Prumnopitys
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''Prumnopitys'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
s belonging to the family
Podocarpaceae Podocarpaceae is a large family of mainly Southern Hemisphere conifers, known in English as podocarps, comprising about 156 species of evergreen trees and shrubs.James E. Eckenwalder. 2009. ''Conifers of the World''. Portland, Oregon: Timber Pre ...
. The nine recognized
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of ''Prumnopitys'' are densely branched,
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s up to 40 metres in height.


Etymology

The name ''Prumnopitys'' comes from the Ancient Greek ' ( ‘hindmost’) and ' ( ‘pine’), referring to the
resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on natu ...
duct being behind the
midrib This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
.


Description

The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are similar to those of the
yew Yew is a common name given to various species of trees. It is most prominently given to any of various coniferous trees and shrubs in the genus ''Taxus'': * European yew or common yew (''Taxus baccata'') * Pacific yew or western yew (''Taxus br ...
, strap-shaped, 1–4 cm long and 2–3 mm broad, with a soft texture; they are green above, and with two blue-green
stomata In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is bor ...
l bands below. The
seed cones A conifer cone (in formal botanical usage: strobilus, plural strobili) is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in conifers an ...
are highly modified, reduced to a central stem 1–5 cm long bearing several scales; from one to five scales are fertile, each with a single
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
surrounded by fleshy scale tissue, resembling a
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
. These berry-like cone scales are eaten by
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s which then disperse the seeds in their droppings.


Distribution

The species are distributed on both sides of the Pacific, in eastern
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
,
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, and
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
, and along the mountain ranges of western
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
from
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
to
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
and
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
. This distribution indicates the origins of ''Prumnopitys'' in the
Antarctic flora Antarctic flora are a distinct community of vascular plants which evolved millions of years ago on the supercontinent of Gondwana. Presently, species of Antarctica flora reside on several now separated areas of the Southern Hemisphere, includin ...
, which evolved from the humid temperate flora of southern
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
, an ancient supercontinent.


Taxonomy

Although the genus ''Prumnopitys'' was first described in 1861, it was only from 1978 that it was widely distinguished as distinct from the allied genus ''
Podocarpus ''Podocarpus'' () is a genus of conifers, the most numerous and widely distributed of the podocarp family, the Podocarpaceae. The name comes from Greek πούς (poús, “foot”) + καρπός (karpós, “fruit”). ''Podocarpus'' species ...
'', despite the marked differences in cone development with different parts of the cone structure becoming fleshy and berry-like. Many older texts still have the species listed under ''Podocarpus''. The Chilean species for which the correct scientific name is ''
Prumnopitys andina ''Prumnopitys andina'', the lleuque or Chilean plum yew,Muñoz-Schick 1999); however this name is illegitimate (Mill & Quinn 2001). Several species of ''Prumnopitys'' are used for timber, though as they are slow-growing, supplies are very limited and over-cutting has led to some having an unfavourable
conservation status The conservation status of a group of organisms (for instance, a species) indicates whether the group still exists and how likely the group is to become extinct in the near future. Many factors are taken into account when assessing conservation ...
.


References


Gymnosperm Database: ''Prumnopitys''
* de Laubenfels, D. J. 1978. The genus ''Prumnopitys'' (Podocarpaceae). ''Blumea'' 24: 189-190. * de Laubenfels, D. J. 1988. Coniferales. in ''Flora Malesiana'', Series I, 10: 337-453. Dordrecht: Kluwer Academic. * Molloy, B. P. J. & Muñoz-Schick, M. 1999. The correct name for the Chilean conifer Lleuque (Podocarpaceae). ''New Zealand J. Bot.'' 37: 189–193. Availabl
online
(pdf file). * Mill, R. R. & Quinn, C. J. 2001. Prumnopitys andina reinstated as the correct name for 'lleuque', the Chilean conifer recently renamed P. spicata (Podocarpaceae). ''Taxon'' 50: 1143 - 1154

{{Taxonbar, from=Q138688 Podocarpaceae genera Dioecious plants