Quadrat (Geometrie)
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A quadrat is a frame used in
ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...
,
geography Geography (from Greek: , ''geographia''. Combination of Greek words ‘Geo’ (The Earth) and ‘Graphien’ (to describe), literally "earth description") is a field of science devoted to the study of the lands, features, inhabitants, and ...
, and
biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary i ...
to isolate a standard unit of area for study of the distribution of an item over a large area. Quadrats are traditionally square, but modern quadrats can be rectangular, circular, or irregular. A quadrat is suitable for sampling or observing
plants Plants are predominantly Photosynthesis, photosynthetic eukaryotes of the Kingdom (biology), kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all curr ...
, slow-moving animals, and some aquatic organisms. A photo-quadrat is a photographic record of the area framed by a quadrat. It may use a physical frame to indicate the area, or may rely on fixed camera distance and lens
field of view The field of view (FoV) is the extent of the observable world that is seen at any given moment. In the case of optical instruments or sensors it is a solid angle through which a detector is sensitive to electromagnetic radiation. Human ...
to automatically cover the specified area of substrate. Parallel laser pointers mounted on the camera can also be used as scale indicators. The photo is taken perpendicular to the surface, or as close as possible to perpendicular for uneven surfaces.


History

The systematic use of quadrats was developed by the pioneering plant ecologists R. Pound and F. E. Clements between 1898 and 1900. The method was then swiftly applied for many purposes in
ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...
, such as the study of
plant succession Ecological succession is the process of change in the species structure of an ecological community over time. The time scale can be decades (for example, after a wildfire) or more or less. Bacteria allows for the cycling of nutrients such as ca ...
. Botanists and ecologists such as
Arthur Tansley Sir Arthur George Tansley FLS, FRS (15 August 1871 – 25 November 1955) was an English botanist and a pioneer in the science of ecology. Educated at Highgate School, University College London and Trinity College, Cambridge, Tansley taught a ...
soon took up and modified the method. The ecologist J. E. Weaver applied the use of quadrats to the teaching of ecology in 1918.


Method

A quadrat can be used by researchers to methodically count organisms within a smaller, representative area in order to extrapolate to a larger
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
when comprehensive sampling is impossible or not practical. The quadrat's size corresponds to the size of the organism being sampled and the overall sampling area. To avoid selection bias, researchers will randomly distribute quadrats throughout the sampling area. For long-term studies, the same quadrats can be revisited after their initial sampling. Methods of precisely relocating the area of study vary widely in accuracy and include measurement from nearby permanent markers, use of
total station A total station (TS) or total station theodolite (TST) is an electronic/optical instrument used for surveying and building construction. It is an electronic transit theodolite integrated with electronic distance measurement (EDM) to measure ...
theodolites, consumer-grade
GPS The Global Positioning System (GPS), originally Navstar GPS, is a satellite-based radionavigation system owned by the United States government and operated by the United States Space Force. It is one of the global navigation satellite sy ...
, and
differential GPS Differential Global Positioning Systems (DGPSs) supplement and enhance the positional data available from global navigation satellite systems (GNSSs). A DGPS for GPS can increase accuracy by about a thousandfold, from approximately to . DGPSs c ...
.


See also

*
Stand level modelling A forest stand is a contiguous community of trees sufficiently uniform in composition, structure, age, size, class, distribution, spatial arrangement, site quality, condition, or location to distinguish it from adjacent communities. A forest is ...


References

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External links


Ecological Sampling Methods
Ecology terminology Environmental statistics Environmental Sampling Equipment