''Retrophyllum rospigliosii'' is a species of

conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...

in the family

Podocarpaceae Podocarpaceae is a large family of mainly Southern Hemisphere conifers, known in English as podocarps, comprising about 156 species of evergreen trees and shrubs.James E. Eckenwalder. 2009. ''Conifers of the World''. Portland, Oregon: Timber Pre ...

. It is a large

evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

native to the montane rainforests of

Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...

Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...

Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...

Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...

and

Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...

Etymology and names

The

specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

''rospigliosii'' honors C. J. Rospigliosi of the

Museum of Natural History A natural history museum or museum of natural history is a scientific institution with natural history collections that include current and historical records of animals, plants, fungi, ecosystems, geology, paleontology, climatology, and more ...

Lima Lima ( ; ), originally founded as Ciudad de Los Reyes (City of The Kings) is the capital and the largest city of Peru. It is located in the valleys of the Chillón River, Chillón, Rímac River, Rímac and Lurín Rivers, in the desert zone of t ...

. The species has multiple common names in the Spanish language. These include ''pino hayuelo'', ''pino laso'', ''pino de montana'', ''pino romeron'' and ''romerillo macho''.

Description

''Retrophyllum rospigliosii'' is an evergreen tree growing up to 45 meters in height and 1,8 meters in trunk diameter. The trunk is usually erect and straight. The

crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...

typically becomes rounded or oval in shape as the tree matures.

Branch A branch, sometimes called a ramus in botany, is a woody structural member connected to the central trunk (botany), trunk of a tree (or sometimes a shrub). Large branches are known as boughs and small branches are known as twigs. The term '' ...

es are generally ascending or spreading, but become pendulous on shaded parts of the crown. The bark is initially brown and weathers dark gray, exfoliating in scale-like flakes. The

leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

are generally flattened with a decurrent base and a spreading blade, but leading shoots may also have appressed scale-like leaves. The leaves are inserted spirally but are twisted on lateral branchlets to appear pectinate and nearly opposite. The pectinate leaves are twisted at their petioles in opposite directions on each side of the shoot causing the adaxial sides of the leaves face up on one side of the shoot and down on the other side. The leaf blades are most commonly 10-18 millimeters long and 3-5 millimeters wide. They are ovate-lanceolate or ovate-elliptic in shape with visible midribs and entire margins. Stomata can be found on both surfaces of the leaf. The species is

dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...

. The male pollen cones are 5-7 millimeters long with spirally arranged triangular sporophylls. The female

seed cones A conifer cone (in formal botanical usage: strobilus, plural strobili) is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in conifers an ...

each have a single subterminal fertile cone scale with a single

ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fe ...

developing into a

seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...

. The seed is covered by a fleshy epimatium. The initially green or glaucous epimatium becomes red at maturity and varies from subglobose to ovoid-pyriform in shape with a length of 18-25 millimeters.

Distribution

''Retrophyllum rospigliosii'' is native to northwestern South America. It occurs naturally in Bolivia,BOLIVIA Retrophyllum rospigliosii, A New Record for Northwest of Pine Mountain Bolivia
(Spanish language) Colombia, Ecuador, Peru and Venezuela.

Habitat and ecology

''Retrophyllum rospigliosii'' is a large

rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...

tree. It occurs in montane tropical rainforests and

cloud forests A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest (TMCF), is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud ...

at altitudes between 1500 and 3750 meters above the sea level. Most commonly the species grows scattered among other trees, most of which are

angiosperms Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

, but on certain sites it may also form pure stands.

References

''Page Notes from the Royal Botanic Garden'', Edinburgh 45: 380. 1989.
{{Taxonbar, from=Q4251304 Podocarpaceae Near threatened plants Trees of Bolivia Trees of Peru Trees of Ecuador Trees of Colombia Trees of Venezuela Taxonomy articles created by Polbot