Ratufa Bicolor
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The black giant squirrel or Malayan giant squirrel (''Ratufa bicolor'') is a large
tree squirrel Tree squirrels are the members of the squirrel Family (biology), family (Sciuridae) commonly just referred to as "squirrels." They include more than 100 arboreal species native to all continents except Antarctica and Oceania. They do not form a ...
in the genus '' Ratufa'' native to the
Indomalayan The Indomalayan realm is one of the eight biogeographic realms. It extends across most of South and Southeast Asia and into the southern parts of East Asia. Also called the Oriental realm by biogeographers, Indomalaya spreads all over the Indi ...
zootope. It is found in forests from northern
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,
northeast India , native_name_lang = mni , settlement_type = , image_skyline = , image_alt = , image_caption = , motto = , image_map = Northeast india.png , ...
, eastern
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...
,
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainous ...
, southern
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,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
,
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
,
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
,
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,
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand t ...
,
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
, and western
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
(
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
,
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
,
Bali Bali () is a province of Indonesia and the westernmost of the Lesser Sunda Islands. East of Java and west of Lombok, the province includes the island of Bali and a few smaller neighbouring islands, notably Nusa Penida, Nusa Lembongan, and Nu ...
and nearby small islands).


Description

The black giant squirrel is one of the largest species of squirrel in the world. On average, an adult black giant squirrel weighs around , has a head–and–body length of , and the tail is long. The
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
''R. b. condorensis'' of Vietnam's
Côn Sơn Island Côn Sơn ( ), also known as Côn Lôn is the largest island of the Côn Đảo archipelago, off the coast of southern Vietnam.Kelley, p 116 Other names Its French variant Grande-Condore was well-known during the times of French Indochina. Mar ...
averages only in head–and–body length and the tail , but otherwise it resembles the typical subspecies. This species is typically distinctly bicoloured with dark upperparts and pale underparts. The back, top of the head, ears and bushy tail are deep brown to black and the underparts are light
buff Buff or BUFF may refer to: People * Buff (surname), a list of people * Buff (nickname), a list of people * Johnny Buff, ring name of American world champion boxer John Lisky (1888–1955) * Buff Bagwell, a ring name of American professional ...
-coloured. In
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
,
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
and
Bali Bali () is a province of Indonesia and the westernmost of the Lesser Sunda Islands. East of Java and west of Lombok, the province includes the island of Bali and a few smaller neighbouring islands, notably Nusa Penida, Nusa Lembongan, and Nu ...
the hairs of the back and tail are light-tipped, making these sections appear relatively pale (however, the back is still distinctly darker than the underparts). On small islands off Myanmar and in the
Strait of Malacca The Strait of Malacca is a narrow stretch of water, 500 mi (800 km) long and from 40 to 155 mi (65–250 km) wide, between the Malay Peninsula (Peninsular Malaysia) to the northeast and the Indonesian island of Sumatra to the southwest, connec ...
the black giant squirrel has reddish-yellowish underparts.


Habitat

''Ratufa bicolor's''
range Range may refer to: Geography * Range (geographic), a chain of hills or mountains; a somewhat linear, complex mountainous or hilly area (cordillera, sierra) ** Mountain range, a group of mountains bordered by lowlands * Range, a term used to i ...
includes a variety of
bioregions A bioregion is an ecology, ecologically and geographically defined area that is smaller than a biogeographic realm, but larger than an ecoregion or an ecosystem, in the World Wide Fund for Nature classification scheme. There is also an attempt to ...
that all share the commonality of being
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
ed. It ranges in elevation from sea level up to at least , in some of the most rugged land in the world. However, in recent decades, ''R. bicolor's'' habitat has been steadily encroached upon by human settlement, timber harvesting and agriculture, which along with overhunting by human predation in parts of its range, has resulted in a total loss of up to 30% of the population in the past ten years. However, in some places this species is protected from hunting by law or tradition. In
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;;;;; ...
''R. bicolor'' dwells among
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extant ...
and broadleaf forests. In
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
''R. bicolor'' lives in tropical
broadleaf evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, ...
and
semi-evergreen Semi-deciduous or semi-evergreen is a botanical term which refers to plants that lose their foliage for a very short period, when old leaves fall off and new foliage growth is starting. This phenomenon occurs in tropical and sub-tropical woody spec ...
forests, but is rarely seen in coniferous forests. In the
tropical rainforest Tropical rainforests are rainforests that occur in areas of tropical rainforest climate in which there is no dry season – all months have an average precipitation of at least 60 mm – and may also be referred to as ''lowland equatori ...
of the
Malay Peninsula The Malay Peninsula (Malay: ''Semenanjung Tanah Melayu'') is a peninsula in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The area ...
and Indonesia, ''R. bicolor'' is not as abundant as elsewhere in its range, which is probably due to competition from other arboreal species (especially
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
s) for food in the upper forest canopy. Among the better places to sight the black giant squirrel is the
Kaziranga National Park Kaziranga National Park is a national park in the Golaghat and Nagaon districts of the state of Assam, India. The park, which hosts two-thirds of the world's great one-horned rhinoceroses, is a World Heritage Site. According to the census held i ...
in the state of
Assam Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
. Several populations are present in the lower range of the
Neora Valley National Park Neora Valley National Park is a national park in Kalimpong district, West Bengal, India that was established in 1986. It spreads over an area of , and is a rich biological zone in eastern India. It is the land of the red panda in the pristine und ...
, Kalimpong, India. A recent study from India showed precipitation during the wettest month of a year is one of the major contributing factor for habitat preference of ''R. bicolor'', along with land use, and vegetation. The species also may be found way beyond 1400 meters in several places. More than 20% of the presence records of the species has been observed above 1500 meters and up to 2700 meters in India. It has been predicted through study that by the year 2050, this species may loose more than 97% of its present suitable habitat due to climate change in India.


Behavior

''R. bicolor'' is diurnal and
arboreal Arboreal locomotion is the Animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. Th ...
, but sometimes climbs down from the forest
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an a ...
to feed on the ground. The black giant squirrel rarely enters
plantation A plantation is an agricultural estate, generally centered on a plantation house, meant for farming that specializes in cash crops, usually mainly planted with a single crop, with perhaps ancillary areas for vegetables for eating and so on. The ...
s or settlements, preferring the wild forest. Its diet consists of seeds, pine cones, fruits, and leaves. It is primarily solitary, and has a litter of from 1 to 2 young, which it raises in a
drey A drey is the nest of a tree squirrel, flying squirrel or ringtail possum. Dreys are usually built of twigs, dry leaves, and grass, and typically assembled in the forks of a tall tree. They are sometimes referred to as "drey nests" to disting ...
(or nest), often located within a hollow space of a tree.


Taxonomy

Further study is required to determine whether ''Ratufa bicolor'' actually represents several similar species. The table below lists the ten recognized subspecies of ''Ratufa bicolor'', along with any synonyms associated with each subspecies:


Gallery

Black giant squirrel (Malayan giant squirrel), Ratufa bicolor - Kaeng Krachan National Park.webm, Fro
Kaeng Krachan National Park
Thailand


References


Bibliography

*Francis, Charles M., Priscilla Barrett. ''A field guide to the mammals of South-East Asia.'' London: New Holland, 2008. , OCLC: 190967851. *Lekhakun, Bunsong, Jeffrey A. McNeely. ''Mammals of Thailand.'' Bangkok: Association for the Conservation of Wildlife, 1977. OCLC: 3953763. *Nowak, Ronald M. ''Walker’s mammals of the world.'' Baltimore: Johns Hopkins University Press, 1999. , OCLC: 39045218. Chapter: "Sciuridae: squirrels, chipmunks, marmots, and prairie dogs" in volume two.


External links



- Ecology Asia page about this species, with beautiful photos and description. {{Taxonbar, from=Q917916 Ratufa Rodents of India Rodents of Bangladesh Mammals of Nepal Mammals of Bhutan Rodents of China Rodents of Myanmar Rodents of Laos Rodents of Thailand Rodents of Cambodia Rodents of Vietnam Rodents of Indonesia Rodents of Malaysia Mammals described in 1778 Articles containing video clips Taxonomy articles created by Polbot Taxa named by Anders Sparrman