrower inflorescence and a preference for non-rocky sites. It differs from '' R. eremaea'' in having leaves that are alternate or opposite (compared to consistently alternate), often spade-shaped, equal length and width or slightly longer than wide (cf. distinctly long relative to their width), with round saucer-like vesicles on the undersurface (compared irregular-shaped vesicles) and have a fishy or foetid odour.


Taxonomy

This saltbush was first formally described in 1810 by Robert Brown who gave it the name ''Rhagodia spinescens'' in ''
Prodromus Florae Novae Hollandiae et Insulae Van Diemen ''Prodromus Florae Novae Hollandiae et Insulae Van Diemen'' (Prodromus of the Flora of New Holland and Van Diemen's Land) is a flora of Australia written by botanist Robert Brown and published in 1810. Often referred to as ''Prodromus Flora Novae ...
''. In 2012, Susy Fuentes Bazán and Thomas Borsch moved all species of ''Rhagodia'' to ''Chenopodium'' within the broader family, ''Amaranthaceae'', a change accepted by
Rafaël Govaerts Rafaël Herman Anna Govaerts (born 1968) is a Belgian botanist. He is particularly noted for his work on plant taxonomy. He has worked at the Royal Botanic Gardens, Kew since the 1990s, and is the principal contributor to the World Checklist of ...
at Plants of the World Online. However, the assignation of this plant to the genus, '' Rhagodia'', and to the family, ''
Chenopodiaceae Amaranthaceae is a family of flowering plants commonly known as the amaranth family, in reference to its type genus '' Amaranthus''. It includes the former goosefoot family Chenopodiaceae and contains about 165 genera and 2,040 species, making i ...
'', is the taxonomy accepted by the
Council of Heads of Australasian Herbaria Council of Heads of Australasian Herbaria (CHAH) is an association of the leaders of herbaria in Australia and New Zealand. It is governed by a constitution. It endorses the taxonomy and nomenclature of the Australian Plant Census, which is the so ...
, the
National Herbarium of New South Wales The National Herbarium of New South Wales was established in 1853. The Herbarium has a collection of more than 1.4 million plant specimens, making it the second largest collection of pressed, dried plant specimens in Australia,Thiers, B. ...
,
Royal Botanic Gardens Victoria Royal Botanic Gardens Victoria are botanic gardens across two sites–Melbourne and Cranbourne. Melbourne Gardens was founded in 1846 when land was reserved on the south side of the Yarra River for a new botanic garden. It extends across ...
,
State Herbarium of South Australia The State Herbarium of South Australia, sometimes called the South Australian Herbarium, and having the herbarium code, AD, is located in Adelaide, South Australia. It is one of several State and Commonwealth herbaria in Australia. The Depa ...
, the Northern Territory Government, Queensland Department of Environment and Science and Western Australian Government
Department of Biodiversity, Conservation and Attractions The Department of Biodiversity, Conservation and Attractions (DBCA) is the Western Australian government department responsible for managing lands and waters described in the ''Conservation and Land Management Act 1984'', the ''Rottnest Island ...
.


Habitat

In the Northern Territory, it occurs on sandplains, alluvial plains, intermittent sandy watercourses, on sandy edges of salt lakes, and rarely on gravelly hillslopes. In Victoria, it is usually found on heavy, saline or subsaline soils at the edges of lakes, on flats, and in dune swales.


Distribution

It is found in the IBRA bioregions of Burt Plain, Central Ranges,
Channel Country The Channel Country is a region of outback Australia mostly in the state of Queensland but also in parts of South Australia, Northern Territory and New South Wales. The name comes from the numerous intertwined rivulets that cross the region, ...
, Coolgardie, Finke,
Flinders Lofty Block Flinders Lofty Block is an interim Australian bioregion located in South Australia. It has an area of , which includes the Mount Lofty Ranges and Flinders Ranges The Flinders Ranges are the largest mountain range in South Australia, whic ...
,
Gawler Gawler is the oldest country town on the Australian mainland in the state of South Australia. It was named after the second Governor (British Vice-Regal representative) of the colony of South Australia, George Gawler. It is about north of the ...
, Great Sandy Desert,
Great Victoria Desert The Great Victoria Desert is a sparsely populated desert ecoregion and interim Australian bioregion in Western Australia and South Australia. History In 1875, British-born Australian explorer Ernest Giles became the first European to cros ...
, MacDonnell Ranges,
Mitchell Grass Downs The Mitchell Grass Downs is a tropical and subtropical grasslands, savannas, and shrublands ecoregion in northeastern Australia. It is a mostly treeless grassland, characterised by Mitchell grasses (''Astrebla'' spp.). Location and descrip ...
,
Nullarbor The Nullarbor Plain ( ; Latin: feminine of , 'no', and , 'tree') is part of the area of flat, almost treeless, arid or semi-arid country of southern Australia, located on the Great Australian Bight coast with the Great Victoria Desert to its ...
,
Riverina The Riverina is an agricultural region of south-western New South Wales, Australia. The Riverina is distinguished from other Australian regions by the combination of flat plains, warm to hot climate and an ample supply of water for irrigation ...
,
Simpson Strzelecki Dunefields The Simpson Strzelecki Dunefields, an interim Australian bioregion, comprises , and is part of four state/territories of Australia: the Northern Territory, South Australia, New South Wales and Queensland
,
Stony Plains Stony Plains, an interim Australian bioregion, comprises , and is part of two state/territories of Australia: the Northern Territory and South Australia.
, and Tanami. (Bioregions from Victoria, Queensland and New South Wales may not be included.)


Propagation

The advice for propagation is to take soft, current season growth tips from actively growing plants; store and transport wrapped in wet newspaper and sealed in a plastic bag, and plant after trimming cuttings to a length of and dipping in 3000 ppm IBA rooting hormone.


References


External links


Occurrence data for ''Rhagodia spinescens'' R.Br.
The Australasian Virtual Herbarium. Retrieved 7 July 2018.
(2017) ''Rhagodia spinescens''. In: Flora of Australia. Australian Biological Resources Study, Department of the Environment and Energy, Canberra.
{{Taxonbar, from=Q17246017, from2=Q15553767 spinescens Caryophyllales of Australia Flora of New South Wales Flora of South Australia Flora of Queensland Flora of Victoria (Australia) Eudicots of Western Australia Plants described in 1810