''Sambucus racemosa'' is a species of

elderberry ''Sambucus'' is a genus of flowering plants in the family Adoxaceae. The various species are commonly called elder or elderberry. The genus was formerly placed in the honeysuckle family, Caprifoliaceae, but was reclassified as Adoxaceae due to ge ...

known by the common names red elderberry and red-berried elder.

Distribution and habitat

It is native to Europe, northern temperate Asia, and North America across

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

and the

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

. It grows in

riparian A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks ar ...

environments,

woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...

s, and other

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s, generally in moist areas.

Description

''Sambucus racemosa'' is often a treelike

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

growing tall. The stems are soft with a

pith Pith, or medulla, is a tissue in the stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other ...

y center. Each individual leaf is composed of 5 to 7 leaflike leaflets, each of which is up to long, lance-shaped to narrowly oval, and irregularly serrated along the edges. The leaflets have a strong disagreeable odor when crushed.Trees, Shrubs, and Woody Vines of North Carolina: Red Elderberry (''Sambucus racemosa'' var. ''pubens'')
/ref> The

inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...

is a vaguely cone-shaped

panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...

of several cymes of flowers blooming from the ends of stem branches. The flower buds are pink when closed, and the open flowers are white, cream, or yellowish. Each flower has small, recurved petals and a star-shaped axis of five white

stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

s tipped in yellow anthers. The flowers are fragrant and visited by

hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics aro ...

s and

butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...

. The fruit is a bright red or sometimes purple

drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...

containing 3 to 5 seeds.

Varieties and subspecies

* ''Sambucus racemosa'' subsp. ''kamtschatica'' — red elder, native to

Northeastern Asia Northeast Asia or Northeastern Asia is a geographical subregion of Asia; its northeastern landmass and islands are bounded by the Pacific Ocean. The term Northeast Asia was popularized during the 1930s by American historian and political scienti ...

. * ''Sambucus racemosa'' var. ''melanocarpa'' — Rocky Mountain elder, native to the

Western United States The Western United States (also called the American West, the Far West, and the West) is the region comprising the westernmost states of the United States. As American settlement in the U.S. expanded westward, the meaning of the term ''the Wes ...

and

Western Canada Western Canada, also referred to as the Western provinces, Canadian West or the Western provinces of Canada, and commonly known within Canada as the West, is a Canadian region that includes the four western provinces just north of the Canada� ...

, including the

Rocky Mountain The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...

s and

Sierra Nevada The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primarily ...

. * ''Sambucus racemosa'' subsp. ''pubens'' — American red elder, native to Eastern North America * ''Sambucus racemosa'' subsp. ''racemosa'' — European red elder. * ''Sambucus racemosa'' var. ''racemosa'' — Pacific red elderberry. * ''Sambucus racemosa'' subsp. ''sibirica'' — red elder, native to

Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...

. * ''Sambucus racemosa'' subsp. ''sieboldiana'' — Japanese red elder

Uses

The stems, roots and foliage are

poisonous Poison is a chemical substance that has a detrimental effect to life. The term is used in a wide range of scientific fields and industries, where it is often specifically defined. It may also be applied colloquially or figuratively, with a broa ...

, and the berries can be toxic or cause nausea if eaten raw.

Medicinal plant

It has been used as a traditional

medicinal plant Medicinal plants, also called medicinal herbs, have been discovered and used in traditional medicine practices since prehistoric times. Plants synthesize hundreds of chemical compounds for various functions, including Plant defense against her ...

by Native Americans, including the Bella Coola,

Carrier Carrier may refer to: Entertainment * ''Carrier'' (album), a 2013 album by The Dodos * ''Carrier'' (board game), a South Pacific World War II board game * ''Carrier'' (TV series), a ten-part documentary miniseries that aired on PBS in April 20 ...

Gitksan Gitxsan (also spelled Gitksan) are an Indigenous people in Canada whose home territory comprises most of the area known as the Skeena Country in English (: means "people of" and : means "the River of Mist"). Gitksan territory encompasses approxim ...

Hesquiaht The Hesquiaht are one of the Nuu-chah-nulth peoples of the West Coast of Vancouver Island, British Columbia, Canada. Today the Hesquiaht are governed mostly by the Hesquiaht First Nation band government, though some are in the Tla-O-Qui-Aht First ...

Menominee The Menominee (; mez, omǣqnomenēwak meaning ''"Menominee People"'', also spelled Menomini, derived from the Ojibwe language word for "Wild Rice People"; known as ''Mamaceqtaw'', "the people", in the Menominee language) are a federally recog ...

Northern Paiute Northern may refer to the following: Geography * North, a point in direction * Northern Europe, the northern part or region of Europe * Northern Highland, a region of Wisconsin, United States * Northern Province, Sri Lanka * Northern Range, a r ...

Ojibwa The Ojibwe, Ojibwa, Chippewa, or Saulteaux are an Anishinaabe people in what is currently southern Canada, the northern Midwestern United States, and Northern Plains. According to the U.S. census, in the United States Ojibwe people are one of ...

Paiute Paiute (; also Piute) refers to three non-contiguous groups of indigenous peoples of the Great Basin. Although their languages are related within the Numic group of Uto-Aztecan languages, these three groups do not form a single set. The term "Pai ...

Potawatomi The Potawatomi , also spelled Pottawatomi and Pottawatomie (among many variations), are a Native American people of the western Great Lakes region, upper Mississippi River and Great Plains. They traditionally speak the Potawatomi language, a m ...

, Tlingit, and Haida peoples.University of Michigan at Dearborn: Native American Ethnobotany for '' Sambucus racemosa''
/ref> The uses included as an

emetic Vomiting (also known as emesis and throwing up) is the involuntary, forceful expulsion of the contents of one's stomach through the mouth and sometimes the nose. Vomiting can be the result of ailments like food poisoning, gastroenteri ...

, antidiarrheal, cold and cough remedy, dermatological and gynecological aid, and a

hemostat A hemostat (also called a hemostatic clamp, arterial forceps, or pean after Jules-Émile Péan) is a surgical tool used in many surgical procedures to control bleeding. For this reason, it is common in the initial phases of surgery for the initia ...

Food

The fruits are reportedly safe to eat when cooked, but are potentially poisonous when raw. They were cooked in a variety of recipes by indigenous peoples, including by the

Apache The Apache () are a group of culturally related Native American tribes in the Southwestern United States, which include the Chiricahua, Jicarilla, Lipan, Mescalero, Mimbreño, Ndendahe (Bedonkohe or Mogollon and Nednhi or Carrizaleño an ...

, Bella Coola,

Gitxsan Gitxsan (also spelled Gitksan) are an Indigenous people in Canada whose home territory comprises most of the area known as the Skeena Country in English (: means "people of" and : means "the River of Mist"). Gitksan territory encompasses approxim ...

Gosiute The Goshutes are a tribe of Western Shoshone Native Americans. There are two federally recognized Goshute tribes today: * Confederated Tribes of the Goshute Reservation, located in Nevada and Utah * Skull Valley Band of Goshute Indians of Utah ...

Makah The Makah (; Klallam: ''màq̓áʔa'')Renker, Ann M., and Gunther, Erna (1990). "Makah". In "Northwest Coast", ed. Wayne Suttles. Vol. 7 of ''Handbook of North American Indians'', ed. William C. Sturtevant. Washington, D.C.: Smithsonian Institut ...

, Ojibwa,

Quileute The Quileute , are a Native American people in western Washington state in the United States, currently numbering approximately 2,000. They are a federally recognized tribe: the ''Quileute Tribe of the Quileute Reservation''. The Quileute peop ...

, Skokomish,

Yurok The Yurok (Karuk language: Yurúkvaarar / Yuru Kyara - "downriver Indian; i.e. Yurok Indian") are an Indigenous people from along the Klamath River and Pacific coast, whose homelands are located in present-day California stretching from Trinidad ...

peoples. The fruits are popular with birds, who also distribute the seeds.Pojar, J. & A. MacKinnon. (1994). ''Plants of the Pacific Northwest''. Lone Pine Publishing. The flowers attract

and

Cultivation

''Sambucus racemosa'' is cultivated as an

ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...

, for use as a

or small tree in traditional and

wildlife garden A wildlife garden (or wild garden) is an environment created by a gardener that serves as a sustainable haven for surrounding wildlife. Wildlife gardens contain a variety of habitats that cater to native and local plants, birds, amphibians, rep ...

s, and

natural landscape A natural landscape is the original landscape that exists before it is acted upon by human culture Culture () is an umbrella term which encompasses the social behavior, institutions, and norms found in human societies, as well as the kn ...

design projects.

Cultivars

Cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...

s in the nursery trade include: *''Sambucus racemosa'' 'Black Lace' — burgundy foliage *''Sambucus racemosa'' 'Lemon Lace' — golden yellow and green foliage *''Sambucus racemosa'' 'Lemony Lace' — golden green foliage, with red new growth *''Sambucus racemosa'' 'Sutherland Gold' — green foliage, with bronze new growth: it has received the

Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nort ...

Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...

Images

File:Sambucus_racemosa_1567.JPG, Fruit (

Mount Baker-Snoqualmie National Forest Mount is often used as part of the name of specific mountains, e.g. Mount Everest. Mount or Mounts may also refer to: Places * Mount, Cornwall, a village in Warleggan parish, England * Mount, Perranzabuloe, a hamlet in Perranzabuloe parish, C ...

) File:0 Sambucus racemosa - Vallorcine.JPG, Leaves and buds of the red elderberry File:Sambucus_racemosa_6269.JPG, Inflorescence and foliage File:Sambucus racemosa ies.jpg, Clusters of berries Sambucus racemosa racemosa & melanocarpa range map 3.png, Natural range of ''S. racemosa'' var. ''racemosa'' (green) and var. ''melanocarpa'' (blue) in western North America.

References

External links

* * * * {{Taxonbar, from=Q157276 racemosa Flora of Europe Flora of temperate Asia Flora of Canada Flora of the Eastern United States Flora of the Western United States Flora of California Plants described in 1753 Taxa named by Carl Linnaeus Berries Bird food plants Butterfly food plants Edible plants Garden plants Flora without expected TNC conservation status