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The prostomium (From Ancient Greek, meaning "before the mouth"; plural: prostomia; sometimes also called the "acron") is the cephalized first body segment in an
annelid The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecolo ...
worm's body at the anterior end. It is in front of (but does not include) the mouth, being usually a small shelf- or lip-like extension over the dorsal side of the mouth. The prostomium together with the
peristomium The peristomium is the first true body segment in an annelid worm's body in the anterior end. It is directly behind the prostomium and contains the mouth, tentacular cirri, and sometimes feeding palps, which may instead occur on the prostomium. If ...
, which includes the mouth and pharynx, make up the annelid head.


Description

The prostomium is part of the head and holds at least part of the brain and often bears sensory structures such as the eyes, antennae and palps. It may function like a kind of overlip when the animal is feeding. The prostomium bears many important taxonomic characters and its shape and composition are important for annelid systematics. In addition to the eyes, antennae and palps, the prostomium can possess appendages such as
tentacle In zoology, a tentacle is a flexible, mobile, and elongated organ present in some species of animals, most of them invertebrates. In animal anatomy, tentacles usually occur in one or more pairs. Anatomically, the tentacles of animals work main ...
s or cirri. Moreover, some
polychaete Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are ma ...
prostomia have a posterior extension or ridge with sensory function, called a caruncle. Another sensory organ, the nuchal organ (or a variation, the nuchal epaulette), is a
chemosensory A chemoreceptor, also known as chemosensor, is a specialized sensory receptor which transduces a chemical substance (endogenous or induced) to generate a biological signal. This signal may be in the form of an action potential, if the chemorecept ...
, ciliated pit or groove at the posterior end of the prostomium.


Appendages

There may be three antennae present: a pair of lateral antennae and one median antennae. These are most commonly simple, tapering structures, but they may also be articulated or ornamented. There are different arrangements of insertion of the lateral and median antennae, with the lateral antenna sometimes inserted ventrally to the median antennae or vice versa, or they may be inserted on the same plane. In some taxa, the lateral antennae may be missing, leaving only a lone median antenna. The palps are paired and innervated structures that tend to be located ventrally and laterally. They are often associated with the mouth and may have a feeding or sensory function. Tentacular cirri can be any sort of elongated, forward (anterior) facing cirri and can occasionally be found on the prostomium.


See also

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Peristomium The peristomium is the first true body segment in an annelid worm's body in the anterior end. It is directly behind the prostomium and contains the mouth, tentacular cirri, and sometimes feeding palps, which may instead occur on the prostomium. If ...
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Pygidium The pygidium (plural pygidia) is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is compo ...


References

{{reflist Annelid anatomy