Pyrgus Cirsii
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The Cinquefoil Skipper (''Pyrgus cirsii'') is a species of skipper butterfly (family Hesperiidae).


Description

This species has a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan o ...
of 26–28 mm and, typically of its
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
, can be very difficult to identify in the field. ''Pyrgus cirsii'' is a medium-sized skipper. There is basically a possibility of confusion with other species of the genus '' Pyrgus'', in particular with '' Pyrgus armoricanus''. Characteristic of ''Pyrgus cirsii'' is a large square discoidal spot on the upper side of the forewing and almost always a second point-like mark over the basal spot on the upper side of the forewing. The hind wing underside is often ocher to reddish brown and stands out from other species of the genus. It is sometimes considered a subspecies of the Carline Skipper (''P. carlinae'') as the two are known to
interbreed In biology, a hybrid is the offspring resulting from combining the qualities of two organisms of different breeds, varieties, species or genera through sexual reproduction. Hybrids are not always intermediates between their parents (such as in ...
where their ranges overlap in the Alps but can usually be recognised by the prominent squarish, rather than c-shaped, white spot near the
costa Costa may refer to: Biology * Rib (Latin: ''costa''), in vertebrate anatomy * Costa (botany), the central strand of a plant leaf or thallus * Costa (coral), a stony rib, part of the skeleton of a coral * Costa (entomology), the leading edge of th ...
of the forewing and the usually deeper red-brown underwings. A reliable separation in this genus, which is difficult to determine according to external characteristics, is possible through genital examination.


Range

''Pyrgus cirsii'' is widespread from Spain to Central Europe. According to current knowledge, there is also evidence from the Caucasus and Eastern Anatolia, as there are several isolated populations in north-eastern
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula in ...
and in
Armenia Armenia (), , group=pron officially the Republic of Armenia,, is a landlocked country in the Armenian Highlands of Western Asia.The UNbr>classification of world regions places Armenia in Western Asia; the CIA World Factbook , , and ' ...
.Butterfly Conservation Armenia http://www.butterfly-conservation-armenia.org/pyrgus-cirsii.html


Habitat

The current habitats of ''Pyrgus cirsii'' are sparsely overgrown, with open ground interspersed, full sun dry grasslands in stony, shallow and base-rich locations. Almost without exception, the current habitats are subject to regular and relatively intensive grazing, and they are also characterized by large pads of ''Potentilla verna'' agg. In the past, extremely sparse pine stands were also populated in Bavaria. In addition to these qualitative characteristics, the overall size of the habitat obviously plays an important role. E. g. all Bavarian occurrences are or were in areas with large, contiguous grassland complexes. A minimum size of the living space with corresponding open ground structures or the very close connection of these dry turf areas is obviously essential.


Ecology

''Pyrgus cirsii'' forms one generation per year, at least in Bavaria. It appears relatively late in the year from late July to mid-September. As a rule, the flight season begins at the end of July / beginning of August with a peak in the second half of August. The last moths, mostly females, can be observed in mid-September. The reports of finds in Bavaria range from 160 m in Main Franconia to around 500 m in the Northern
Franconian Jura The Franconian Jura ( , , or ) is an upland in Franconia, Bavaria, Germany. Located between two rivers, the Danube in the south and the Main in the north, its peaks reach elevations of up to and it has an area of some 7053.8 km2. Emil Meyn ...
, with a focus of 280 m to 400 m. When visiting flowers, the imagos (males and females) behave opportunistically and use a wide range of different colored flowers. The males also like to suckle on moist soil. ''Pyrgus cirsii'' can occur in numbers in suitable habitats. According to current knowledge, ''Pyrgus cirsii'' caterpillars probably lives exclusively on cinquefoil species (
Potentilla ''Potentilla'' is a genus containing over 300Guillén, A., et al. (2005)Reproductive biology of the Iberian species of ''Potentilla'' L. (Rosaceae).''Anales del Jardín Botánico de Madrid'' 1(62) 9–21. species of annual, biennial and perenni ...
, Rosaceae). In Central Europe by far the most important is the spring cinquefoil (''Potentilla verna'' agg. = ''
Potentilla tabernaemontani ''Potentilla neumanniana'', the spring cinquefoil or spotted cinquefoil, is a perennial flowering plant in the rose family (Rosaceae). It may grow up to the height of 5–15 cm. It was first scientifically described by H.G.L. Reichenbach i ...
''). Other proven caterpillar forage plants are '' Potentilla cinerea, Potentilla incana'' ''Potentilla arenaria'' '' Potentilla hirta'' and '' Potentilla pusilla''. ''Potentilla verna'' agg. has so far been proven as a larval food and egg-laying plant from
Bavaria Bavaria ( ; ), officially the Free State of Bavaria (german: Freistaat Bayern, link=no ), is a state in the south-east of Germany. With an area of , Bavaria is the largest German state by land area, comprising roughly a fifth of the total lan ...
. ''Potentilla verna'' agg. is also mentioned as a host plant from
Baden-Württemberg Baden-Württemberg (; ), commonly shortened to BW or BaWü, is a German state () in Southwest Germany, east of the Rhine, which forms the southern part of Germany's western border with France. With more than 11.07 million inhabitants across a ...
and Switzerland. '' Potentilla incana'' is given for
Alsace Alsace (, ; ; Low Alemannic German/ gsw-FR, Elsàss ; german: Elsass ; la, Alsatia) is a cultural region and a territorial collectivity in eastern France, on the west bank of the upper Rhine next to Germany and Switzerland. In 2020, it had ...
. After laying eggs in August and September, the caterpillar overwinters until the coming spring. The first newly hatched caterpillars were found on the Swabian Alb in March. The caterpillars live in the leaf bags characteristic of the '' Pyrgus'' species and can therefore be searched for in a targeted manner. The development of the caterpillar to the imagos takes a relatively long time with over three months. Both sexes of ''Pyrgus cirsii'' are based on the structure of the terrain and specifically seek out the areas with little vegetation. The males occupy territories here and defend them against their male counterparts. Almost all imagos that fly by are checked for a short period of time. Females that are ready to lay eggs fly very low and land on the sparse vegetation of the raw soil in search of places to lay their eggs. Recognizing the right food plant is obviously not possible "from the air", which is shown by the chance landing on very different plant species. After landing, it immediately checks the suitability of the plant by lowering the feelers on the leaves. If it is not a cinquefoil, the female continues to look for neighboring plants in the area, which are checked again in this way. If the search for suitable cinquefoil plants is successful, various leaves of the same are thoroughly drummed with the antennae. If a leaf is finally assessed as suitable, the eggs are usually laid on the underside of the leaf, sometimes also on the stem of the food plant.


Conservation

Like most ''Pyrgus'' species, ''Pyrgus cirsii'' needs a fairly large habitat and colonizes xerothermal limestone grasslands. The caterpillars are only found in the leanest, driest places. These places are in decline due to eutrophication, among other things from the air, decline in migratory sheep farming, in addition to the general loss of habitat due to loss of all kinds of grassland communities and climate changes. Increasing isolation might be a main reason for the strongly regressive population development of the species. As soon as the vegetation becomes denser and ''
Potentilla ''Potentilla'' is a genus containing over 300Guillén, A., et al. (2005)Reproductive biology of the Iberian species of ''Potentilla'' L. (Rosaceae).''Anales del Jardín Botánico de Madrid'' 1(62) 9–21. species of annual, biennial and perenni ...
'' stands between denser, higher plants, no successful larval development and usually no more oviposition can be expected. For ''Pyrgus cirsii'', protective measures have been introduced and maintenance measures have been carried out in the
Steigerwald The Steigerwald is a hill region up to in the Bavarian-Franconian part of the South German Scarplands between Würzburg and Nuremberg. It is part of the Keuper Uplands, and within it, it is continued to the north-northeast and right of the river ...
since 1998 as part of a species aid program. This resulted in clearing of bushes and clearing of grasslands to enlarge the current habitats. At the same time, de-bushed areas were reintegrated into regular grazing and are now becoming more or more intensive, i.e. grazed at a higher frequency. The current habitats are all grazed with sheep and goats at least twice a year, usually much more frequently.


Taxonomy

Some authors only accept the separation of ''P. carlinae'' and ''P. cirsii'' at the subspecies level.


External links


Fauna EuropaeaLepiforum.deButterfly Conservation Armenia


References

; ;Bibliography * Database entry includes justification for why this species is vulnerable. Hesperiidae Pyrgus Butterflies described in 1839 Butterflies of Europe {{Pyrginae-stub