''Quinchamalium'' is a genus of

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

in the family

Schoepfiaceae Schoepfiaceae is a family of flowering plants recognized in the APG III system of 2009. The family was previously only recognized by few taxonomists; the plants in question usually being assigned to family Olacaceae and Santalaceae. The genus '' ...

, with a single species ''Quinchamalium chilense'', native to

Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...

and

Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...

. Depending on the latitude, it can be found from sea level to 3,800 m in altitude.

Etymology

It is locally called ''quinchamalí''. The word ''kincha'' translates from

Quechua Quechua may refer to: *Quechua people, several indigenous ethnic groups in South America, especially in Peru *Quechuan languages, a Native South American language family spoken primarily in the Andes, derived from a common ancestral language **So ...

as 'hedge of sticks', as often made from dead old

quinoa Quinoa (''Chenopodium quinoa''; , from Quechua ' or ') is a flowering plant in the amaranth family. It is a herbaceous annual plant grown as a crop primarily for its edible seeds; the seeds are rich in protein, dietary fiber, B vitamins, and ...

plants (see

quincha Quincha is a traditional construction system that uses, fundamentally, wood and cane or giant reed forming an earthquake-proof framework that is covered in mud and plaster. History Quincha is a Spanish term widely known in Latin America, borrow ...

Description

''Quinchamalium chilense'' is a

herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...

hemiparasitic A parasitic plant is a plant that derives some or all of its nutritional requirements from another living plant. They make up about 1% of angiosperms and are found in almost every biome. All parasitic plants develop a specialized organ called the ...

perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

with a

rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...

, from which smaller rootlets with

haustoria In botany and mycology, a haustorium (plural haustoria) is a rootlike structure that grows into or around another structure to absorb water or nutrients. For example, in mistletoe or members of the broomrape family, the structure penetrates t ...

grow. From the rhizome a number of rarely-branching stems grow each year. These can be decumbent (creeping over the ground) or rise to over 2 metres in height. The flowers are produced in a cluster at the end of the stems, they are pentamerous and usually orangey-yellow in colour. It is extremely variable in height and flower size and colour, so much so that at one time 21 species were recognised, but in 2015 these were all reduced to

synonymy A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...

with ''Q. chilense''. It is heterostylous; some individuals bear 'thrum flowers', flowers which have short styles. This is somewhat correlated with altitude and cold. The leaves are also variable among individuals, they can be narrow or broader, the plants with more narrow leaves correlated with higher temperatures. The size of the plant appears correlated with annual precipitation. This species has rather bizarre embryos, these appear to be parasitic upon their mother plant before even being

pollinated Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, ...

. As the embryo develops in the ovule, the two

synergid In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the sporangium, megasporangium), ...

cells elongate into tubes which penetrate through the

micropyle Micropyle may refer to: * Micropyle (botany) In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (o ...

and grow about a third up the length of the

style Style is a manner of doing or presenting things and may refer to: * Architectural style, the features that make a building or structure historically identifiable * Design, the process of creating something * Fashion, a prevailing mode of clothing ...

where they form haustoria, organs for sucking nutrients from a host. It also creates a lateral caecum (pouch) close to the other end of the embryo, the chalazal end, which extends in the basal region of funiculus. This rare situation is also known to occur in pampas grasses of the genus ''

Cortaderia ''Cortaderia'' is a genus of South American and Central American plants in the Poaceae grass family. Etymology The common name pampas grass, though strictly referring to ''C. selloana'', is frequently applied to all species in the genus (and so ...

'' and some species of the ''

Olax ''Olax'' is a plant genus in the family Olacaceae. The name derives from the Latin, ''olax'' (malodorous), and refers to the unpleasant scent of some of the ''Olax'' species. ''Olax'' is an Old World genus represented by several climbers, so ...

'' genus. As noted above, the individuals of this species can have 'thrum flowers' or 'pin flowers', which differ in form and length of the styles, and often have both. In most heterostylous plants, this is associated with self-incompatibility, but in ''Q. chilense'' both flower forms are bisexual and self-fertile. Nonetheless, the smaller pin flowers appear to create more pollen and have a higher seed set. Why these plants have different sized flowers is probably caused by some sort of pollinator relationship, but the gist of it is still unknown.

Ecology

''Quinchamalium chilense'' is a dominant plant species along with '' Festuca scabriuscula'', '' Poa obvalata'' and '' Adesmia emarginata'' in a type of

Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and gl ...

steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate grasslands, ...

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

with scattered trees of ''

Araucaria araucana ''Araucaria araucana'' (commonly called the monkey puzzle tree, monkey tail tree, piñonero, pewen or Chilean pine) is an evergreen tree growing to a trunk diameter of 1–1.5 m (3–5 ft) and a height of 30–40 m (100–130 ft). ...

'' found along the upper course of the

Biobío River The Biobío River (also known as Bío Bío or Bio-Bio) is the second largest river in Chile. It originates from Icalma and Galletué lakes in the Andes and flows 380 km to the Gulf of Arauco (in Spanish) on the Pacific Ocean. The major tri ...

in the

Araucanía Region The Araucanía ( ), La Araucanía Region ( es, Región de La Araucanía ) is one of Chile's 16 first-order administrative divisions, and comprises two provinces: Malleco in the north and Cautín in the south. Its capital and largest city is Te ...

of Chile. This is a cold region at 1,500m altitude, with a dry season of one to two months, sometimes heavy rains, and temperatures which can fall below freezing. The fly '' Mitrodetus dentitarsis'' is known to visit the flowers in Chile, among other plants it visits.

Uses

In southern Chile the plant is used in

folk medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...

References

{{Taxonbar, from1=Q9065330, from2=Q15551662 Schoepfiaceae Monotypic Santalales genera