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The red spitting cobra (''Naja pallida'') is a species of
spitting cobra A spitting cobra is any of several species of cobras that can defensively spray a toxic secretion - functioning as both a venom (that can be injected via a wound) and a toxungen (that can be sprayed on the target surface) - from their fangs in ...
native to
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
.


Description

This medium-sized cobra attains lengths between , but may grow to a maximum length of around in very rare cases. It is often thought of as an attractive species; it is usually bright salmon-red contrasted with a broad black throat band and subocular teardrop markings. However, the color of this species does have variation, which usually depends on where in Africa a particular specimen is found. For example, specimens from southern Kenya and northern Tanzania have an orange-red colour, with a broad, dark blue or black throat band. Some specimens may have two or three throat bands, but this is uncommon for specimens from
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
. The ventral side is also reddish in colour, sometimes the throat area may be a creamy white. Specimens from other areas can be yellow, pinkish, pink-grey, pale red or steel grey. Most specimens will have a throat band, but it will fade or even sometimes disappear in larger adults. The true red specimens will become reddish-brown in colour as they age and grow in size. The body of this snake is slightly depressed, tapered and moderately slender with a medium-length tail. It is slightly compressed dorsoventrally and subcylindrical posteriorly. The head is broad, flattened and slightly distinct from the neck. The canthus is distinct and the snout is rounded. The eyes are medium to large in size with round pupils. Dorsal scales are smooth and strongly oblique.


Scalation

Dorsal scales on the midbody are in 21–27, ventrals are in 197–228, and subcaudals are in 61–72 rows, subscales are paired, and the anal scale is single. There are seven upper labials, one upper labial enters the eye; two preoculars, three postoculars, and the lower labials are usually eight in number (range 7-9).


Distribution and habitat

The red spitting cobra is mainly found in
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
, including
Djibouti Djibouti, ar, جيبوتي ', french: link=no, Djibouti, so, Jabuuti officially the Republic of Djibouti, is a country in the Horn of Africa, bordered by Somalia to the south, Ethiopia to the southwest, Eritrea in the north, and the Red ...
,
Eritrea Eritrea ( ; ti, ኤርትራ, Ertra, ; ar, إرتريا, ʾIritriyā), officially the State of Eritrea, is a country in the Horn of Africa region of Eastern Africa, with its capital and largest city at Asmara. It is bordered by Ethiopia ...
,
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...
, southern
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Mediter ...
, northern and eastern
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
, and northern
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
and northern
Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...
. It is also widespread in the dry country of eastern and northern
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
. It primarily inhabits dry savanna and semidesert areas of East Africa up to an elevation of about above sea level. They can usually be found near water holes.


Behavior

Red spitting cobras are terrestrial, fast and alert snakes. Adult specimens of this species are nocturnal, while juveniles are more active during the day. Adults like to hide in termite mounds, old logs, holes, brush piles or any other ground cover during the day. They are also known to be
cannibalistic Cannibalism is the act of consuming another individual of the same species as food. Cannibalism is a common ecological interaction in the animal kingdom and has been recorded in more than 1,500 species In biology, a species is the basic ...
; this could be the reason juveniles and smaller specimens are diurnal, while adults are nocturnal. When threatened, this cobra rears up and displays a typical cobra hood. It may also hiss loudly. If the intruder does not retreat, it may spray jets of venom to the face of the intruder. Venom in the eyes can cause burning pain and blindness. Even so, this snake seldom causes fatalities in humans.


Diet

This species has a wide range of prey, but they prefer amphibians such as toads and frogs, if and when they are available. However, they will prey on rodents, birds and probably other snakes. They are known to raid chickens in the region.


Reproduction

The red spitting cobra is
oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...
. The female will lay 6 to 21 eggs.


Venom

The venom of this species, like most spitting cobras, contains a mixture of
neurotoxins Neurotoxins are toxins that are destructive to nerve tissue (causing neurotoxicity). Neurotoxins are an extensive class of exogenous chemical neurological insultsSpencer 2000 that can adversely affect function in both developing and mature nerv ...
and
cytotoxins Cytotoxicity is the quality of being toxic to cells. Examples of toxic agents are an immune cell or some types of venom, e.g. from the puff adder (''Bitis arietans'') or brown recluse spider (''Loxosceles reclusa''). Cell physiology Treating cell ...
. Bite symptoms include slight pain around the wound and numbness of the lips, fingers and tongue. Although it rarely causes human fatalities, survivors are usually disfigured. The
murine The Old World rats and mice, part of the subfamily Murinae in the family Muridae, comprise at least 519 species. Members of this subfamily are called murines. In terms of species richness, this subfamily is larger than all mammal families excep ...
IP value for this snake is 2 mg/kg. In September 2011, a snake keeper from the
Eastern Cape The Eastern Cape is one of the provinces of South Africa. Its capital is Bhisho, but its two largest cities are East London and Gqeberha. The second largest province in the country (at 168,966 km2) after Northern Cape, it was formed in ...
in
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
died shortly after a red spitting cobra sprayed venom into his face while he and a friend were photographing the snake. Some of the venom was believed to have entered his nasal passages and led to
anaphylactic shock Anaphylaxis is a serious, potentially fatal allergic reaction and medical emergency that is rapid in onset and requires immediate medical attention regardless of use of emergency medication on site. It typically causes more than one of the follow ...
. His asthma is believed to have contributed to the fatal reaction. An autopsy done on him by the eastern cape provincial pathologist revealed no biting or damage either in his nasal cavities, throat or lungs.


Taxonomy

This species was formerly considered to be a subspecies of ''
Naja mossambica The Mozambique spitting cobra (''Naja mossambica'') is a highly venomous species of spitting cobra native to Africa. It is largely found in Angola, Botswana, Malawi, Mozambique, Namibia, South Africa, Tanzania, Zambia, and Zimbabwe. Taxonomy Germ ...
'', ''N. m. pallida'', but is now categorized as a separate species. Indeed, ''N. mossambica'' is more closely related to '' N. nigricollis'' than to this species. The red spitting cobra is closely related and forms a
sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
with the
Nubian spitting cobra The Nubian spitting cobra (''Naja nubiae'') is a species of spitting cobra native to Africa. Description A relatively small spitting cobra. Maximum recorded length 148 cm. Colour and pattern: Brownish-grey overall, scale bases and skin betwe ...
(''Naja nubiae'') of northeastern Africa, which was originally considered a northern population of ''N. m. pallida'' before being recognized as a distinct species in 2003.


References

{{Taxonbar, from=Q1469485 Naja Snakes of Africa Reptiles described in 1896