Progonocimicidae
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The Progonocimicidae are an extinct
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
true bugs Hemiptera (; ) is an order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from to aroun ...
in the suborder
Coleorrhyncha Coleorrhyncha or Peloridiomorpha, also known as moss bugs or beetle bugs, are a suborder of Hemiptera and represent an ancient lineage of moss-feeding insects. They show some similarities to the Heteroptera but have been considered distinct. It ha ...
. Progonocimicidae fossils have been found in Europe, Asia, Australia, and South America. Families preserved in the fossil record of Coleorrhyncha include the early Progonocimicidae, and the later Karabasiidae and Hoploridiidae. The only extant family in Coleorrhyncha is
Peloridiidae The Peloridiidae or moss bugs are a family of true bugs, comprising eighteen genera and thirty-four species. They are small, ranging in length from 2 to 4 mm, rarely seen, peculiarly lumpy, flattened bugs found in Patagonia (Argentina and Chi ...
. The family is thought to have evolved from the Permian Ingruidae.


Genera

These 27 genera belong to the family Progonocimicidae: * †subfamily Cicadocorinae Bekker-Migdisova 1958 **† '' Absoluta'' Becker-Migdisova, 1962 Dzhil Formation, Kyrgyzstan,
Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (My ...
** † '' Archicercopis'' Handlirsch, 1939 Green Series, Germany,
Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toarcian ...
** † '' Cicadocoris'' Becker-Migdisova, 1958 Dzhil Formation, Kyrgyzstan, Hettangian
Abasheva Formation Abashev (russian: Аба́шев; masculine) or Abasheva (; feminine) is a Russian last name.Ganzhina, p. 10 Variants of this last name include Abashenko (), Abashin/Abashina (/), Abashkin/Abashkina (/), Abashichev/Abashicheva (/), Abashkov/A ...
, Russia,
Pliensbachian The Pliensbachian is an age of the geologic timescale and stage in the stratigraphic column. It is part of the Early or Lower Jurassic Epoch or Series and spans the time between 190.8 ± 1.5 Ma and 182.7 ± 1.5 Ma (million years ago). The Plien ...
, Sagul Formation, Kyrgyzstan, Toarcian,
Itat Formation The Itat Formation (Russian: итатская свита) is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from t ...
, Russia,
Bajocian In the geologic timescale, the Bajocian is an age and stage in the Middle Jurassic. It lasted from approximately 170.3 Ma to around 168.3 Ma (million years ago). The Bajocian Age succeeds the Aalenian Age and precedes the Bathonian Age. Stratig ...
/
Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Strat ...
Daohugou The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerates ...
, China,
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
Ichetuy Formation, Russia, Oxfordian,
Kalgan Formation Zhangjiakou (; ; ) also known as Kalgan and by several other names, is a prefecture-level city in northwestern Hebei province in Northern China, bordering Beijing to the southeast, Inner Mongolia to the north and west, and Shanxi to the southw ...
, Russia,
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
**†'' Gakasha'' Jiang et al. 2018
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...
, Myanmar,
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
**† '' Ilahulgabalus'' Szwedo, Azar & Ziade, 2011
Lebanese amber Lebanese amber is fossilized resin found in Lebanon and southwest Syria. It dates back approximately 130-125 million years to the Barremian of the Early Cretaceous. It formed on what was then the northern coast of Gondwana, believed to be a tropi ...
,
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
**† '' Ildavia'' Popov, 1993
Weald Clay Weald Clay or the Weald Clay Formation is a Lower Cretaceous sedimentary rock unit underlying areas of South East England, between the North and South Downs, in an area called the Weald Basin. It is the uppermost unit of the Wealden Group of ro ...
, United Kingdom,
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.9 ± 2 Ma and 129.4 ± 1.5 Ma (million years ago). The Hauterivian is preceded by the Va ...
**†'' Mesocimex'' Hong 1983 Daohugou,
Haifanggou Formation The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerates ...
, China, Callovian **†'' Olgamartynovia'' Becker-Migdisova 1958 Dzhil Formation, Kyrgyzstan, Hettangian Abasheva Formation, Russia, Pliensbachian, Sagul Formation, Kyrgyzstan, Toarcian, Itat Formation, Russia, Bajocian/Bathonian Daohugou, China, Callovian Ichetuy Formation, Russia, Oxfordian, Kalgan Formation, Russia, Kimmeridgian **† '' Onokhoia'' Popov, 1988 Godymboyskaya Formation, Russia,
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
**† '' Valdiscytina'' Popov, 1993 Weald Clay, United Kingdom, Hauterivian ** † '' Yuripopovia'' Jarzembowski, 1991 Weald Clay, United Kingdom, Hauterivian,
Vectis Formation The Vectis Formation is a geological formation on the Isle of Wight and Swanage, England whose strata were formed in the lowermost Aptian, approximately 125 million years ago."Magnetostratigraphy of the Lower Cretaceous Vectis Formation (Wealden ...
, United Kingdom, Aptian
*† '' Actinoscytina'' Tillyard, 1926 Croudace Bay Formation, Australia,
Changhsingian In the geologic time scale, the Changhsingian or Changxingian is the latest age or uppermost stage of the Permian. It is also the upper or latest of two subdivisions of the Lopingian Epoch or Series. The Changhsingian lasted from to 251.902 mill ...
* † '' Eocercopis'' Handlirsch, 1939 Green Series, Germany, Toarcian * † '' Heterojassus'' Evans, 1961 Mount Crosby Insect Bed, Australia,
Norian The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic defi ...
* † '' Heteronella'' Evans, 1961 Mount Crosby Insect Bed, Australia, Norian * † '' Heteroscytina'' Evans, 1956 * † '' Hexascytina'' Wootton, 1963 Mount Crosby Insect Bed, Australia, Norian * † '' Indutionomarus'' Szwedo, 2011 Bascharage, Luxembourg, Toarcian * † '' Microscytinella'' Wootton, 1963 Mount Crosby Insect Bed, Australia, Norian * † '' Ovicimex'' Hong & Wang, 1990
Laiyang Formation Laiyang city () is a county-level city within Yantai bordering Qingdao, located in the middle of the Shandong Peninsula, in Shandong province, China. The majority (70%) of its population are farmers and it is famous for producing the Laiyang pe ...
, China, Aptian
* † '' Pelorisca'' Popov & Shcherbakov, 1991
Madygen Formation The Madygen Formation ( Russian: Madygen Svita) is a Late Triassic (Carnian) geologic formation and Lagerstätte in the Batken and Osh Regions of western Kyrgyzstan, with minor outcrops in neighboring Tajikistan and Uzbekistan. The conglomerates, ...
, Kyrgyzstan,
Ladinian The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic). ...
* † '' Platyscytinella'' Evans, 1956 Mount Crosby Insect Bed, Australia, Norian * † '' Popovigocimex'' Martins-Neto & Galego, 2003 Los Rastros Formation, Argentina,
Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by t ...
* † '' Popovus'' Özdikmen & Demir, 2007 Gurvan-Eren Formation, Mongolia, Aptian * † '' Progonocimex'' Handlirsch, 1906 Green Series, Germany, Toarcian *†'' Pseudipsvicia'' Handlirsch 1939 Blackstone Formation, Australia, Norian * † '' Triassodoecus'' Evans, 1963
Hawkesbury Sandstone Sydney sandstone is the common name for Sydney Basin Hawkesbury Sandstone, one variety of which is historically known as Yellowblock, and also as "yellow gold" a sedimentary rock named after the Hawkesbury River north of Sydney, where this ...
, Australia,
Anisian In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from million years ago until million years ago. The Anisian Age succeeds the Olenekian Age (part of the Lower Triassic Ep ...
* † '' Triscytina'' Evans, 1956 Mount Crosby Insect Bed, Australia, Norian * † '' Woottonia'' Popov & Shcherbakov, 1991 Tologoi Formation, Kazakhstan, Norian * † '' Yurigocimex'' Martins-Neto & Galego, 2003 Los Rastros Formation, Argentina, Carnian *


References


External links

Prehistoric insect families Extinct Hemiptera {{Paleo-insect-stub