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The red kangaroo (''Osphranter rufus'') is the largest of all
kangaroo Kangaroos are four marsupials from the family Macropodidae (macropods, meaning "large foot"). In common use the term is used to describe the largest species from this family, the red kangaroo, as well as the antilopine kangaroo, eastern gre ...
s, the largest terrestrial
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
native to
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, and the
largest Large means of great size. Large may also refer to: Mathematics * Arbitrarily large, a phrase in mathematics * Large cardinal, a property of certain transfinite numbers * Large category, a category with a proper class of objects and morphisms (o ...
extant
marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a po ...
. It is found across mainland
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, except for the more fertile areas, such as southern
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
, the eastern and southeastern coasts, and the rainforests along the northern coast.


Taxonomy

The initial description of the species by A.G. Desmarest was published in 1822. The type location was given as an unknown location west of the Blue Mountains. The author assigned the new species to the genus ''Kangurus''. In 1842, Gould reassigned the species to the genus ''
Osphranter ''Osphranter'' is a genus of large marsupials in the family Macropodidae, commonly known as kangaroos and wallaroos (among other species). It contains the largest extant marsupial, the red kangaroo (''O. rufus''). In 2019, a reassessment of mac ...
'', a taxon later submerged as a subgenus of ''
Macropus ''Macropus'' is a marsupial genus in the family Macropodidae. It has two extant species of large terrestrial kangaroos. The term is derived from the Ancient Greek μάκρος, ''makros'' "long" and πους, ''pous'' "foot". Thirteen known ext ...
''. A taxonomic restructure in 2015 in ''Taxonomy of Australian Mammals'' by Jackson and Groves promoted ''Osphranter'' back to genus level, redefining the red kangaroo, among others, as species within the genus ''Osphranter''. This was further supported by genetic analysis in 2019.


Description

This species is a very large kangaroo with long, pointed ears and a square shaped muzzle nout/nose They are
sexually dimorphic Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
as the males have short, red-brown fur, fading to pale buff below and on the limbs. Females are smaller than males and are blue-grey with a brown tinge, pale grey below, although arid zone females are coloured more like males. It has two forelimbs with small claws, two muscular hind-limbs, which are used for jumping, and a strong tail which is often used to create a tripod when standing upright. Males grow up to a head-and-body length of with a tail that adds a further to the total length and are referred to by Australians as "Big Reds". Females are considerably smaller, with a head-and-body length of and tail length of .Yue, M. (2001
''Macropus rufus''
Animaldiversity.ummz.umich.edu. Retrieved on 2015-09-25.
Red kangaroo videos, photos and facts – ''Macropus rufus''
. ARKive. Retrieved on 2015-09-25.
Females can weigh from , while males typically weigh about twice as much at . The average red kangaroo stands approximately tall to the top of the head in upright posture. Large mature males can stand more than tall, with the largest confirmed one having been around tall and weighed . The red kangaroo maintains its internal temperature at a point of homeostasis about using a variety of physical, physiological, and behavioural adaptations. These include having an insulating layer of fur, being less active and staying in the shade when temperatures are high, panting, sweating, and licking its forelimbs. The red kangaroo's range of vision is approximately 300° (324° with about 25° overlap), due to the position of its eyes.


Locomotion

The red kangaroo's legs work much like a rubber band, with the Achilles tendon stretching as the animal comes down, then releasing its energy to propel the animal up and forward, enabling the characteristic bouncing locomotion. They can reach speeds of around 60 km/h (37 mph). The males can cover 8–9 m (26.2–29.5 ft) in one leap while reaching heights of 1.8–3 m (5.9–9.8 ft), though the average is 1.2–1.9 m (3.9–6.2 ft).


Ecology and habitat

The red kangaroo ranges throughout western and
central Australia Central Australia, also sometimes referred to as the Red Centre, is an inexactly defined region associated with the geographic centre of Australia. In its narrowest sense it describes a region that is limited to the town of Alice Springs and i ...
. Its range encompasses scrubland, grassland, and desert habitats. It typically inhabits open habitats with some trees for shade. Red kangaroos are capable of conserving enough water and selecting enough fresh vegetation to survive in an arid environment. The kangaroo's kidneys efficiently concentrate urine, particularly during summer. Red kangaroo primarily eat green vegetation, particularly fresh grasses and
forb A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...
s, and can get enough even when most plants look brown and dry. One study of kangaroos in Central Australia found that green grass makes up 75–95% of the diet, with '' Eragrostis setifolia'' dominating at 54%. This grass continues to be green into the dry season. Kangaroos also primarily consumed this species, along with ''Enneapogon avanaceus'', in western New South Wales where they comprised much as 21–69% of its diet according to a study. During dry times, kangaroos search for green plants by staying on open grassland and near watercourses. While grasses and forbs are preferred, red kangaroos will also eat certain species of chenopods, like ''
Bassia ''Bassia'' is a genus of flowering plants in the family Amaranthaceae. They are distributed in the western Mediterranean to eastern Asia. Some occur outside their native ranges as introduced species.Maireana ''Maireana '' is a genus of around 57 species of perennial shrubs and herbs in the family Amaranthaceae which are endemic to Australia. Species in this genus were formerly classified within the genus ''Kochia''. The genus was described in 1840 ...
pyramidata'', and will even browse shrubs when its favoured foods are scarce. However, some perennial chenopods, such as round-leaf chenopod ''
Kochia ''Kochia'' ''Sunset Western Garden Book,'' 1995:606–607 is a synonym of the genus ''Bassia'', which belongs to the subfamily Camphorosmoideae of family Amaranthaceae Amaranthaceae is a family of flowering plants commonly known as the amarant ...
'' are avoided even when abundant. At times, red kangaroos congregate in large numbers; in areas with much forage, these groups can number as much as 1,500 individuals. Red kangaroos are mostly
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of daylig ...
and nocturnal, resting in the shade during the day. However, they sometimes move about during the day. Red kangaroos rely on small saltbushes or mulga bushes for shelter in extreme heat rather than rocky outcrops or caves. Grazing takes up most of their daily activities. Like most kangaroo species, they are mostly sedentary, staying within a relatively well-defined home range. However, great environmental changes can cause them to travel great distances. Kangaroos in New South Wales have weekly home ranges of 258–560 ha, with the larger areas belonging to adult males. When forage is poor and rainfall patchy, kangaroos will travel 25–30 km to more favourable feeding grounds. Another study of kangaroos in central Australia found that most of them stay close to remaining vegetation but disperse to find fresh plants after it rains. The red kangaroo is too big to be subject to significant non-human predation. They can use their robust legs and clawed feet to defend themselves from attackers with kicks and blows. However,
dingo The dingo (''Canis familiaris'', ''Canis familiaris dingo'', ''Canis dingo'', or ''Canis lupus dingo'') is an ancient (Basal (phylogenetics), basal) lineage of dog found in Australia (continent), Australia. Its taxonomic classification is de ...
es and
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, just ...
s can kill and eat
joey Joey may refer to: People *Joey (name) Animals * Joey (marsupial), an infant marsupial * Joey, a Blue-fronted Amazon parrot who was one of the Blue Peter pets Film and television * ''Joey'' (1977 film), an American film directed by Horac ...
s. Joeys are thus protected in their mother's pouch. The red kangaroo does have some major predators, like
crocodile Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term crocodile is sometimes used even more loosely to inclu ...
s. Extinct predators include the
marsupial lion ''Thylacoleo'' ("pouch lion") is an extinct genus of carnivorous marsupials that lived in Australia from the late Pliocene to the late Pleistocene (2 million to 46 thousand years ago). Some of these marsupial lions were the largest mammalian pred ...
, the
thylacine The thylacine ( , or , also ) (''Thylacinus cynocephalus'') is an extinct carnivorous marsupial that was native to the Australian mainland and the islands of Tasmania and New Guinea. The last known live animal was captured in 1930 in Tasman ...
, ''
Megalania Megalania (''Varanus priscus'') is an extinct species of giant monitor lizard, part of the megafaunal assemblage that inhabited Australia during the Pleistocene. It is the largest terrestrial lizard known to have existed, reaching an estimated ...
'', the locally extinct
komodo dragon The Komodo dragon (''Varanus komodoensis''), also known as the Komodo monitor, is a member of the monitor lizard family Varanidae that is endemic to the Indonesian islands of Komodo, Rinca, Flores, and Gili Motang. It is the largest extant ...
and the
wonambi ''Wonambi'' is an extinct genus of madtsoiid snakes that lived in late Neogene to late Quaternary Australia. Species of ''Wonambi'' were constrictor snakes unrelated to Australian pythons. Description ''Wonambi'' was a fairly large snake, ...
. Kangaroos are adept
swimmers Swimming is the self-propulsion of a person through water, or other liquid, usually for recreation, sport, exercise, or survival. Locomotion is achieved through coordinated movement of the limbs and the body to achieve hydrodynamic thrust that r ...
, and often flee into waterways if threatened by a predator. If pursued into the water, a kangaroo may use its forepaws to hold the predator underwater so as to
drown Drowning is a type of suffocation induced by the submersion of the mouth and nose in a liquid. Most instances of fatal drowning occur alone or in situations where others present are either unaware of the victim's situation or unable to offer as ...
it.Canadian Museum of Nature – Kangaroo
Retrieved 6 January 2007.


Behaviour

Red kangaroos live in groups of 2–4 members. The most common groups are females and their young. Larger groups can be found in densely populated areas and females are usually with a male. Membership of these groups is very flexible, and males (boomers) are not territorial, fighting only over females (flyers) that come into heat. Males develop proportionately much larger shoulders and arms than females. Most agonistic interactions occur between young males, which engage in ritualised fighting known as ''boxing''. They usually stand up on their hind limbs and attempt to push their opponent off balance by jabbing him or locking forearms. If the fight escalates, they will begin to kick each other. Using their tail to support their weight, they deliver kicks with their powerful hind legs. Compared to other kangaroo species, fights between red kangaroo males tend to involve more wrestling. Fights establish dominance relationships among males, and determine who gets access to estrous females. Dominant males make agonistic behaviours and more sexual behaviours until they are overthrown. Displaced males live alone and avoid close contact with others.


Reproduction

The red kangaroo breeds all year round. The females have the unusual ability to delay the birth of their baby until their previous
joey Joey may refer to: People *Joey (name) Animals * Joey (marsupial), an infant marsupial * Joey, a Blue-fronted Amazon parrot who was one of the Blue Peter pets Film and television * ''Joey'' (1977 film), an American film directed by Horac ...
has left the pouch. This is known as
embryonic diapause Embryonic diapause (from late 19th century English: dia- ‘through’ + pause- 'delay') (aka delayed implantation in mammals) is an evolutionary reproductive strategy used by several animal species across a number of kingdoms, including approximate ...
. Copulation may last 25 minutes. The red kangaroo has the typical reproductive system of a kangaroo. The neonate emerges after only 33 days. Usually only one young is born at a time. It is blind, hairless, and only a few centimetres long. Its hind legs are mere stumps; it instead uses its more developed forelegs to climb its way through the thick fur on its mother's
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
into the pouch, which takes about three to five minutes. Once in the pouch, it fastens onto one of the two teats and starts to feed. Almost immediately, the mother's sexual cycle starts again. Another egg descends into the uterus and she becomes sexually receptive. Then, if she mates and a second egg is fertilised, its development is temporarily halted. Meanwhile, the neonate in the pouch grows rapidly. After approximately 190 days, the baby (called a
joey Joey may refer to: People *Joey (name) Animals * Joey (marsupial), an infant marsupial * Joey, a Blue-fronted Amazon parrot who was one of the Blue Peter pets Film and television * ''Joey'' (1977 film), an American film directed by Horac ...
) is sufficiently large and developed to make its full emergence out of the pouch, after sticking its head out for a few weeks until it eventually feels safe enough to fully emerge. From then on, it spends increasing time in the outside world and eventually, after around 235 days, it leaves the pouch for the last time. While the young joey will permanently leave the pouch at around 235 days old, it will continue to suckle until it reaches about 12 months of age. A doe may first reproduce as early as 18 months of age and as late as five years during drought, but normally she is two and a half years old before she begins to breed. The female red kangaroo is usually permanently
pregnant Pregnancy is the time during which one or more offspring develops (gestation, gestates) inside a woman, woman's uterus (womb). A multiple birth, multiple pregnancy involves more than one offspring, such as with twins. Pregnancy usually occur ...
except on the day she gives birth; however, she has the ability to freeze the development of an embryo until the previous joey is able to leave the pouch. This is known as
embryonic diapause Embryonic diapause (from late 19th century English: dia- ‘through’ + pause- 'delay') (aka delayed implantation in mammals) is an evolutionary reproductive strategy used by several animal species across a number of kingdoms, including approximate ...
, and will occur in times of drought and in areas with poor food sources. The composition of the milk produced by the mother varies according to the needs of the joey. In addition, red kangaroo mothers may "have up to three generations of offspring simultaneously; a young-at-foot suckling from an elongated teat, a young in the pouch attached to a second teat and a blastula in arrested development in the uterus".McCullough, Dale R. and McCullough, Yvette (2000) ''Kangaroos in Outback Australia'', Columbia University Press. . The red kangaroo has also been observed to engage in
alloparental care Alloparenting (also referred to as alloparental care) is a term used to classify any form of parental care provided by an individual towards young that aren't its own direct offspring. These are often referred to as "non-descendant" young, even th ...
, a behaviour in which a female may adopt another female's joey. This is a common parenting behaviour seen in many other animal species like wolves, elephants and
fathead minnow Fathead minnow (''Pimephales promelas''), also known as fathead or tuffy, is a species of temperate freshwater fish belonging to the genus ''Pimephales'' of the cyprinid family. The natural geographic range extends throughout much of North Amer ...
s.


Relationship with humans

The red kangaroo is an abundant species and has even benefited from the spread of agriculture and creation of man-made waterholes. However, competition with livestock and rabbits poses a threat. It is also sometimes shot by farmers as a pest, although a "destruction permit" is required from the relevant state government. Kangaroos dazzled by headlights or startled by engine noise often leap in front of vehicles, severely damaging or destroying smaller or unprotected vehicles. The risk of harm to vehicle occupants is greatly increased if the windscreen is the point of impact. As a result, "kangaroo crossing" signs are commonplace in Australia. Peak times for kangaroo/vehicle crashes are between 5:00 PM and 10:00 PM, in winter, and after extended dry-weather spells.


Commercial use

Like all
Australian wildlife The fauna of Australia consists of a huge variety of animals; some 46% of birds, 69% of mammals, 94% of amphibians, and 93% of reptiles that inhabit the continent are endemic to it. This high level of endemism can be attributed to the continen ...
, the red kangaroo is protected by legislation, but it is so numerous that there is regulated harvest of its hide and
meat Meat is animal flesh that is eaten as food. Humans have hunted, farmed, and scavenged animals for meat since prehistoric times. The establishment of settlements in the Neolithic Revolution allowed the domestication of animals such as chic ...
. Hunting permits and commercial harvesting are controlled under nationally approved management plans, which aim to maintain red kangaroo populations and manage them as a renewable resource. Harvesting of kangaroos is controversial, particularly due to the animal's popularity. In the year 2000, 1,173,242 animals were killed. In 2009 the government put a limit of 1,611,216 for the number of red kangaroos available for commercial use. The
kangaroo industry The kangaroo industry in Australia is based on the regulated harvesting of the large, abundant species of kangaroos. Limitation of the numbers of kangaroo can have environmental benefits, and is a way of meat production that does not involve int ...
is worth about A$270 million each year, and employs over 4000 people. The kangaroos provide meat for both humans and pet food. Kangaroo meat is very lean with only about 2% fat. Their skins are used for leather.


See also

* Roo bar


References


External links

* {{Taxonbar , from = Q53462 Macropods Mammals of Western Australia Mammals of South Australia Mammals of the Northern Territory Mammals of New South Wales Mammals of Queensland Mammals of Victoria (Australia) Marsupials of Australia Mammals described in 1822 Extant Gelasian first appearances