Pacullidae is a

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

araneomorph The Araneomorphae (also called the Labidognatha) are an infraorder of spiders. They are distinguishable by chelicerae (fangs) that point diagonally forward and cross in a pinching action, in contrast to the Mygalomorphae (tarantulas and their cl ...

spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...

s first described by

Eugène Simon Eugène Louis Simon (; 30 April 1848 – 17 November 1924) was a French naturalist who worked particularly on insects and spiders, but also on birds and plants. He is by far the most prolific spider taxonomist in history, describing over 4, ...

in 1894. It was merged into

Tetrablemmidae Tetrablemmidae, sometimes called armored spiders, is a family of tropical araneomorph spiders first described by Octavius Pickard-Cambridge in 1873. It contains 126 described species in 29 genera from southeast Asia, with a few that occur in Afric ...

in 1958, then raised back to family status after a large phylogenetic study in 2017.

Description

The family Pacullidae contains three-clawed spiders with six eyes, lacking a

cribellum Cribellum literally means "little sieve", and in biology the term generally applies to anatomical structures in the form of tiny perforated plates. In certain groups of diatoms it refers to microscopically punctured regions of the frustule, or ...

. They resemble spiders from the family Tetrablemmidae in some respects but are much larger, always exceeding 5 mm long, have a very wrinkled (rugose) cuticle, and females do not have large membranous receptacles.

Phylogeny

Pacullidae falls within the

Synspermiata Synspermiata is a clade of araneomorph spiders, comprising most of the former " haplogynes". They are united by having simpler genitalia than other araneomorph spiders, lacking a cribellum, and sharing an evolutionary history of synspermia – a ...

clade, a clade of former

haplogyne The Haplogynae or haplogynes are one of the two main groups into which araneomorph spiders have traditionally been divided, the other being the Entelegynae. Morphological phylogenetic studies suggested that the Haplogynae formed a clade; more recen ...

spiders with "synsperm" – encapsulated groups of 2–4 fused sperm cells. Within this clade, it groups with four other families, including Tetrablemmidae, but is distinct from the latter, being most closely related to

Diguetidae Coneweb spiders (Diguetidae) are six-eyed haplogyne spiders that live in tangled space webs, fashioning a cone-like central retreat where they hide and lay eggs. It is a small family, containing only two genera with fifteen species and is confin ...

. Together with

Pholcidae The Pholcidae are a family of araneomorph spiders. The family contains over 1,800 individual species of pholcids, including those commonly known as cellar spider, daddy long-legs spider, carpenter spider, daddy long-legger, vibrating spid ...

, these four families are placed in the "lost trachea clade", a group of families that have lost their posterior respiratory system.

Genera and species

, the

World Spider Catalog The World Spider Catalog (WSC) is an online searchable database concerned with spider taxonomy. It aims to list all accepted families, genera and species, as well as provide access to the related taxonomic literature. The WSC began as a series of ...

accepts the following genera and species: *'' Lamania'' Lehtinen, 1981 **'' Lamania bernhardi'' (Deeleman-Reinhold, 1980) — Borneo **'' Lamania bokor'' Schwendinger & Košulič, 2015 — Cambodia **'' Lamania gracilis'' Schwendinger, 1989 — Bali **'' Lamania inornata'' (Deeleman-Reinhold, 1980) — Borneo **'' Lamania kraui'' (Shear, 1978) — Thailand, Malaysia **'' Lamania lipsae'' Dierkens, 2011 — Borneo **'' Lamania nirmala'' Lehtinen, 1981 — Borneo **'' Lamania sheari'' (Brignoli, 1980) — Indonesia (Sulawesi) *'' Paculla'' Simon, 1887 **'' Paculla bukittimahensis'' Lin & Li, 2017 — Singapore **'' Paculla cameronensis'' Shear, 1978 — Malaysia **'' Paculla globosa'' Lin & Li, 2017 — Singapore **'' Paculla granulosa'' (Thorell, 1881) — New Guinea **'' Paculla mului'' Bourne, 1981 — Borneo **'' Paculla negara'' Shear, 1978 — Malaysia **'' Paculla sulaimani'' Lehtinen, 1981 — Malaysia **'' Paculla wanlessi'' Bourne, 1981 — Borneo *'' Perania'' Thorell, 1890 **'' Perania annam'' Schwendinger & Košulič, 2015 — Vietnam **'' Perania armata'' (Thorell, 1890) — Indonesia (Sumatra) **'' Perania birmanica'' (Thorell, 1898) — Myanmar **'' Perania cerastes'' Schwendinger, 1994 — Malaysia **'' Perania coryne'' Schwendinger, 1994 — Malaysia **'' Perania deelemanae'' Schwendinger, 2013 — Indonesia (Sumatra) **'' Perania egregia'' Schwendinger, 2013 — Thailand **'' Perania ferox'' Schwendinger, 2013 — Thailand **'' Perania harau'' Schwendinger, 2013 — Indonesia (Sumatra) **'' Perania korinchica'' Hogg, 1919 — Indonesia (Sumatra) **'' Perania nasicornis'' Schwendinger, 1994 — Thailand **'' Perania nasuta'' Schwendinger, 1989 — Thailand **'' Perania nigra'' (Thorell, 1890) — Indonesia (Sumatra) **'' Perania picea'' (Thorell, 1890) — Indonesia (Sumatra) **'' Perania quadrifurcata'' Schwendinger, 2013 — Thailand **'' Perania robusta'' Schwendinger, 1989 — China, Thailand **'' Perania selatan'' Schwendinger, 2013 — Indonesia (Sumatra) **'' Perania siamensis'' Schwendinger, 1994 — Thailand **'' Perania tumida'' Schwendinger, 2013 — Thailand **'' Perania utara'' Schwendinger, 2013 — Indonesia (Sumatra) *'' Sabahya'' Deeleman-Reinhold, 1980 **'' Sabahya bispinosa'' Deeleman-Reinhold, 1980 — Borneo **'' Sabahya kinabaluana'' Deeleman-Reinhold, 1980 — Borneo *†'' Furcembolus'' Wunderlich 2008 9 spp,

Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...

, Myanmar,

Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...

References

{{Taxonbar, from=Q33134682 Araneomorphae families