Pyrgus Cacaliae
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The Dusky Grizzled Skipper (''Pyrgus cacaliae'') is a species of skipper butterfly (family Hesperiidae).


Description

''Pyrgus cacaliae'' is comparatively large and the white spots on the upper side of the forewing are greatly reduced. The hind wing underside has a faded olive-green color with a long and a short point on the inner edge (similar to an exclamation mark). This feature also shows ''Pyrgus andromedae'', but in ''Pyrgus cacaliae'' it does not stand out so clearly from the subsurface. Also the Alpine grizzled skipper is usually encountered at lower altitudes. It can also be distinguished by the smaller, fainter pale markings on the
forewing Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwi ...
and the less bold markings on the underwing. In cases of doubt, a genital examination is essential for a reliable determination. Both sexes have the same markings, but the males are slightly darker in color. The caterpillar is usually dark in color with a darker topline. In ex ovo breeding, clay-yellow colored caterpillars were often observed in the last two stages. The head shield is black. The head and wing sheaths of the pupa are bluish with frosting; in the case of the abdomen, which is brown in its basic color, only the segment boundaries are left out. The ventral side has a distinctive drawing, consisting of a black central bar and each side of which consists of two black dots. The back side also shows black dots and lines. The dorsal cremaster is red-brown in colo


Range

The species is widespread in the Central
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to ...
and from the western to the eastern
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Swi ...
(from
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan area ...
to
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
, Switzerland,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
and
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
). In addition, ''Pyrgus cacaliae'' occurs in the
Balkans The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
in the Dinaric Mountains of Bosnia-Herzegovina, in the Central
Carpathians The Carpathian Mountains or Carpathians () are a range of mountains forming an arc across Central Europe. Roughly long, it is the third-longest European mountain range after the Urals at and the Scandinavian Mountains at . The range stretches ...
of
Romania Romania ( ; ro, România ) is a country located at the crossroads of Central, Eastern, and Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, Serbia to the southwest, Moldova to the east, and ...
and in the Rila,
Pirin , photo=Pirin-mountains-Bansko.jpg , photo_caption=Pirin scenery in winter , country= Bulgaria, , parent= , geology= granite, gneiss, marble, limestone , area_km2=2585 , range_coordinates = , length_km=80 , length_orientation= north-s ...
and Stara Planina Mountains of
Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Macedo ...
.


Habitat

On the one hand, ''Pyrgus cacaliae'' colonizes humid habitats such as rye bushes along streams, peatlands and springs. On the other hand, there is evidence from dry alpine lawns, mostly interlocking with rock debris. In the Alps, the maximum distribution is between 1700 and 2500 meters above sea level. In Bavaria this species is recorded at altitudes of 1500-2300 m.


Ecology

The flight period in the Bavarian Alpine region extends from the end of June to the beginning of August. ''Pyrgus cacaliae'' can be detected in both even and odd years. According to observations from Switzerland and Austria, the eggs are laid individually on the underside of the leaves of the host plants: ''
Potentilla erecta ''Potentilla erecta'' (syn. ''Tormentilla erecta'', ''Potentilla laeta'', ''Potentilla tormentilla'', known as the (common) tormentil, septfoil or erect cinquefoil ) is a herbaceous perennial plant belonging to the rose family ( Rosaceae). Descr ...
,
Potentilla crantzii ''Potentilla crantzii'', the alpine cinquefoil, is a flowering plant in the family Rosaceae. It is native to North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. ...
'' and '' Potentilla aurea''. The wintering was observed in Switzerland as a caterpillar in the penultimate stage. Other observations show that after an egg rest of eleven days, the caterpillars hatch, only eat plants for a few days and then prepare for wintering. Observations from the
Rätikon The Rätikon is a mountain range of the Central Eastern Alps, located at the border between Vorarlberg, Liechtenstein and Graubünden. It is the geological border between the Eastern and Western Alps and stretches from the Montafon as far as ...
and
Silvretta The Silvretta Alps are a mountain range of the Central Eastern Alps shared by Tirol, Vorarlberg (both in Austria) and Graubünden ( Switzerland). The Austrian states of Tirol and Vorarlberg are connected by a pass road ( Silvretta Hochalpenst ...
areas in
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
show a two-year development cycle with the first wintering in the first larval stage and a second wintering in the last larval or pupal stage. Imagos can be found every year due to a partial third hibernation.


Conservation

According to the current state of knowledge, the species is not endangered in the Alps, as the occurrence of ''Pyrgus cacaliae'' is mostly outside of agricultural areas or on extensively grazed mountain pastures.


Etymology

"'' Cacalia'', in German Roßlattich". "Roßlattich" is now mostly regarded as one of many German names for ''Tussilago farfara''. Originally, '' Adenostyles alpina'' ''Cacalia alpina, Tussilago cacalia''was very probably meant here. As with most ''Pyrgus'' species, it has nothing to do with the food of the caterpillar.


References


External links


Barcodes of lifeLepiforum.de
Pyrgus Butterflies described in 1839 Butterflies of Europe {{Pyrginae-stub