Papilio Zagreus
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''Papilio zagreus'' is a
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
of the family
Papilionidae Swallowtail butterflies are large, colorful butterflies in the family Papilionidae, and include over 550 species. Though the majority are tropical, members of the family inhabit every continent except Antarctica. The family includes the larges ...
(swallowtails). It is found in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, including
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
,
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
,
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
,
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
and western
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
.


Description

A powerfully built insect with strong neuration in the forewing. The frons is either quite black or bears a yellow mesial line, never a yellow lateral streak along the eye. The antennae are long, yellow, with thin club; the frons has a yellow mesial stripe, the breast is diagonally streaked with yellow, the abdomen is for the most part yellow, the costal margin of the forewing is not dentate, the cell of the forewing is broad and the hindwing is rounded, without a tail. The spots of the forewing orange, the marginal ones yellow; hindwing orange, a marginal band enclosing a yellow submarginal spot, a basal subcostal area, a patch in the extremity of the cell, as well as several spots on the disc, black. The
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...
is 110–130 mm.


Biology

''Papilio zagreus'' is a palatable
Batesian mimic Batesian mimicry is a form of mimicry where a harmless species has evolved to imitate the warning signals of a harmful species directed at a predator of them both. It is named after the English naturalist Henry Walter Bates, after his work on bu ...
of various ''
Heliconius ''Heliconius'' comprises a colorful and widespread genus of brush-footed butterflies commonly known as the longwings or heliconians. This genus is distributed throughout the tropical and subtropical regions of the New World, from South America a ...
'' butterfly species.


Subspecies

*''Papilio zagreus zagreus'' (Colombia, Venezuela, Amazonas to Bolivia) *''Papilio zagreus ascolius'' C. & R. Felder, 1864 (Ecuador, Colombia, Panama) Hindwing without black spots in the cell and on the disc, basal area of the cell of the forewing always pure pale yellow, between the 3. radial and 1. median two spots touching the cell: cell of the hindwing and the adjoining parts of the disc orange.
Batesian mimic Batesian mimicry is a form of mimicry where a harmless species has evolved to imitate the warning signals of a harmful species directed at a predator of them both. It is named after the English naturalist Henry Walter Bates, after his work on bu ...
of various ''
Ithomiini Ithomiini is a butterfly tribe in the nymphalid subfamily Danainae. It is sometimes referred to as the tribe of clearwing butterflies or glasswing butterflies. Some authors consider the group to be a subfamily (Ithomiinae). These butterflies are ...
''. *'' Papilio zagreus bachus'' C. & R. Felder, 1865 (Colombia, Ecuador) The orange area of the hindwing of the preceding subspecies is here only represented by a narrow variable band: however, the veins are often more or less yellowish on the underside. *''Papilio zagreus zalates'' Godman & Salvin, 1890 (Panama, Nicaragua, Costa Rica) The most northern form. The cell area of the forewing is dusted with black, the subapical cell-spot is narrow, the discal spots are short, the marginal area of the hindwing narrower than in the other forms and the hindwing beneath deeper orange. *''Papilio zagreus rosenbergi'' Druce, 1903 (Ecuador) The subapical cell-spot of the forewing large, the discal spot between the 1. and 2. radial usually small, sometimes absent, rarely large, the following discal spots on the whole larger than in the preceding forms, whilst the posterior submarginal spots are smaller. Named for the London-based insect dealer W.F.M. Rosenberg. *''Papilio zagreus chrysomelus'' Rothschild & Jordan, 1906 (Peru, Bolivia, Ecuador) The forewing orange above and beneath from the base to the disc, at the costal margin more or less pale yellow. *''Papilio zagreus daguanus'' Rothschild & Jordan, 1906 (Colombia) The cell-spot of the forewing as in ''zalates'', the discal spots on the contrary as in ''ascolius'', the discal spot between the 1. and 2. radial much shorter than the one placed behind it; hindwing pale, much less orange than in ''ascolius'', the black basal streak broad, entering the cell, behind this streak a large black spot on the disc. *''Papilio zagreus batesi'' (Racheli & Racheli, 1996) (northern Peru) *''Papilio zagreus chrysoxanthus'' Fruhstorfer, 1915 (central Peru, southern Peru to Bolivia) *''Papilio zagreus nigroapicalis'' (Bollino & Sala, 1998) (Peru) *''Papilio zagreus baueri'' (Möhn, 2001) (Amazonas)


Taxonomy

''Papilio zagreus'' is in the ''Papilio zagreus''
species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
. This
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
has two members. *''Papilio zagreus'' *'' Papilio neyi'' (may be a form of ''zagreus'') ''P.ascolius'' is ranked as a species in earlier works. The status was changed to subspecies by Racheli and Parise in 1992 and this was accepted by Tyler, Brown and Wilson in 1994.Tyler, H.A., Brown, Jr., K.S. & Wilson, K.H. 1994. ''Swallowtail Butterflies of the Americas''. Scientific Publishers, Gainesville, Florida


References

*Lewis, H. L., 1974 ''Butterflies of the World'' Page 25, figure 17


External links


Butterflycorner
* {{Taxonbar, from=Q3014675
zagreus In ancient Greek religion and mythology, Zagreus ( grc-gre, Ζαγρεύς) was sometimes identified with a god worshipped by the followers of Orphism, the "first Dionysus", a son of Zeus and Persephone, who was dismembered by the Titans and reb ...
Papilionidae of South America Butterflies described in 1847