Pyrgus Serratulae
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The Olive Skipper (''Pyrgus serratulae'') is a species of skipper (family
Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy ...
).


Description

This is a relatively distinctive species by the standards of the genus, the upperside tending to be plainer than most of its congeners with only tiny white marks on the forewings and almost unmarked on the hindwings. The underside is usually mostly olive-green with paler markings. An oval basal spot on the underside of the hind wing is a mostly well-developed feature. Alpine individuals often have a strong, almost completely reduced pattern of bright points on the upper side of the wing. The
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...
is 24–28 mm which means that ''Pyrgus serratulae'' is a medium-sized ''
Pyrgus ''Pyrgus'' is a genus in the skippers butterfly family, Hesperiidae, known as the grizzled skippers. The name "checkered" or "chequered skipper" may also be applied to some species, but also refers to species in the genera '' Burnsius'' and '' ...
'' species. Basically, there is a great possibility of confusion with butterflies of the ''
Pyrgus alveus The Large Grizzled skipper (''Pyrgus alveus'') is a species of skipper butterfly (family Hesperiidae). Description This is a rather variable species with a wingspan of 22–32 mm (not always particularly "large") with several subspecies having ...
'' complex, which can also occur syntopically with ''P. serratulae'', as well as the other species of the genus ''Pyrgus''. Safe separation is possible through the genitals.


Range

It is a species of the mountainous areas of Europe but is absent from the British Isles and
Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion#Europe, subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, ...
. ''P. serratulae'' is distributed from the southern European mountains through western, central and eastern Europe via south-western Siberia to
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
and northern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. In addition, Asia Minor,
Transcaucasia The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and Western Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Arme ...
as far as north-west
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
and north-east
Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq ...
are populated. In north-west
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
the occurrences reach as far as
East Westphalia Ostwestfalen-Lippe (, literally ''East(ern) Westphalia-Lippe'', abbreviation OWL) is the eastern region of the German state of North Rhine-Westphalia, congruent with the administrative region of Detmold and containing the eastern part of Westph ...
and in north-east Germany to the vicinity of the
Baltic Sea The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden and the North and Central European Plain. The sea stretches from 53°N to 66°N latitude and from ...
.


Habitat

Habitats are limestone grasslands or alpine grasslands in shallow and often rock-interspersed locations as well as rock corridors on mostly alkaline rocks. Avalanche tracks and erosion channels are often occupied in the Alps. In addition, the species is also found in ''Nardus stricta'' dominated grasslands.


Ecology

The adults are on the wing in July and August. The phenology of ''Pyrgus serratulae'' differs over a long period of time due to its wide altitude distribution. While the flight time in the Alps begins from mid-June (in above-average warm springs from the beginning of June) with the peak in July and numerous records still in August to the beginning of September, the imagos appear in the lower elevations of northern Bavaria a month earlier, usually in May (in very warm springs from the end of April) with a focus on June. The last imagos can be found in some years until the end of July. The imagos of ''Pyrgus serratulae'' obviously orient themselves optically on smaller or larger rocks, rock edges or banks or on other open ground structures. The oviposition takes place in very gappy or vegetation-poor locations, usually on individually standing cinquefoil plants. Proven caterpillar forage plants are '' Potentilla tabernaemontani'' ''Potentilla neumanniana'', ''Potentilla verna'' ''
Potentilla heptaphylla ''Potentilla'' is a genus containing over 300Guillén, A., et al. (2005)Reproductive biology of the Iberian species of ''Potentilla'' L. (Rosaceae).''Anales del Jardín Botánico de Madrid'' 1(62) 9–21. species of annual, biennial and peren ...
'', ''
Potentilla recta ''Potentilla recta'', the sulphur cinquefoil or rough-fruited cinquefoil, is a species of cinquefoil. It is native to Eurasia but it is present in North America as an introduced species, ranging through almost the entire continent except the no ...
'', ''
Potentilla aurea ''Potentilla aurea'', the golden cinquefoil, is a species of flowering plant in the family Rosaceae. It is native to the mountains of mainland Europe, and has been introduced to Turkey. A number of cultivars are available, including 'Aurantiaca' ...
'', ''
Potentilla crantzii ''Potentilla crantzii'', the alpine cinquefoil, is a flowering plant in the family Rosaceae. It is native to North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It ...
'', ''
Potentilla hirta ''Potentilla'' is a genus containing over 300Guillén, A., et al. (2005)Reproductive biology of the Iberian species of ''Potentilla'' L. (Rosaceae).''Anales del Jardín Botánico de Madrid'' 1(62) 9–21. species of annual, biennial and perenni ...
'', ''
Potentilla pusilla ''Potentilla'' is a genus containing over 300Guillén, A., et al. (2005)Reproductive biology of the Iberian species of ''Potentilla'' L. (Rosaceae).''Anales del Jardín Botánico de Madrid'' 1(62) 9–21. species of annual, biennial and perenni ...
'', '' Potentilla reptans'', '' Sibbaldia montanumbens'' and '' Heliantum reptanumbens''. The eggs are deposited individually on the underside of the leaves. As the oviposition observations and caterpillar finds show, larval development takes place in full sun and quickly warming locations. Cinquefoil plants on or immediately next to rocks are often used. Furthermore, open locations with little vegetation with a rocky-gritty soil matrix or along unpaved paths are used for larval development. The caterpillars spin the typical ''
Pyrgus ''Pyrgus'' is a genus in the skippers butterfly family, Hesperiidae, known as the grizzled skippers. The name "checkered" or "chequered skipper" may also be applied to some species, but also refers to species in the genera '' Burnsius'' and '' ...
'' leaf bags. The most important food plant in the extra-alpine area is the species group ''Potentilla verna agg''. While so far only a few egg-laying observations have been made on ''
Potentilla erecta ''Potentilla erecta'' (syn. ''Tormentilla erecta'', ''Potentilla laeta'', ''Potentilla tormentilla'', known as the (common) tormentil, septfoil or erect cinquefoil ) is a herbaceous perennial plant belonging to the rose family (Rosaceae). Descri ...
''. In the
Swabian Alb The Swabian Jura (german: Schwäbische Alb , more rarely ), sometimes also named Swabian Alps in English, is a mountain range in Baden-Württemberg, Germany, extending from southwest to northeast and in width. It is named after the region of ...
, ''
Potentilla heptaphylla ''Potentilla'' is a genus containing over 300Guillén, A., et al. (2005)Reproductive biology of the Iberian species of ''Potentilla'' L. (Rosaceae).''Anales del Jardín Botánico de Madrid'' 1(62) 9–21. species of annual, biennial and peren ...
'' is preferred over ''Potentilla verna''. In the Alps, in particular, other ''
Potentilla ''Potentilla'' is a genus containing over 300Guillén, A., et al. (2005)Reproductive biology of the Iberian species of ''Potentilla'' L. (Rosaceae).''Anales del Jardín Botánico de Madrid'' 1(62) 9–21. species of annual, biennial and perenni ...
'' species should be considered as caterpillar food. The northern Bavarian caterpillars overwinter in the penultimate caterpillar stadium and can already be found adult in April. A longer-range in larval development can possibly be expected in the Alps. The males of ''Pyrgus serratulae'' like to suckle on moist ground as well as on sweat. When visiting flowers, the imagos are opportunistic and visit flowers from several families and with different colors.


Conservation

In the Alps, no special protective measures currently appear necessary. The preservation of the extra-alpine occurrences, on the other hand, is heavily dependent on maintenance measures such as rock clearing and regular grazing or clearing to keep the limestone grasslands open. For this purpose, steep rock heaths, as well as shallow and rock-interspersed grasslands, are to be integrated.


Etymology

"''
Serratula ''Serratula'' is a genus of plants in the tribe Cardueae within the family Asteraceae native to Eurasia. Plumeless saw-wort is a common name for plants in this genus. ''Serratula'' as traditionally defined contains at least two groups: one of whi ...
''" As with most ''Pyrgus'' species, the plant genus mentioned in the species name has nothing to do with the caterpillar's diet.


References


External links


Lepiforum.deFauna EuropaeaButterfly Conservation Armenia
{{Taxonbar, from=Q1904094 Pyrgus Butterflies described in 1839 Butterflies of Europe