botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...

, ramification is the divergence of the

stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushro ...

and limbs of a

plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...

into smaller ones, i.e., trunk into

branch A branch, sometimes called a ramus in botany, is a woody structural member connected to the central trunk (botany), trunk of a tree (or sometimes a shrub). Large branches are known as boughs and small branches are known as twigs. The term '' ...

es, branches into increasingly smaller branches, and so on. Gardeners stimulate the process of ramification through

pruning Pruning is a horticultural, arboricultural, and silvicultural practice involving the selective removal of certain parts of a plant, such as branches, buds, or roots. The practice entails the ''targeted'' removal of diseased, damaged, dead, ...

, thereby making

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

s, and other plants bushier and denser. Short

internodes A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, stores nutrien ...

(the section of stem between nodes, i.e., areas where

leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

are produced) help increase ramification in those plants that form branches at these nodes. Long internodes (which may be the result of over-watering, the over-use of

fertilizer A fertilizer (American English) or fertiliser (British English; see spelling differences) is any material of natural or synthetic origin that is applied to soil or to plant tissues to supply plant nutrients. Fertilizers may be distinct from ...

, or a seasonal "growth spurt") decrease a gardener's ability to induce ramification in a plant. A high degree of ramification is essential for the creation of

topiary Topiary is the horticultural practice of training perennial plants by clipping the foliage and twigs of trees, shrubs and subshrubs to develop and maintain clearly defined shapes, whether geometric or fanciful. The term also refers to plants w ...

as it enables the topiary artist to carve a bush or

hedge A hedge or hedgerow is a line of closely spaced shrubs and sometimes trees, planted and trained to form a barrier or to mark the boundary of an area, such as between neighbouring properties. Hedges that are used to separate a road from adjoini ...

into a shape with an even surface. Ramification is also essential to practitioners of the art of bonsai as it helps re-create the form and habit of a full-size tree in a small tree grown in a container. The pruning practices of

coppicing Coppicing is a traditional method of woodland management Forest management is a branch of forestry concerned with overall administrative, legal, economic, and social aspects, as well as scientific and technical aspects, such as silviculture, pro ...

and

pollarding Pollarding is a pruning system involving the removal of the upper branches of a tree, which promotes the growth of a dense head of foliage and branches. In ancient Rome, Propertius mentioned pollarding during the 1st century BCE. The practice oc ...

induce ramification by removing most of a tree's mass above the root.

Fruit tree pruning Fruit tree pruning is the cutting and removing of selected parts of a fruit tree. It spans a number of horticultural techniques. Pruning often means cutting branches back, sometimes removing smaller limbs entirely. It may also mean removal of y ...

increases the yield of

orchards An orchard is an intentional plantation of trees or shrubs that is maintained for food production. Orchards comprise fruit- or nut-producing trees which are generally grown for commercial production. Orchards are also sometimes a feature of lar ...

by inducing ramification and thereby creating many vigorous, fruitful branches in the place of a few less-fruitful ones.

External links

Annals of Botany 91: 559-569, 2003

Plant physiology Horticulture Plant anatomy Plant morphology {{botany-stub