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''Prunus virginiana'', commonly called bitter-berry, chokecherry, Virginia bird cherry, and western chokecherry (also black chokecherry for ''P. virginiana'' var. ''demissa''), is a species of bird cherry (''Prunus'' subgenus ''Padus'') native to North America.


Description

Chokecherry is a suckering shrub or small
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
growing to tall, rarely to and exceptionally with a trunk as thick as . The leaves are
oval An oval () is a closed curve in a plane which resembles the outline of an egg. The term is not very specific, but in some areas (projective geometry, technical drawing, etc.) it is given a more precise definition, which may include either one ...
, long and wide, with a serrated margin. The stems rarely exceed in length. The
flowers A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...
are produced in racemes long in late spring (well after leaf emergence), eventually growing up to 15 cm. They are across. The fruits ( drupes) are about in diameter, range in color from bright red to black, and possess a very
astringent An astringent (sometimes called adstringent) is a chemical that shrinks or constricts body tissues. The word derives from the Latin ''adstringere'', which means "to bind fast". Calamine lotion, witch hazel, and yerba mansa, a Californian pla ...
taste, being both somewhat sour and somewhat bitter. They get darker and marginally sweeter as they ripen. They each contain a large stone.


Chemistry

Chokecherries are very high in antioxidant
pigment A pigment is a colored material that is completely or nearly insoluble in water. In contrast, dyes are typically soluble, at least at some stage in their use. Generally dyes are often organic compounds whereas pigments are often inorganic compou ...
compounds, such as
anthocyanins Anthocyanins (), also called anthocyans, are water-soluble vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart gave the name Anthokyan to a chemical com ...
. They share this property with
chokeberries ''Aronia'' is a genus of deciduous shrubs, the chokeberries, in the family Rosaceae native to eastern North America and most commonly found in wet woods and swamps. The genus Aronia is considered to have 3 species. The most common and wide ...
, further contributing to confusion.


Similar species

The chokecherry is closely related to the
black cherry ''Prunus serotina'', commonly called black cherry,World Economic Plants: A Standard Reference, Second Edition'. CRC Press; 19 April 2016. . p. 833–. wild black cherry, rum cherry, or mountain black cherry, is a deciduous tree or shrub of the ...
(''Prunus serotina'') of eastern North America, which can reach tall, have larger leaves, and darker fruit. The chokecherry leaf has a finely serrated margin and is dark green above with a paler underside, while the black cherry leaf has numerous blunt edges along its margin and is dark green and smooth.


Taxonomy

The name chokecherry is also used for the related Manchurian cherry or Amur chokecherry (''Prunus maackii'').


Varieties

* ''Prunus virginiana'' var. ''virginiana'' (eastern chokecherry) * ''Prunus virginiana'' var. ''demissa'' (Nutt. ex Torr. & A.Gray) Torr. (western chokecherry) * ''Prunus virginiana'' var. ''melanocarpa'' (A.Nelson) Sarg.


Distribution

The natural historic range of ''P. virginiana'' includes most of Canada (including Northwest Territories, but excluding
Yukon Yukon (; ; formerly called Yukon Territory and also referred to as the Yukon) is the smallest and westernmost of Canada's three territories. It also is the second-least populated province or territory in Canada, with a population of 43,964 as ...
, Nunavut, and Labrador), most of the United States (including
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S. ...
, but excluding some states in the
Southeast The points of the compass are a set of horizontal, radially arrayed compass directions (or azimuths) used in navigation and cartography. A compass rose is primarily composed of four cardinal directions—north, east, south, and west—each sepa ...
), and northern Mexico ( Sonora, Chihuahua,
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
, Durango,
Zacatecas , image_map = Zacatecas in Mexico (location map scheme).svg , map_caption = State of Zacatecas within Mexico , coordinates = , coor_pinpoint = , coordinates_footnotes = , subdivision_type ...
, Coahuila, and Nuevo León).


Ecology

The wild chokecherry is often considered a pest, as it is a host for the
tent caterpillar Tent caterpillars are moderately sized caterpillars, or moth larvae, belonging to the genus ''Malacosoma'' in the family Lasiocampidae. Twenty-six species have been described, six of which occur in North America and the rest in Eurasia. Some s ...
, a threat to other fruit plants. It is also a larval host to the black-waved flannel moth, the blinded sphinx, the cecropia moth, the coral hairstreak, the cynthia moth, the elm sphinx, Glover's silkmoth, the hummingbird clearwing moth, the imperial moth, the Io moth, the polyphemus moth, the promethea moth, the red-spotted purple, the small-eyed sphinx, the spring azure, the striped hairstreak, the tiger swallowtail, the twin-spotted sphinx, and Weidemeyer's admiral. Many wildlife, including birds and game animals, eat the berries.
Moose The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult ma ...
, elk, mountain sheep,
deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the re ...
and rabbits eat the foliage, twigs, leaves, and buds. Deer and elk sometimes browse the twigs profusely, not letting the plant grow above knee height. The leaves serve as food for caterpillars of various Lepidoptera.


Cultivation

The chokecherry has a number of
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s. 'Canada Red' and 'Schubert' have leaves that mature to purple and turn orange and red in the autumn. 'Goertz' has a nonastringent, so palatable, fruit. Research at the
University of Saskatchewan A university () is an institution of higher (or tertiary) education and research which awards academic degrees in several academic disciplines. Universities typically offer both undergraduate and postgraduate programs. In the United States, ...
seeks to find or create new cultivars to increase production and processing.


Toxicity

The stone of the fruit is poisonous. Chokecherry is toxic to
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
s, moose,
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult ma ...
,
goats The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of th ...
, deer, and other animals with segmented stomachs (
rumen The rumen, also known as a paunch, is the largest stomach compartment in ruminants and the larger part of the reticulorumen, which is the first chamber in the alimentary canal of ruminant animals. The rumen's microbial favoring environment al ...
s), especially after the leaves have wilted (such as after a frost or after branches have been broken), because wilting releases
cyanide Cyanide is a naturally occurring, rapidly acting, toxic chemical that can exist in many different forms. In chemistry, a cyanide () is a chemical compound that contains a functional group. This group, known as the cyano group, consists of ...
and makes the plant sweet. The leaves themselves being poisonous, about of foliage can be fatal. Symptoms of a horse that has been poisoned include heavy breathing, agitation, and weakness.


Uses

For many Native American tribes of the Northern Rockies,
Northern Plains The Great Plains (french: Grandes Plaines), sometimes simply "the Plains", is a broad expanse of flatland in North America. It is located west of the Mississippi River and east of the Rocky Mountains, much of it covered in prairie, steppe, and ...
, and
boreal forest Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruc ...
region of Canada and the United States, chokecherries are the most important fruit in their traditional diets and are part of
pemmican Pemmican (also pemican in older sources) is a mixture of tallow, dried meat, and sometimes dried berries. A calorie-rich food, it can be used as a key component in prepared meals or eaten raw. Historically, it was an important part of indigenou ...
, a staple traditional food. The bark of chokecherry root is made into an asperous-textured concoction used to ward off or treat colds, fever and stomach maladies by Native Americans. The inner bark of the chokecherry, as well as
red osier dogwood ''Cornus sericea'', the red osier or red-osier dogwood, is a species of flowering plant in the family Cornaceae, native to much of North America. It has sometimes been considered a synonym of the Asian species '' Cornus alba''. Other names inclu ...
, or
alder Alders are trees comprising the genus ''Alnus'' in the birch family Betulaceae. The genus comprises about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few sp ...
, is also used by some tribes in ceremonial smoking mixtures, known as
kinnikinnick Kinnikinnick is a Native American and First Nations herbal smoking mixture, made from a traditional combination of leaves or barks. Recipes for the mixture vary, as do the uses, from social, to spiritual to medicinal. Etymology The term "kinni ...
. The chokecherry fruit can be eaten when fully ripe, but otherwise contains a toxin. The fruit can be used to make jam or syrup, but the bitter nature of the fruit requires sugar to sweeten the preserves. The Plains Indians pound up the whole fruits—including the toxic pits—in a mortar, from which they made sun-baked cakes. Chokecherry is also used to craft wine in the Western United States, mainly in the Dakotas and
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
, as well as in
Manitoba , image_map = Manitoba in Canada 2.svg , map_alt = Map showing Manitoba's location in the centre of Southern Canada , Label_map = yes , coordinates = , capital = Winn ...
, Canada.


Culture

In 2007,
North Dakota North Dakota () is a U.S. state in the Upper Midwest, named after the indigenous Dakota Sioux. North Dakota is bordered by the Canadian provinces of Saskatchewan and Manitoba to the north and by the U.S. states of Minnesota to the east, So ...
governor
John Hoeven John Henry Hoeven III ( ; born March 13, 1957) is an American banker and politician serving as the senior U.S. senator from North Dakota, a seat he has held since 2011. A member of the Republican Party, Hoeven served as the 31st governor of Nort ...
signed a bill naming the chokecherry the state's official fruit, in part because its remains have been found at more archeological sites in the Dakotas than anywhere else.


See also

* Choke pear


References


External links

*
North Dakota State University Agriculture, Chokecherry

Nutrition Facts for Chokecherries

Flora of Pennsylvania
{{Taxonbar, from=Q162829 Bird cherries virginiana Flora of North America Edible nuts and seeds Garden plants of North America Plants used in Native American cuisine Plants used in traditional Native American medicine Plants described in 1753 Taxa named by Carl Linnaeus