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The house sparrow (''Passer domesticus'') is a
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
of the
sparrow Sparrow may refer to: Birds * Old World sparrows, family Passeridae * New World sparrows, family Passerellidae * two species in the Passerine family Estrildidae: ** Java sparrow ** Timor sparrow * Hedge sparrow, also known as the dunnock or hed ...
family Passeridae, found in most parts of the world. It is a small bird that has a typical length of and a mass of . Females and young birds are coloured pale brown and grey, and males have brighter black, white, and brown markings. One of about 25 
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''
Passer ''Passer'' is a genus of sparrows, also known as the true sparrows. The genus contains 28 species and includes the house sparrow and the Eurasian tree sparrow, two of the most common birds in the world. They are small birds with thick bills for ...
'', the house sparrow is native to most of Europe, the
Mediterranean Basin In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and wa ...
, and a large part of Asia. Its intentional or accidental introductions to many regions, including parts of Australasia, Africa, and the Americas, make it the most widely distributed wild bird. The house sparrow is strongly associated with human habitation, and can live in urban or rural settings. Though found in widely varied habitats and climates, it typically avoids extensive
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
s,
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
s, and
desert A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...
s away from human development. It feeds mostly on the seeds of grains and weeds, but it is an opportunistic eater and commonly eats insects and many other foods. Its predators include
domestic cats The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...
,
hawk Hawks are birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. This subfa ...
s, and many other predatory birds and
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...
s. Because of its numbers, ubiquity, and association with human settlements, the house sparrow is culturally prominent. It is extensively, and usually unsuccessfully, persecuted as an agricultural pest. It has also often been kept as a pet, as well as being a food item and a symbol of lust, sexual potency, commonness, and vulgarity. Though it is widespread and abundant, its numbers have declined in some areas. The animal's
conservation status The conservation status of a group of organisms (for instance, a species) indicates whether the group still exists and how likely the group is to become extinct in the near future. Many factors are taken into account when assessing conservation ...
is listed as
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
.


Description


Measurements and shape

The house sparrow is typically about long, ranging from . The house sparrow is a compact bird with a full chest and a large, rounded head. Its bill is stout and conical with a culmen length of , strongly built as an adaptation for eating seeds. Its tail is short, at long. The wing chord is , and the tarsus is . Wingspan ranges from . In mass, the house sparrow ranges from . Females usually are slightly smaller than males. The median mass on the European continent for both sexes is about , and in more southerly subspecies is around . Younger birds are smaller, males are larger during the winter, and females are larger during the breeding season. Birds at higher latitudes, colder climates, and sometimes higher altitudes are larger (under
Bergmann's rule Bergmann's rule is an ecogeographical rule that states that within a broadly distributed taxonomic clade, populations and species of larger size are found in colder environments, while populations and species of smaller size are found in warmer r ...
), both between and within
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
.


Plumage

The
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
of the house sparrow is mostly different shades of grey and brown. The sexes exhibit strong dimorphism: the female is mostly buffish above and below, while the male has boldly coloured head markings, a reddish back, and grey underparts. The male has a dark grey
crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...
from the top of its bill to its back, and chestnut brown flanking its crown on the sides of its head. It has black around its bill, on its throat, and on the spaces between its bill and eyes ( lores). It has a small white stripe between the lores and crown and small white spots immediately behind the eyes (postoculars), with black patches below and above them. The underparts are pale grey or white, as are the cheeks, ear
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are ...
, and stripes at the base of the head. The upper back and mantle are a warm brown, with broad black streaks, while the lower back, rump and upper tail coverts are greyish brown. The male is duller in fresh nonbreeding plumage, with whitish tips on many feathers. Wear and preening expose many of the bright brown and black markings, including most of the black throat and chest patch, called the "bib" or "badge". The badge is variable in width and general size, and may signal social status or fitness. This hypothesis has led to a "veritable 'cottage industry'" of studies, which have only conclusively shown that patches increase in size with age. The male's bill is dark grey, but black in the breeding season. The female has no black markings or grey crown. Its upperparts and head are brown with darker streaks around the mantle and a distinct pale
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...
. Its underparts are pale grey-brown. The female's bill is brownish-grey and becomes darker in breeding plumage approaching the black of the male's bill. Juveniles are similar to the adult female, but deeper brown below and paler above, with paler and less defined supercilia. Juveniles have broader buff feather edges, and tend to have looser, scruffier plumage, like moulting adults. Juvenile males tend to have darker throats and white postoculars like adult males, while juvenile females tend to have white throats. However, juveniles cannot be reliably sexed by plumage: some juvenile males lack any markings of the adult male, and some juvenile females have male features. The bills of young birds are light yellow to
straw Straw is an agricultural byproduct consisting of the dry stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the yield of cereal crops such as barley, oats, rice, rye and wheat. It has a number ...
, paler than the female's bill. Immature males have paler versions of the adult male's markings, which can be very indistinct in fresh plumage. By their first breeding season, young birds generally are indistinguishable from other adults, though they may still be paler during their first year.


Voice

Most house sparrow vocalisations are variations on its short and frequent chirping call. Transcribed as ''chirrup'', ''tschilp'', or ''philip'', this note is made as a contact call by flocking or resting birds; or by males to proclaim nest ownership and invite pairing. In the breeding season, the male gives this call repetitively, with emphasis and speed, but not much rhythm, forming what is described either as a song or an "ecstatic call" similar to a song. Young birds also give a true song, especially in captivity, a warbling similar to that of the
European greenfinch The European greenfinch or simply the greenfinch (''Chloris chloris'') is a small passerine bird in the finch family Fringillidae. This bird is widespread throughout Europe, North Africa and Southwest Asia. It is mainly resident, but some n ...
. Aggressive males give a trilled version of their call, transcribed as "chur-''chur-r-r-it-it-it-it''". This call is also used by females in the breeding season, to establish dominance over males while displacing them to feed young or incubate eggs. House sparrows give a nasal alarm call, the basic sound of which is transcribed as ''quer'', and a shrill ''chree'' call in great distress. Another vocalisation is the "appeasement call", a soft ''quee'' given to inhibit aggression, usually given between birds of a mated pair. These vocalisations are not unique to the house sparrow, but are shared, with small variations, by all sparrows.


Variation

Some variation is seen in the 12 subspecies of house sparrows, which are divided into two groups, the
Oriental The Orient is a term for the East in relation to Europe, traditionally comprising anything belonging to the Eastern world. It is the antonym of '' Occident'', the Western World. In English, it is largely a metonym for, and coterminous with, the ...
''P. d. indicus'' group, and the
Palaearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
''P. d. domesticus'' group. Birds of the ''P. d. domesticus'' group have grey cheeks, while ''P. d. indicus'' group birds have white cheeks, as well as bright colouration on the crown, a smaller bill, and a longer black bib. The subspecies ''P. d. tingitanus'' differs little from the nominate subspecies, except in the worn breeding plumage of the male, in which the head is speckled with black and underparts are paler. ''P. d. balearoibericus'' is slightly paler than the nominate, but darker than ''P. d. bibilicus''. ''P. d. bibilicus'' is paler than most subspecies, but has the grey cheeks of ''P. d. domesticus'' group birds. The similar ''P. d. persicus'' is paler and smaller, and ''P. d. niloticus'' is nearly identical but smaller. Of the less widespread ''P. d. indicus'' group subspecies, ''P. d. hyrcanus'' is larger than ''P. d. indicus'', ''P. d. hufufae'' is paler, ''P. d. bactrianus'' is larger and paler, and ''P. d. parkini'' is larger and darker with more black on the breast than any other subspecies.


Identification

The house sparrow can be confused with a number of other seed-eating birds, especially its relatives in the genus ''Passer''. Many of these relatives are smaller, with an appearance that is neater or "cuter", as with the
Dead Sea sparrow The Dead Sea sparrow (''Passer moabiticus'') is a species of bird in the Old World sparrow family Passeridae, with one subspecies breeding in parts of the Middle East and another in western Afghanistan and eastern Iran. The eastern subspecies '' ...
. The dull-coloured female can often not be distinguished from other females, and is nearly identical to those of the
Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many Latin American countries **Spanish cuisine Other places * Spanish, Ontario, Can ...
and Italian sparrows. The
Eurasian tree sparrow The Eurasian tree sparrow (''Passer montanus'') is a passerine bird in the sparrow family with a rich chestnut crown and nape, and a black patch on each pure white cheek. The sexes are similarly plumaged, and young birds are a duller version ...
is smaller and slenderer with a chestnut crown and a black patch on each cheek. The male Spanish sparrow and Italian sparrow are distinguished by their chestnut crowns. The Sind sparrow is very similar but smaller, with less black on the male's throat and a distinct pale supercilium on the female.


Taxonomy and systematics


Names

The house sparrow was among the first animals to be given a
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
in the modern system of
biological classification In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are give ...
, since it was described by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
, in the 1758
10th edition 1 (one, unit, unity) is a number representing a single or the only entity. 1 is also a numerical digit and represents a single unit of counting or measurement. For example, a line segment of ''unit length'' is a line segment of length 1. ...
of ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial ...
''. It was described from a
type specimen In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes th ...
collected in
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic countries, Nordic c ...
, with the name ''Fringilla domestica''. Later, the genus name '' Fringilla'' came to be used only for the common chaffinch and its relatives, and the house sparrow has usually been placed in the genus ''Passer'' created by French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published wo ...
in 1760. The bird's scientific name and its usual English name have the same meaning. The
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
word ''passer'', like the English word "sparrow", is a term for small active birds, coming from a root word referring to speed. The Latin word ''domesticus'' means "belonging to the house", like the common name a reference to its association with humans. The house sparrow is also called by a number of alternative English names, including ''English sparrow'', chiefly in North America; and ''Indian sparrow'' or ''Indian house sparrow'', for the birds of the
Indian subcontinent The Indian subcontinent is a physiographical region in Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian Ocean from the Himalayas. Geopolitically, it includes the countries of Bangladesh, Bhutan, In ...
and
Central Asia Central Asia, also known as Middle Asia, is a region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes the fo ...
. Dialectal names include ''sparr'', ''sparrer'', ''spadger'', ''spadgick'', and ''philip'', mainly in southern England; ''spug'' and ''spuggy'', mainly in northern England; ''spur'' and ''sprig'', mainly in
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to ...
; and ''spatzie'' or ''spotsie'', from the German ''Spatz'', in North America.


Taxonomy

The genus ''Passer'' contains about 25 species, depending on the authority, 26 according to the ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...
''. Most ''Passer'' species are dull-coloured birds with short, square tails and stubby, conical beaks, between long.
Mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
studies suggest that
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution withi ...
in the genus occurred during the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
and earlier, while other evidence suggests speciation occurred 25,000 to 15,000 years ago. Within ''Passer'', the house sparrow is part of the "
Palaearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
black-bibbed sparrows" group and a close relative of the Mediterranean "willow sparrows". The taxonomy of the house sparrow and its Mediterranean relatives is complicated. The common type of "willow sparrow" is the Spanish sparrow, which resembles the house sparrow in many respects. It frequently prefers wetter habitats than the house sparrow, and it is often colonial and nomadic. In most of the Mediterranean, one or both species occur, with some degree of hybridisation. In North Africa, the two species hybridise extensively, forming highly variable mixed populations with a full range of characters from pure house sparrows to pure Spanish sparrows. In most of Italy, the breeding species is the Italian sparrow, which has an appearance intermediate between those of the house and Spanish sparrows. Its specific status and origin are the subject of much debate, but it may be a case of long-ago
hybrid speciation Hybrid speciation is a form of speciation where hybridization between two different species leads to a new species, reproductively isolated from the parent species. Previously, reproductive isolation between two species and their parents was thou ...
. In the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
, the Italian sparrow intergrades over a narrow roughly strip with the house sparrow, and some house sparrows migrate into the Italian sparrow's range in winter. On the Mediterranean islands of
Malta Malta ( , , ), officially the Republic of Malta ( mt, Repubblika ta' Malta ), is an island country in the Mediterranean Sea. It consists of an archipelago, between Italy and Libya, and is often considered a part of Southern Europe. It lies ...
,
Gozo Gozo (, ), Maltese: ''Għawdex'' () and in antiquity known as Gaulos ( xpu, 𐤂𐤅𐤋, ; grc, Γαῦλος, Gaúlos), is an island in the Maltese archipelago in the Mediterranean Sea. The island is part of the Republic of Malta. After ...
,
Crete Crete ( el, Κρήτη, translit=, Modern: , Ancient: ) is the largest and most populous of the Greek islands, the 88th largest island in the world and the fifth largest island in the Mediterranean Sea, after Sicily, Sardinia, Cyprus, ...
,
Rhodes Rhodes (; el, Ρόδος , translit=Ródos ) is the largest and the historical capital of the Dodecanese islands of Greece. Administratively, the island forms a separate municipality within the Rhodes regional unit, which is part of the S ...
, and
Karpathos Karpathos ( el, Κάρπαθος, ), also Carpathos, is the second largest of the Greek Dodecanese islands, in the southeastern Aegean Sea. Together with the neighboring smaller Saria Island it forms the municipality of Karpathos, which is part of ...
, other apparently intermediate birds are of unknown status.


Subspecies

A large number of subspecies have been named, of which 12 were recognised in the ''Handbook of the Birds of the World''. These subspecies are divided into two groups, the Palaearctic ''P. d. domesticus'' group, and the Oriental ''P. d. indicus'' group. Several Middle Eastern subspecies, including ''P. d. biblicus'', are sometimes considered a third, intermediate group. The subspecies ''P. d. indicus'' was described as a species, and was considered to be distinct by many ornithologists during the 19th century. Migratory birds of the subspecies ''P. d. bactrianus'' in the ''P. d. indicus'' group were recorded overlapping with ''P. d. domesticus'' birds without hybridising in the 1970s, so the Soviet scientists Edward I. Gavrilov and M. N. Korelov proposed the separation of the ''P. d. indicus'' group as a separate species. However, ''P. d. indicus'' group and ''P. d. domesticus'' group birds intergrade in a large part of
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
, so this split is rarely recognised. In North America, house sparrow populations are more differentiated than those in Europe. This variation follows predictable patterns, with birds at higher latitudes being larger and darker and those in arid areas being smaller and paler. However, how much this is caused by evolution or by environment is not clear. Similar observations have been made in New Zealand and in South Africa. The introduced house sparrow populations may be distinct enough to merit subspecies status, especially in North America and southern Africa, and American ornithologist
Harry Church Oberholser Harry Church Oberholser (June 25, 1870 – December 25, 1963) was an American ornithologist. Biography Harry Oberholser was born to Jacob and Lavera S. Oberholser on June 25, 1870, in Brooklyn, New York. He attended Columbia University, but did ...
even gave the subspecies name ''P. d. plecticus'' to the paler birds of western North America. ;''P. d. domesticus'' group * ''P. d. domesticus'' Linnaeus, 1758, the
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all speci ...
, is found in most of Europe, across northern Asia to
Sakhalin Sakhalin ( rus, Сахали́н, r=Sakhalín, p=səxɐˈlʲin; ja, 樺太 ''Karafuto''; zh, c=, p=Kùyèdǎo, s=库页岛, t=庫頁島; Manchu: ᠰᠠᡥᠠᠯᡳᠶᠠᠨ, ''Sahaliyan''; Orok: Бугата на̄, ''Bugata nā''; Nivkh ...
and
Kamchatka The Kamchatka Peninsula (russian: полуостров Камчатка, Poluostrov Kamchatka, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and west ...
. It is the most widely introduced subspecies. * ''P. d. balearoibericus'' von Jordans, 1923, described from
Majorca Mallorca, or Majorca, is the largest island in the Balearic Islands, which are part of Spain and located in the Mediterranean. The capital of the island, Palma, is also the capital of the autonomous community of the Balearic Islands. The Bale ...
, is found in the
Balearic Islands The Balearic Islands ( es, Islas Baleares ; or ca, Illes Balears ) are an archipelago in the Balearic Sea, near the eastern coast of the Iberian Peninsula. The archipelago is an autonomous community and a province of Spain; its capital is ...
, southern France, the
Balkans The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
, and
Anatolia Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...
. * ''P. d. tingitanus'' ( Loche, 1867), described from
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
, is found in the
Maghreb The Maghreb (; ar, الْمَغْرِب, al-Maghrib, lit=the west), also known as the Arab Maghreb ( ar, المغرب العربي) and Northwest Africa, is the western part of North Africa and the Arab world. The region includes Algeria, ...
from
Ajdabiya Ajdabiya ( ; ar, أجدابيا, Aǧdābiyā) is a town in and capital of the Al Wahat District in northeastern Libya. It is some south of Benghazi. From 2001 to 2007 it was part of and capital of the Ajdabiya District. The town is divided in ...
in
Libya Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to the east, Suda ...
to
Béni Abbès Béni Abbès ( ar, بني عباس), also known as the ''Pearl of the Saoura'', and also as the ''White Oasis'', is a town and Communes of Algeria, commune located in western Algeria in Béchar Province, far from the provincial capital Béchar, ...
in Algeria, and to
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to A ...
's Atlantic coast. It hybridises extensively with the Spanish sparrow, especially in the eastern part of its range. * ''P. d. niloticus'' Nicoll and Bonhote, 1909, described from
Faiyum Faiyum ( ar, الفيوم ' , borrowed from cop,  ̀Ⲫⲓⲟⲙ or Ⲫⲓⲱⲙ ' from egy, pꜣ ym "the Sea, Lake") is a city in Middle Egypt. Located southwest of Cairo, in the Faiyum Oasis, it is the capital of the modern Faiyum ...
, Egypt, is found along the
Nile The Nile, , Bohairic , lg, Kiira , Nobiin: Áman Dawū is a major north-flowing river in northeastern Africa. It flows into the Mediterranean Sea. The Nile is the longest river in Africa and has historically been considered the longest riv ...
north of
Wadi Halfa Wādī Ḥalfā ( ar, وادي حلفا) is a city in the Northern state of Sudan on the shores of Lake Nubia near the border with Egypt. It is the terminus of a rail line from Khartoum and the point where goods are transferred from rail to fer ...
, Sudan. It intergrades with ''bibilicus'' in the Sinai, and with ''rufidorsalis'' in a narrow zone around
Wadi Halfa Wādī Ḥalfā ( ar, وادي حلفا) is a city in the Northern state of Sudan on the shores of Lake Nubia near the border with Egypt. It is the terminus of a rail line from Khartoum and the point where goods are transferred from rail to fer ...
. It has been recorded in
Somaliland Somaliland,; ar, صوماليلاند ', ' officially the Republic of Somaliland,, ar, جمهورية صوماليلاند, link=no ''Jumhūrīyat Ṣūmālīlānd'' is a ''de facto'' sovereign state in the Horn of Africa, still conside ...
. * ''P. d. persicus'' Zarudny and Kudashev, 1916, described from the
Karun River The Karun ( fa, کارون, ) is the Iranian river with the highest water flow, and its only navigable river. It is long. It rises in the Zard Kuh mountains of the Bakhtiari district in the Zagros Range, receiving many tributaries, such as t ...
in
Khuzestan Khuzestan Province (also spelled Xuzestan; fa, استان خوزستان ''Ostān-e Xūzestān'') is one of the 31 provinces of Iran. It is in the southwest of the country, bordering Iraq and the Persian Gulf. Its capital is Ahvaz and it cover ...
, Iran, is found in the western and central Iran south of the
Alborz The Alborz ( fa, البرز) range, also spelled as Alburz, Elburz or Elborz, is a mountain range in northern Iran that stretches from the border of Azerbaijan along the western and entire southern coast of the Caspian Sea and finally runs nort ...
mountains, intergrading with ''indicus'' in eastern Iran, and Afghanistan. * ''P. d. biblicus'' Hartert, 1910, described from Palestine, is found in the Middle East from
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is ...
and southeastern Turkey to the Sinai in the west and from
Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transcontinental country located at the boundary of Eastern Europe and Western Asia. It is a part of th ...
to
Kuwait Kuwait (; ar, الكويت ', or ), officially the State of Kuwait ( ar, دولة الكويت '), is a country in Western Asia. It is situated in the northern edge of Eastern Arabia at the tip of the Persian Gulf, bordering Iraq to Iraq–Ku ...
in the east. ;''P. d. indicus'' group * ''P. d. hyrcanus'' Zarudny and Kudashev, 1916, described from
Gorgan Gorgan ( fa, گرگان ; also romanized as ''Gorgān'', ''Gurgān'', and ''Gurgan''), formerly Esterabad ( ; also romanized as ''Astarābād'', ''Asterabad'', and ''Esterābād''), is the capital city of Golestan Province, Iran. It lies appro ...
, Iran, is found along the southern coast of the
Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad steppe of Central A ...
from Gorgan to southeastern Azerbaijan. It intergrades with ''P. d. persicus'' in the Alborz mountains, and with ''P. d. bibilicus'' to the west. It is the subspecies with the smallest range. * ''P. d. bactrianus'' Zarudny and Kudashev, 1916, described from
Tashkent Tashkent (, uz, Toshkent, Тошкент/, ) (from russian: Ташкент), or Toshkent (; ), also historically known as Chach is the capital and largest city of Uzbekistan. It is the most populous city in Central Asia, with a population of 2 ...
, is found in southern Kazakhstan to the
Tian Shan The Tian Shan,, , otk, 𐰴𐰣 𐱅𐰭𐰼𐰃, , tr, Tanrı Dağı, mn, Тэнгэр уул, , ug, تەڭرىتاغ, , , kk, Тәңіртауы / Алатау, , , ky, Теңир-Тоо / Ала-Тоо, , , uz, Tyan-Shan / Tangritog‘ ...
and northern Iran and
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is borde ...
. It intergrades with ''persicus'' in
Baluchistan Balochistan ( ; bal, بلۏچستان; also romanised as Baluchistan and Baluchestan) is a historical region in Western Asia, Western and South Asia, located in the Iranian plateau's far southeast and bordering the Indian Plate and the Arabian S ...
and with ''indicus'' across central Afghanistan. Unlike most other house sparrow subspecies, it is almost entirely migratory, wintering in the plains of the northern Indian subcontinent. It is found in open country rather than in settlements, which are occupied by the Eurasian tree sparrow in its range. There is an exceptional record from Sudan. * ''P. d. parkini'' Whistler, 1920, described from
Srinagar Srinagar (English: , ) is the largest city and the summer capital of Jammu and Kashmir, India. It lies in the Kashmir Valley on the banks of the Jhelum River, a tributary of the Indus, and Dal and Anchar lakes. The city is known for its ...
,
Kashmir Kashmir () is the northernmost geographical region of the Indian subcontinent. Until the mid-19th century, the term "Kashmir" denoted only the Kashmir Valley between the Great Himalayas and the Pir Panjal Range. Today, the term encompas ...
, is found in the western
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...
from the
Pamir Mountains The Pamir Mountains are a mountain range between Central Asia and Pakistan. It is located at a junction with other notable mountains, namely the Tian Shan, Karakoram, Kunlun, Hindu Kush and the Himalaya mountain ranges. They are among the wor ...
to southeastern
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...
. It is migratory, like ''P. d. bactrianus''. * ''P. d. indicus''
Jardine Jardine is a surname. Notable people with the surname include: * Al Jardine (born 1942), member of the Beach Boys * Alexander Jardine (Medal of Honor) (1874–1949), American Medal of Honor recipient * Antonio Jardine (born 1988), NCAA college b ...
and
Selby Selby is a market town and civil parish in the Selby District of North Yorkshire, England, south of York on the River Ouse, with a population at the 2011 census of 14,731. The town was historically part of the West Riding of Yorkshire until ...
, 1831
, described from
Bangalore Bangalore (), officially Bengaluru (), is the capital and largest city of the Indian state of Karnataka. It has a population of more than and a metropolitan population of around , making it the third most populous city and fifth most ...
, is found in the Indian subcontinent south of the Himalayas, in
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
, western
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...
, eastern Iran, southwestern Arabia and southern
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
. * ''P. d. hufufae''
Ticehurst Ticehurst is both a village and a large civil parish in the Rother district of East Sussex, England. The parish lies in the upper reaches of both the Bewl stream before it enters Bewl Water and in the upper reaches of the River Rother flowi ...
and Cheeseman, 1924
, described from
Hofuf Al-Hofuf ( ar, ٱلْهُفُوف ', also spelled Hofuf or Hufuf, also known as "Al-Hasa", "Al-Ahsa" or "Al-Hassa") is the major urban city in the Al-Ahsa Oasis in the Eastern Province of Saudi Arabia, with a population of 858,395 (as of 202 ...
in
Saudi Arabia Saudi Arabia, officially the Kingdom of Saudi Arabia (KSA), is a country in Western Asia. It covers the bulk of the Arabian Peninsula, and has a land area of about , making it the fifth-largest country in Asia, the second-largest in the Ara ...
, is found in northeastern Arabia. * ''P. d. rufidorsalis'' C. L. Brehm, 1855, described from
Khartoum Khartoum or Khartum ( ; ar, الخرطوم, Al-Khurṭūm, din, Kaartuɔ̈m) is the capital of Sudan. With a population of 5,274,321, its metropolitan area is the largest in Sudan. It is located at the confluence of the White Nile, flowing n ...
, Sudan, is found in the Nile valley from Wadi Halfa south to Renk in northern
South Sudan South Sudan (; din, Paguot Thudän), officially the Republic of South Sudan ( din, Paankɔc Cuëny Thudän), is a landlocked country in East Africa. It is bordered by Ethiopia, Sudan, Central African Republic, Democratic Republic of th ...
, and in eastern Sudan, northern Ethiopia to the
Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; ...
coast in
Eritrea Eritrea ( ; ti, ኤርትራ, Ertra, ; ar, إرتريا, ʾIritriyā), officially the State of Eritrea, is a country in the Horn of Africa region of Eastern Africa, with its capital and largest city at Asmara. It is bordered by Ethiopi ...
. It has also been introduced to
Mohéli Mohéli , also known as Mwali, is an autonomous island that forms part of the Union of the Comoros. It is the smallest of the three major islands in the country. It is located in the Indian Ocean off the coast of Africa and it is the smallest o ...
in the
Comoros The Comoros,, ' officially the Union of the Comoros,; ar, الاتحاد القمري ' is an independent country made up of three islands in southeastern Africa, located at the northern end of the Mozambique Channel in the Indian Ocean. It ...
.


Distribution and habitat

The house sparrow originated in the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabian Peninsula, Arabia (including the Arabian Peninsula and Bahrain), Anatolia, Asia Minor (Asian part of Turkey except Hatay Pro ...
and spread, along with agriculture, to most of Eurasia and parts of North Africa. Since the mid-19th century, it has reached most of the world, chiefly due to deliberate introductions, but also through natural and shipborne dispersal. Its introduced range encompasses most of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
(including
Bermuda ) , anthem = "God Save the King" , song_type = National song , song = "Hail to Bermuda" , image_map = , map_caption = , image_map2 = , mapsize2 = , map_caption2 = , subdivision_type = Sovereign state , subdivision_name = , es ...
),
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, southern South America,
southern Africa Southern Africa is the southernmost subregion of the African continent, south of the Congo and Tanzania. The physical location is the large part of Africa to the south of the extensive Congo River basin. Southern Africa is home to a number o ...
, part of
West Africa West Africa or Western Africa is the westernmost region of Africa. The United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Mali ...
,
Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...
,
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island coun ...
, and islands throughout the world. It has greatly extended its range in northern Eurasia since the 1850s, and continues to do so, as was shown by its colonisation around 1990 of
Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its ...
and
Rishiri Island is a high island in the Sea of Japan off the coast of Hokkaido, Japan. Administratively the island is part of Hokkaido Prefecture, and is divided between two towns, Rishiri and Rishirifuji. The island is formed by the cone-shaped extinct ...
, Japan. The extent of its range makes it the most widely distributed wild bird on the planet.


Introduction

The house sparrow has become highly successful in most parts of the world where it has been introduced. This is mostly due to its early adaptation to living with humans, and its adaptability to a wide range of conditions. Other factors may include its robust immune response, compared to the
Eurasian tree sparrow The Eurasian tree sparrow (''Passer montanus'') is a passerine bird in the sparrow family with a rich chestnut crown and nape, and a black patch on each pure white cheek. The sexes are similarly plumaged, and young birds are a duller version ...
. Where introduced, it can extend its range quickly, sometimes at a rate over per year. In many parts of the world, it has been characterised as a pest, and poses a threat to native birds. A few introductions have died out or been of limited success, such as those to
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland ...
and
Cape Verde , national_anthem = () , official_languages = Portuguese , national_languages = Cape Verdean Creole , capital = Praia , coordinates = , largest_city = capital , demonym ...
. The first of many successful introductions to North America occurred when birds from England were released in New York City, in 1852, intended to control the ravages of the linden moth. In North America, the house sparrow now occurs from the
Northwest Territories The Northwest Territories (abbreviated ''NT'' or ''NWT''; french: Territoires du Nord-Ouest, formerly ''North-Western Territory'' and ''North-West Territories'' and namely shortened as ''Northwest Territory'') is a federal territory of Canada. ...
of Canada to southern Panama, and it is one of the most abundant birds of the continent. The house sparrow was first introduced to Australia in 1863 at Melbourne and is common throughout the eastern part of the continent as far north as Cape York Peninsula, Cape York, but has been prevented from establishing itself in Western Australia, where every house sparrow found in the state is killed. House sparrows were introduced in New Zealand in 1859, and from there reached many of the Pacific islands, including Hawaii. In southern Africa, birds of both the European subspecies (''P. d. domesticus'') and the Indian subspecies (''P. d. indicus'') were introduced around 1900. Birds of ''P. d. domesticus'' ancestry are confined to a few towns, while ''P. d. indicus'' birds have spread rapidly, reaching Tanzania in the 1980s. Despite this rapid spread, native relatives such as the Cape sparrow also occur and thrive in urban habitats. In South America, it was first introduced near Buenos Aires around 1870, and quickly became common in most of the southern part of the continent. It now occurs almost continuously from Tierra del Fuego to the fringes of the Amazon basin, with isolated populations as far north as coastal Venezuela.


Habitat

The house sparrow is closely associated with human habitation and cultivation. It is not an obligate Commensalism, commensal of humans as some have suggested: birds of the migratory Central Asian subspecies usually breed away from humans in open country, and birds elsewhere are occasionally found away from humans. The only terrestrial habitats that the house sparrow does not inhabit are dense forest and tundra. Well adapted to living around humans, it frequently lives and even breeds indoors, especially in factories, warehouses, and zoos. It has been recorded breeding in an English coal mine below ground, and feeding on the Empire State Building's observation deck at night. It reaches its greatest densities in urban centres, but its reproductive success is greater in suburbs, where insects are more abundant. On a larger scale, it is most abundant in wheat-growing areas such as the Midwestern United States. It tolerates a variety of climates, but prefers drier conditions, especially in moist tropical climates. It has several adaptations to dry areas, including a high salt tolerance and an ability to survive without water by ingesting berry, berries. In most of eastern Asia, the house sparrow is entirely absent, replaced by the Eurasian tree sparrow. Where these two species overlap, the house sparrow is usually more common than the Eurasian tree sparrow, but one species may replace the other in a manner that ornithologist Maud Doria Haviland described as "random, or even capricious". In most of its range, the house sparrow is extremely common, despite some declines, but in marginal habitats such as rainforest or mountain ranges, its distribution can be spotty.


Behaviour


Social behaviour

The house sparrow is a very social bird. It is gregarious during all seasons when feeding, often forming flocks with other species of birds. It Roosting, roosts communally and while breeding nests are usually grouped together in clumps. House sparrows also engage in social activities such as dust bath, dust or water bathing and "social singing", in which birds call together in bushes. The house sparrow feeds mostly on the ground, but it flocks in trees and bushes. At feeding stations and nests, female house sparrows are dominant despite their smaller size, and they can fight over males in the breeding season.


Sleep and roosting

House sparrows sleep with the bill tucked underneath the scapular feathers. Outside of the reproductive season, they often roost communally in trees or shrubs. Much communal chirping occurs before and after the birds settle in the roost in the evening, as well as before the birds leave the roost in the morning. Some congregating sites separate from the roost may be visited by the birds prior to settling in for the night.


Body maintenance

Dust or water bathing is common and often occurs in groups. Anting (bird activity), Anting is rare. Head scratching is done with the leg over the drooped wing.


Feeding

As an adult, the house sparrow mostly feeds on the seeds of cereal, grains and weeds, but it is opportunistic and adaptable, and eats whatever foods are available. In towns and cities, it often scavenges for food in garbage containers and congregates in the outdoors of restaurants and other eating establishments to feed on leftover food and crumbs. It can perform complex tasks to obtain food, such as opening automatic doors to enter supermarkets, clinging to hotel walls to watch vacationers on their balconies, and nectar robbing kowhai flowers. In common with many other birds, the house sparrow requires gastrolith, grit to digest the harder items in its diet. Grit can be either stone, often grains of masonry, or the shells of eggs or snails; oblong and rough grains are preferred. Several studies of the house sparrow in temperate agricultural areas have found the proportion of seeds in its diet to be about 90%. It will eat almost any seeds, but where it has a choice, it prefers cereal, corn: oats, wheat or maize. Rural birds tend to eat more waste seed from animal dung and seed from fields while urban birds tend to eat more commercial bird seed and weed seed. In urban areas, the house sparrow also feeds largely on food provided directly or indirectly by humans, such as bread, though it prefers raw seeds. The house sparrow also eats some plant matter besides seeds, including buds, berries, and fruits such as grapes and cherries. In temperate areas, the house sparrow has an unusual habit of tearing flowers, especially yellow ones, in the spring. Animals form another important part of the house sparrow's diet, chiefly insects, of which beetles, caterpillars, dipteran flies, and aphids are especially important. Various noninsect arthropods are eaten, as are molluscs and crustaceans where available, earthworms, and even vertebrates such as lizards and frogs. Young house sparrows are fed mostly on insects until about 15 days after hatching. They are also given small quantities of seeds, spiders, and grit. In most places, grasshoppers and cricket (insect), crickets are the most abundant foods of nestlings. Hemiptera, True bugs, ants, sawfly, sawflies, and beetles are also important, but house sparrows take advantage of whatever foods are abundant to feed their young. House sparrows have been observed Kleptoparasitism, stealing prey from other birds, including American robins. The gut microbiota of house sparrows differs between chicks and adults, with Pseudomonadota (formerly Proteobacteria) decreasing in chicks when they get to around 9 days old, whilst the relative abundance of Bacillota increase.


Locomotion

The house sparrow's flight is direct (not undulating) and flapping, averaging and about 15 wingbeats per second. On the ground, the house sparrow typically hops rather than walks. It can swim when pressed to do so by pursuit from predators. Captive birds have been recorded diving and swimming short distances under water.


Dispersal and migration

Most house sparrows do not move more than a few kilometres during their lifetimes. However, limited migration occurs in all regions. Some young birds disperse long distances, especially on coasts, and mountain birds move to lower elevations in winter. Two subspecies, ''P. d. bactrianus'' and ''P. d. parkini'', are predominantly bird migration, migratory. Unlike the birds in sedentary populations that migrate, birds of migratory subspecies prepare for migration by putting on weight.


Breeding

House sparrows can breed in the breeding season immediately following their hatching, and sometimes attempt to do so. Some birds breeding for the first time in tropical areas are only a few months old and still have juvenile plumage. Birds breeding for the first time are rarely successful in raising young, and reproductive success increases with age, as older birds breed earlier in the breeding season, and fledge more young. As the breeding season approaches, hormone releases trigger enormous increases in the size of the sexual organs and changes in day length lead males to start calling by nesting sites. The timing of mating and egg-laying varies geographically, and between specific locations and years because a sufficient supply of insects is needed for egg formation and feeding nestlings. Males take up nesting sites before the breeding season, by frequently calling beside them. Unmated males start nest construction and call particularly frequently to attract females. When a female approaches a male during this period, the male courtship display, displays by moving up and down while drooping and shivering his wings, pushing up his head, raising and spreading his tail, and showing his bib. Males may try to mate with females while calling or displaying. In response, a female will adopt a threatening posture and attack a male before flying away, pursued by the male. The male displays in front of her, attracting other males, which also pursue and display to the female. This group display usually does not immediately result in copulations. Other males usually do not copulate with the female. Copulation is typically initiated by the female giving a soft ''dee-dee-dee'' call to the male. Birds of a pair copulation (zoology), copulate frequently until the female is laying eggs, and the male mounts the female repeatedly each time a pair mates. The house sparrow is monogamous, and typically mates for life, but birds from pairs often engage in extra-pair copulations, so about 15% of house sparrow fledglings are unrelated to their mother's mate. Males guard their mates carefully to avoid being cuckolded, and most extra-pair copulation occurs away from nest sites. Males may sometimes have multiple mates, and bigamy is mostly limited by aggression between females. Many birds do not find a nest and a mate, and instead may serve as helpers at the nest, helpers around the nest for mated pairs, a role which increases the chances of being chosen to replace a lost mate. Lost mates of both sexes can be replaced quickly during the breeding season. The formation of a pair and the bond between the two birds is tied to the holding of a nest site, though paired house sparrows can recognise each other away from the nest.


Nesting

Nest sites are varied, though cavities are preferred. Nests are most frequently built in the eaves and other crevices of houses. Holes in cliffs and banks, and tree hollows, are also used. A sparrow sometimes excavates its own nests in sandy banks or rotten branches, but more frequently uses the nests of other birds such as those of swallows in banks and cliffs, and old tree cavity nests. It usually uses deserted nests, though sometimes it usurps active ones by driving away or killing the occupants. Tree hollows are more commonly used in North America than in Europe, putting the sparrows in competition with bluebirds and other North American cavity nesters, and thereby contributing to their population declines. Especially in warmer areas, the house sparrow may build its nests in the open, on the branches of trees, especially evergreens and hawthorns, or in the nests of large birds such as storks or magpies. In open nesting sites, breeding success tends to be lower, since breeding begins late and the nest can easily be destroyed or damaged by storms. Less common nesting sites include street lights and neon signs, favoured for their warmth; and the old open-topped nests of other songbirds, which are then domed over. Usually the couples repeat copulation many times. Every copulation is followed by some break of 3 to 4 seconds, and in that time both pair change their position by some distance. The nest is usually domed, though it may lack a roof in enclosed sites. It has an outer layer of stems and roots, a middle layer of dead grass and leaves, and a lining of feathers, as well as of paper and other soft materials. Nests typically have external dimensions of 20 × 30 cm (8 × 12 in), but their size varies greatly. The building of the nest is initiated by the unmated male while displaying to females. The female assists in building, but is less active than the male. Some nest building occurs throughout the year, especially after moult in autumn. In colder areas house sparrows build specially created roost nests, or roost in street lights, to avoid losing heat during the winter. House sparrows do not hold territories, but they defend their nests aggressively against intruders of the same sex. House sparrows' nests support a wide range of scavenging insects, including nest flies such as ''Neottiophilum praestum'', ''Protocalliphora'' blowflies, and over 1,400 species of beetle.


Eggs and young

Clutch (eggs), Clutches usually comprise four or five egg (biology), eggs, though numbers from one to 10 have been recorded. At least two clutches are usually laid, and up to seven a year may be laid in the tropics or four a year in temperate latitudes. When fewer clutches are laid in a year, especially at higher latitudes, the number of eggs per clutch is greater. Central Asian house sparrows, which migrate and have only one clutch a year, average 6.5 eggs in a clutch. Clutch size is also affected by environmental and seasonal conditions, female age, and breeding density. Some intraspecific brood parasite, brood parasitism occurs, and instances of unusually large numbers of eggs in a nest may be the result of females laying eggs in the nests of their neighbours. Such foreign eggs are sometimes recognised and ejected by females. The house sparrow is a victim of interspecific brood parasites, but only rarely, since it usually uses nests in holes too small for parasites to enter, and it feeds its young foods unsuitable for young parasites. In turn, the house sparrow has once been recorded as a brood parasite of the American cliff swallow. The eggs are white, bluish white, or greenish white, spotted with brown or grey. Subelliptical in shape, they range from in length and in width, have an average mass of , and an average surface area of . Eggs from the tropical subspecies are distinctly smaller. Eggs begin to develop with the deposition of Egg yolk, yolk in the ovary a few days before ovulation. In the day between ovulation and laying, egg white forms, followed by eggshell. Eggs laid later in a clutch are larger, as are those laid by larger females, and egg size is hereditary. Eggs decrease slightly in size from laying to hatching. The yolk comprises 25% of the egg, the egg white 68%, and the shell 7%. Eggs are watery, being 79% liquid, and otherwise mostly protein. The female develops a brood patch of bare skin and plays the main part in avian incubation, incubating the eggs. The male helps, but can only cover the eggs rather than truly incubate them. The female spends the night incubating during this period, while the male roosts near the nest. Eggs hatch at the same time, after a short incubation period lasting 11–14 days, and exceptionally for as many as 17 or as few as 9. The length of the incubation period decreases as ambient temperature increases later in the breeding season. Young house sparrows remain in the nest for 11 to 23 days, normally 14 to 16 days. During this time, they are fed by both parents. As newly hatched house sparrows do not have sufficient insulation, they are brooded for a few days, or longer in cold conditions. The parents swallow the droppings produced by the hatchlings during the first few days; later, the droppings are moved up to away from the nest. The chicks' eyes open after about 4 days and, at an age of about 8 days, the young birds get their first down feather, down. If both parents perish, the ensuing intensive begging sounds of the young often attract replacement parents which feed them until they can sustain themselves. All the young in the nest leave it during the same period of a few hours. At this stage, they are normally able to fly. They start feeding themselves partly after 1 or 2 days, and sustain themselves completely after 7 to 10 days, 14 at the latest.


Survival

In adult house sparrows, annual survival is 45–65%. After fledging and leaving the care of their parents, young sparrows have a high mortality rate, which lessens as they grow older and more experienced. Only about 20–25% of birds hatched survive to their first breeding season. The oldest known wild house sparrow lived for nearly two decades; it was found dead 19 years and 9 months after it was bird ringing, ringed in Denmark. The oldest recorded captive house sparrow lived for 23 years. The typical ratio of males to females in a population is uncertain due to problems in collecting data, but a very slight preponderance of males at all ages is usual.


Predation

The house sparrow's main predators are cats and Bird of prey, birds of prey, but many other animals prey on them, including Corvidae, corvids, squirrels, and even humans—the house sparrow has been consumed in the past by people in many parts of the world, and it still is in parts of the Mediterranean. Most species of birds of prey have been recorded preying on the house sparrow in places where records are extensive. Accipiters and the Merlin (bird), merlin in particular are major predators, though cats are likely to have a greater impact on house sparrow populations. The house sparrow is also a common victim of roadkill; on European roads, it is the bird most frequently found dead.


Parasites and disease

The house sparrow is host to a huge number of parasites and diseases, and the effect of most is unknown. Ornithologist Ted R. Anderson listed thousands, noting that his list was incomplete. The commonly recorded bacterial pathogens of the house sparrow are often those common in humans, and include ''Salmonella'' and ''Escherichia coli''. ''Salmonella'' is common in the house sparrow, and a comprehensive study of house sparrow disease found it in 13% of sparrows tested. ''Salmonella'' epidemics in the spring and winter can kill large numbers of sparrows. The house sparrow hosts avian pox and avian malaria, which it has spread to the native forest birds of Hawaii. Many of the diseases hosted by the house sparrow are also present in humans and domestic animals, for which the house sparrow acts as a natural reservoir, reservoir host. Arbovirus infection, Arboviruses such as the West Nile virus, which most commonly infect insects and mammals, survive winters in temperate areas by going dormant in birds such as the house sparrow. A few records indicate disease extirpating house sparrow populations, especially from Scottish islands, but this seems to be rare. House sparrows are also infected by Haemosporida, haemosporidian parasites, but less so in urban than in rural areas ''Toxoplasma gondii'' has been detected in sparrows in northwestern China where they pose a risk due to their meat being consumed in the region. The house sparrow is infested by a number of external parasites, which usually cause little harm to adult sparrows. In Europe, the most common mite found on sparrows is ''Proctophyllodes'', the most common ticks are ''Argas reflexus'' and ''Ixodes arboricola'', and the most common flea on the house sparrow is ''Ceratophyllus gallinae''. ''Dermanyssus'' blood-feeding mites are also common ectoparasites of house sparrows, and these mites can enter human habitation and bite humans, causing a condition known as gamasoidosis. A number of chewing louse, chewing lice occupy different Ecological niche, niches on the house sparrow's body. ''Menacanthus'' lice occur across the house sparrow's body, where they feed on blood and feathers, while ''Brueelia'' lice feed on feathers and ''Philopterus fringillae'' occurs on the head.


Physiology

House sparrows express strong circadian rhythms of activity in the laboratory. They were among the first bird species to be seriously studied in terms of their circadian activity and photoperiodism, in part because of their availability and adaptability in captivity, but also because they can "find their way" and remain rhythmic in constant darkness. Such studies have found that the pineal gland is a central part of the house sparrow's circadian system: removal of the pineal eliminates the circadian rhythm of activity, and transplant of the pineal into another individual confers to this individual the rhythm phase of the donor bird. The suprachiasmatic nuclei of the hypothalamus have also been shown to be an important component of the circadian system of house sparrows. The photoreceptors involved in the synchronisation of the circadian clock to the external light-dark cycle are located in the brain and can be stimulated by light reaching them directly though the skull, as revealed by experiments in which blind sparrows, which normally can still synchronise to the light-dark cycle, failed to do so once India ink was injected as a screen under the skin on top of their skulls. Similarly, even when blind, house sparrows continue to be photoperiodic, i.e. show reproductive development when the days are long, but not when the days are short. This response is stronger when the feathers on top of the head are plucked, and is eliminated when India ink is injected under the skin at the top of the head, showing that the photoreceptors involved in the photoperiodic response to day length are located inside the brain. House sparrows have also been used in studies of nonphotic entrainment (i.e. synchronisation to an external cycle other than light and dark): for example, in constant darkness, a situation in which the birds would normally reveal their endogenous, non-24-hour, "free-running" rhythms of activity, they instead show 24-hour periodicity if they are exposed to two hours of chirp playbacks every 24 hours, matching their daily activity onsets with the daily playback onsets. House sparrows in constant dim light can also be entrained to a daily cycle based on the presence of food. Finally, house sparrows in constant darkness could be entrained to a cycle of high and low temperature, but only if the difference in temperature was large (38 °C versus 6 °C); some of the tested sparrows matched their activity to the warm phase, and others to the cold phase.


Relationships with humans

The house sparrow is closely associated with humans. They are believed to have become associated with humans around 10,000 years ago. The Turkestan subspecies (''P. d. bactrianus'') is least associated with humans and considered to be evolutionarily closer to the ancestral noncommensal populations. Usually, the house sparrow is regarded as a pest, since it consumes agricultural products and spreads disease to humans and their domestic animals. Even Birdwatching, birdwatchers often hold it in little regard because of its molestation of other birds. In most of the world, the house sparrow is not protected by law. Attempts to control house sparrows include the trapping, poisoning, or shooting of adults; the destruction of their nests and eggs; or less directly, blocking nest holes and scaring off sparrows with noise, glue, or porcupine wire. However, the house sparrow can be beneficial to humans, as well, especially by eating insect pests, and attempts at the large-scale control of the house sparrow have failed. The house sparrow has long been used as a food item. From around 1560 to at least the 19th century in northern Europe, earthenware "sparrow pots" were hung from eaves to attract nesting birds so the young could be readily harvested. Wild birds were trapped in nets in large numbers, and sparrow pie was a traditional dish, thought, because of the association of sparrows with Lascivious#Lechery, lechery, to have aphrodisiac properties. A traditional Indian medicine, ''Ciṭṭukkuruvi Lehyam, lēkiyam'' in Tamil language, Tamil, was sold with similar aphrodisiac claims. Sparrows were also trapped as food for falconry, falconers' birds and zoo animals. During the 1870s, there were debates on the damaging effects of sparrows in the House of Commons in England. In the early part of the 20th century, sparrow clubs culled many millions of birds and eggs in an attempt to control numbers of this perceived pest, but with only a localised impact on numbers. House sparrows have been kept as pets at many times in history, though they have no bright plumage or attractive songs, and raising them is difficult. The house sparrow has an extremely large range and population, so it is assessed as least concern for conservation on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
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Population decline

The IUCN estimates for the global population runs up to nearly 1.4 billion individuals, second among all birds perhaps only to the red-billed quelea in abundance (although the quelea is, unlike the sparrow, restricted to a single continent and has never been subject to human introductions). However, populations have been declining in many parts of the world, especially near its Eurasian places of origin. These declines were first noticed in North America, where they were initially attributed to the spread of the house finch, but have been most severe in Western Europe. Declines have not been universal, as no serious declines have been reported from Eastern Europe, but have even occurred in Australia, where the house sparrow was introduced recently. In Great Britain, populations peaked in the early 1970s, but have since declined by 68% overall, and about 90% in some regions. The RSPB lists the house sparrow's UK conservation status as red. In London, the house sparrow almost disappeared from the central city. The numbers of house sparrows in the Netherlands have dropped in half since the 1980s, so the house sparrow is even considered an endangered species. This status came to widespread attention after a female house sparrow, referred to as the "Domino Day 2005 sparrow, Dominomus", was killed after knocking down dominoes arranged as part of an attempt to set a world record. These declines are not unprecedented, as similar reductions in population occurred when the internal combustion engine replaced horses in the 1920s and a major source of food in the form of grain spillage was lost. Declines have been particularly apparent even in North America, where the house sparrow is invasive in some states. Introduced to Philadelphia initially in 1852 the house sparrow rapidly spread across the nation. However, the bird has largely disappeared from the city nowadays and overall, it is estimated to have declined in North America by 84% since 1966. In South Asia, the house sparrow has largely vanished from major cities such as Karachi, Kolkata, Mumbai, New Delhi, and Lahore. Various causes for the dramatic decreases in population have been proposed, including predation, in particular by Eurasian sparrowhawks; electromagnetic radiation from mobile phones; and diseases such as avian malaria. A shortage of nesting sites caused by changes in urban building design is probably a factor, and conservation organisations have encouraged the use of special nest boxes for sparrows. A primary cause of the decline seems to be an insufficient supply of insect food for nestling sparrows. Declines in insect populations result from an increase of monoculture crops, the heavy use of pesticides, the replacement of native plants in cities with introduced plants and parking areas, and possibly the introduction of unleaded petrol, which produces toxic compounds such as methyl nitrite. Protecting insect habitats on farms and planting native plants in cities benefit the house sparrow, as does establishing urban green spaces. To raise awareness of threats to the house sparrow, World Sparrow Day has been celebrated on 20 March across the world since 2010. Over the recent years, the house sparrow population has been on the decline in many Asian countries, and this decline is quite evident in India. To promote the conservation of these birds, in 2012, the house sparrow was declared as the state bird of Delhi.


Cultural associations

To many people across the world, the house sparrow is the most familiar wild animal and, because of its association with humans and familiarity, it is frequently used to represent the common and vulgar, or the lewd. One of the reasons for the introduction of house sparrows throughout the world was their association with the European homeland of many immigrants. Birds usually described later as sparrows are referred to in many works of ancient literature and religious texts in Europe and western Asia. These references may not always refer specifically to the house sparrow, or even to small, seed-eating birds, but later writers who were inspired by these texts often had the house sparrow in mind. In particular, sparrows were associated by the ancient Greeks with Aphrodite, the goddess of love, due to their perceived lustfulness, an association echoed by later writers such as Chaucer and Shakespeare. Jesus's use of "sparrows" as an example of divine providence in the Gospel of Matthew also inspired later references, such as that in Shakespeare's ''Hamlet'' and the Gospel music, Gospel hymn ''His Eye Is on the Sparrow''.
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The house sparrow is very rarely represented in ancient Egyptian art, but an Egyptian hieroglyph is based on it. The sparrow hieroglyph had no phonetic value and was used as a determinative in words to indicate small, narrow, or bad. An alternative view is that the hieroglyph meant "a prolific man" or "the revolution of a year".


See also

*House bunting *House finch *House wren


References


Works cited

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External links

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House sparrow
at the Royal Society for the Protection of Birds website
Indian sparrow
an
house sparrow
at Birds of Kazakhstan
World Sparrow Day
{{Authority control Passer, house sparrow Cosmopolitan birds Stored-product pests Birds described in 1758, house sparrow Taxa named by Carl Linnaeus, house sparrow Articles containing video clips