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''Pycnanthus angolensis'' is a species of
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
in the
nutmeg Nutmeg is the seed or ground spice of several species of the genus ''Myristica''. ''Myristica fragrans'' (fragrant nutmeg or true nutmeg) is a dark-leaved evergreen tree cultivated for two spices derived from its fruit: nutmeg, from its seed, an ...
family,
Myristicaceae The Myristicaceae are a family of flowering plants native to Africa, Asia, Pacific islands, and the Americas and has been recognized by most taxonomists. It is sometimes called the "nutmeg family", after its most famous member, ''Myristica fragra ...
. It is native to
Tropical Africa Although tropical Africa is mostly familiar to the West for its rainforests, this biogeographic realm of Africa is far more diverse. While the tropics are thought of as regions with hot moist climates, which are caused by latitude and the tropi ...
. Its English language common names include African nutmeg, false nutmeg, boxboard, and cardboard.''Pycnanthus angolensis''.
AgroForestryTree Database. World Agroforestry Centre.
In Africa it is widely known as ''ilomba''.Richter, H. G. and M. J. Dallwitz. 2000 onwards

Commercial timbers: descriptions, illustrations, identification, and information retrieval. DELTA – DEscription Language for TAxonomy. June, 2009.


Description

This
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
tree grows up to 40 meters tall and usually up to a meter wide, sometimes up to 1.5 meters or more. The trunk is straight and cylindrical with fissures and flaking bark. The
sap Sap is a fluid transported in xylem cells (vessel elements or tracheids) or phloem sieve tube elements of a plant. These cells transport water and nutrients throughout the plant. Sap is distinct from latex, resin, or cell sap; it is a separa ...
is honey-colored and turns red in time. The branches are in whorls. The leathery leaves are up to 31 centimeters long by 9 wide. The blades have pointed tips, heart-shaped bases, and thick midribs. They are hairless on top and coated with
rusty Rusty may refer to something covered with rust or with a rust (color). Rusty is also a nickname for people who have red hair, have a rust-hued skin tone, or have the given name Russell. Rusty may also refer to: People *Rusty Anderson (born 1959 ...
,
felt Felt is a textile material that is produced by matting, condensing and pressing fibers together. Felt can be made of natural fibers such as wool or animal fur, or from synthetic fibers such as petroleum-based acrylic or acrylonitrile or wood ...
like hairs on the undersides. The leaves usually bear signs of insect damage, a feature so common it is considered characteristic of the species. The flowers are arranged in dense, rusty
panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...
s up to 15 centimeters long. The individual flowers are difficult to see in the tight panicle until the
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s develop, being only about a millimeter long. The flowers are hairy and fragrant.''Pycnanthus angolensis'' (Welw.) Warb.
In: Stannard, B. L. ''Flora Zambesiaca'' Vol. 9, Part 2: Myristicaceae. 1997. Royal Botanic Gardens, Kew.
The fruit is a rounded
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
reaching over 3 centimeters long and wide, borne in clusters. It is hairy brown when new, turning yellow-orange, and has cartilaginous flesh that dries woody. It contains a black seed with a red
aril An aril (pronounced ), also called an arillus, is a specialized outgrowth from a seed that partly or completely covers the seed. An arillode or false aril is sometimes distinguished: whereas an aril grows from the attachment point of the see ...
which resembles that of
nutmeg Nutmeg is the seed or ground spice of several species of the genus ''Myristica''. ''Myristica fragrans'' (fragrant nutmeg or true nutmeg) is a dark-leaved evergreen tree cultivated for two spices derived from its fruit: nutmeg, from its seed, an ...
.''Pycnanthus angolensis''.
PROTA.
The fruit ripens over a long period continuing into the next flowering season, which begins around October.


Ecology

The tree grows in moist
rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...
s up to about 1200 meters in elevation among other evergreens and semideciduous trees. It occurs in
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has re-grown after a timber harvest or clearing for agriculture, until a long enough period has passed so that the effects of the disturbance are no longer evident. ...
, sometimes taking hold in new canopy gaps or clearings. It thrives in sunny locations. It grows in
riparian A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks ar ...
forests such as
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
s, and in some regions it can be found in
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s. It occurs in areas receiving between about 1300 and 1800 millimeters of rain per year, but the optimal seems to be about 2000 millimeters. Many animals feed on the fruits, such as
hornbill Hornbills (Bucerotidae) are a family (biology), family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a Casque (an ...
s.''Pycnanthus angolensis''.
An Introduction to the Trees from the North of the Republic of Congo. Royal Botanic Garden, Edinburgh.


Uses

This species has a wide variety of human uses. It is harvested for its
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin th ...
, which is light, soft, and whitish gray or pink-tinged in color. Its popularity rose after
World War II World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the vast majority of the world's countries—including all of the great powers—forming two opposin ...
when
plywood Plywood is a material manufactured from thin layers or "plies" of wood veneer that are glued together with adjacent layers having their wood grain rotated up to 90 degrees to one another. It is an engineered wood from the family of manufactured ...
was in demand, and during the mid-20th century it was one of the more valuable timbers in Central Africa. It is not very durable and tends to
warp Warp, warped or warping may refer to: Arts and entertainment Books and comics * WaRP Graphics, an alternative comics publisher * ''Warp'' (First Comics), comic book series published by First Comics based on the play ''Warp!'' * Warp (comics), a ...
, but it is easy to cut and work and can be used for many purposes. It is suitable for
furniture Furniture refers to movable objects intended to support various human activities such as seating (e.g., stools, chairs, and sofas), eating (tables), storing items, eating and/or working with an item, and sleeping (e.g., beds and hammocks). Fu ...
, and in house construction as
panelling Panelling (or paneling in the U.S.) is a millwork wall covering constructed from rigid or semi-rigid components. These are traditionally interlocking wood, but could be plastic or other materials. Panelling was developed in antiquity to make roo ...
,
siding Siding may refer to: * Siding (construction), the outer covering or cladding of a house * Siding (rail) A siding, in rail terminology, is a low-speed track section distinct from a running line or through route such as a main line, branch l ...
,
roof shingle Roof shingles are a roof covering consisting of individual overlapping elements. These elements are typically flat, rectangular shapes laid in courses from the bottom edge of the roof up, with each successive course overlapping the joints below. ...
s, and framing. It is used for
fuel A fuel is any material that can be made to react with other substances so that it releases energy as thermal energy or to be used for work. The concept was originally applied solely to those materials capable of releasing chemical energy but ...
and
paper pulp Pulp is a lignocellulosic fibrous material prepared by chemically or mechanically separating cellulose fibers from wood, fiber crops, waste paper, or rags. Mixed with water and other chemical or plant-based additives, pulp is the major raw mate ...
. The wood can be vulnerable to
termites Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattode ...
,
powderpost beetle Powderpost beetles are a group of seventy species of woodboring beetles classified in the insect subfamily Lyctinae. These beetles, along with spider beetles, death watch beetles, common furniture beetles, Dermestidae, skin beetles, and others ...
s, and other
pests PESTS was an anonymous American activist group formed in 1986 to critique racism, tokenism, and exclusion in the art world. PESTS produced newsletters, posters, and other print material highlighting examples of discrimination in gallery represent ...
. The yellowish or reddish
fat In nutrition science, nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such chemical compound, compounds, most commonly those that occur in living beings or in food. The term often refers spec ...
from the aromatic seed is called "kombo butter" or "Angola tallow". A seed can be up to 70% fat. It is used as a
fuel A fuel is any material that can be made to react with other substances so that it releases energy as thermal energy or to be used for work. The concept was originally applied solely to those materials capable of releasing chemical energy but ...
for lighting and is made into
soap Soap is a salt of a fatty acid used in a variety of cleansing and lubricating products. In a domestic setting, soaps are surfactants usually used for washing, bathing, and other types of housekeeping. In industrial settings, soaps are use ...
. Seed remnants are used in
compost Compost is a mixture of ingredients used as plant fertilizer and to improve soil's physical, chemical and biological properties. It is commonly prepared by decomposing plant, food waste, recycling organic materials and manure. The resulting m ...
. When ignited, the oily seed burns slowly and can be used as a
candle A candle is an ignitable wick embedded in wax, or another flammable solid substance such as tallow, that provides light, and in some cases, a fragrance. A candle can also provide heat or a method of keeping time. A person who makes candles i ...
. In
Uganda }), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...
the tree is grown in
banana A banana is an elongated, edible fruit – botanically a berry – produced by several kinds of large herbaceous flowering plants in the genus ''Musa''. In some countries, bananas used for cooking may be called "plantains", distinguis ...
,
coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulant, stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. S ...
, and
cocoa Cocoa may refer to: Chocolate * Chocolate * ''Theobroma cacao'', the cocoa tree * Cocoa bean, seed of ''Theobroma cacao'' * Chocolate liquor, or cocoa liquor, pure, liquid chocolate extracted from the cocoa bean, including both cocoa butter and ...
plantations to shade the crops. Most parts of the tree have been used in
traditional African medicine Traditional African medicine is a range of traditional medicine disciplines involving Indigenous peoples of Africa, indigenous herbalism and Traditional African religions, African spirituality, typically including divination, diviners, midwives, ...
. The sap has been used to control bleeding. The bark has been used as a poison
antidote An antidote is a substance that can counteract a form of poisoning. The term ultimately derives from the Greek term φάρμακον ἀντίδοτον ''(pharmakon) antidoton'', "(medicine) given as a remedy". Antidotes for anticoagulants are s ...
and a treatment for
leprosy Leprosy, also known as Hansen's disease (HD), is a long-term infection by the bacteria ''Mycobacterium leprae'' or ''Mycobacterium lepromatosis''. Infection can lead to damage of the nerves, respiratory tract, skin, and eyes. This nerve damag ...
,
anemia Anemia or anaemia (British English) is a blood disorder in which the blood has a reduced ability to carry oxygen due to a lower than normal number of red blood cells, or a reduction in the amount of hemoglobin. When anemia comes on slowly, th ...
,
infertility Infertility is the inability of a person, animal or plant to reproduce by natural means. It is usually not the natural state of a healthy adult, except notably among certain eusocial species (mostly haplodiploid insects). It is the normal state ...
,
gonorrhea Gonorrhea, colloquially known as the clap, is a sexually transmitted infection (STI) caused by the bacterium '' Neisseria gonorrhoeae''. Infection may involve the genitals, mouth, or rectum. Infected men may experience pain or burning with ...
, and
malaria Malaria is a mosquito-borne infectious disease that affects humans and other animals. Malaria causes symptoms that typically include fever, tiredness, vomiting, and headaches. In severe cases, it can cause jaundice, seizures, coma, or death. S ...
. Leaf extracts are consumed or used in an
enema An enema, also known as a clyster, is an injection of fluid into the lower bowel by way of the rectum.Cullingworth, ''A Manual of Nursing, Medical and Surgical'':155 The word enema can also refer to the liquid injected, as well as to a device ...
to treat
edema Edema, also spelled oedema, and also known as fluid retention, dropsy, hydropsy and swelling, is the build-up of fluid in the body's Tissue (biology), tissue. Most commonly, the legs or arms are affected. Symptoms may include skin which feels t ...
. Root extracts are used to treat parasitic infections, such as
schistosomiasis Schistosomiasis, also known as snail fever, bilharzia, and Katayama fever, is a disease caused by parasitic flatworms called schistosomes. The urinary tract or the intestines may be infected. Symptoms include abdominal pain, diarrhea, bloody s ...
. The seed oil is used to treat thrush. Like the fat of nutmeg, kombo butter is mostly
myristic acid Myristic acid (IUPAC name: tetradecanoic acid) is a common saturated fatty acid with the molecular formula CH3(CH2)12COOH. Its salts and esters are commonly referred to as myristates or tetradecanoates. It is named after the binomial name for nutm ...
, with a high amount of
myristoleic acid Myristoleic acid, or 9-tetradecenoic acid, is an omega-5 fatty acid. It is biosynthesized from myristic acid by the enzyme Stearoyl-CoA desaturase-1, but it is uncommon in nature. One of the major sources of this fatty acid is the seed oil from ...
, as well. It contains the unique compound kombic acid, which was named for the tree under its nomenclatural synonym, ''P. kombo''.


Cultivation

This tree is cultivated for its products. Seeds are sown in the field, and seedlings are grown in
plant nurseries A nursery is a place where plants are propagated and grown to a desired size. Mostly the plants concerned are for gardening, forestry or conservation biology, rather than agriculture. They include retail nurseries, which sell to the general p ...
until they have large
taproot A taproot is a large, central, and dominant root from which other roots sprout laterally. Typically a taproot is somewhat straight and very thick, is tapering in shape, and grows directly downward. In some plants, such as the carrot, the taproo ...
s. They cannot tolerate
drought A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, an ...
. After a year the tree is about half a meter tall, and within 4 years it can reach 4 meters. A 20-year-old tree can be 25 meters tall. Stands of cultivated trees are pruned and thinned periodically. Pest insects such as ''
Monochamus scabiosus ''Monochamus scabiosus'' is a species of beetle in the family Cerambycidae. It was described by Quedenfeldt in 1882, originally under the genus ''Monohammus ''Monochamus'' is a genus of longhorn beetles found throughout the world. They are co ...
'' and '' Mallodon downesi'' and
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
such as ''
Ophiostoma ''Ophiostoma'' is a genus of fungi within the family Ophiostomataceae. It was circumscribed in 1919 by mycologists Hans Sydow and Paul Sydow. Species *''Ophiostoma adjuncti'' *'' Ophiostoma ainoae'' *'' Ophiostoma allantosporum'' *'' Ophiosto ...
'' sp. have been observed, but they are not severe problems. The freshly harvested wood may become discolored due to
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among ...
l growth, the light-colored wood developing dark brown staining. Trees can be cut when they reach a diameter of 50 centimeters, at around 30 years old.
Agricultural cooperative An agricultural cooperative, also known as a farmers' co-op, is a cooperative in which farmers pool their resources in certain areas of activity. A broad typology of agricultural cooperatives distinguishes between agricultural service cooperati ...
s in Ghana cultivate the trees for the kombo butter.


Common names

There are
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
s for the plant in many languages. It is called ''mkungu mwitu'' in Swahili, ''akwa-mili'' and ''oje'' in
Igbo Igbo may refer to: * Igbo people, an ethnic group of Nigeria * Igbo language, their language * anything related to Igboland, a cultural region in Nigeria See also * Ibo (disambiguation) * Igbo mythology * Igbo music * Igbo art * * Igbo-Ukwu, a ...
, ''lunaba'' and ''munaba'' in
Luganda The Ganda language or Luganda (, , ) is a Bantu language spoken in the African Great Lakes region. It is one of the major languages in Uganda and is spoken by more than 10 million Baganda and other people principally in central Uganda including ...
, ''akomu'' in
Yoruba The Yoruba people (, , ) are a West African ethnic group that mainly inhabit parts of Nigeria, Benin, and Togo. The areas of these countries primarily inhabited by Yoruba are often collectively referred to as Yorubaland. The Yoruba constitute ...
, ''calabo'' in Spanish, and ''arbre à suif'' and ''faux muscadier'' in French. Local names include ''pó casson'' in
São Tomé and Príncipe São Tomé and Príncipe (; pt, São Tomé e Príncipe (); English: " Saint Thomas and Prince"), officially the Democratic Republic of São Tomé and Príncipe ( pt, República Democrática de São Tomé e Príncipe), is a Portuguese-speaking i ...
, ''gboyei'' in
Sierra Leone Sierra Leone,)]. officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered by Liberia to the southeast and Guinea surrounds the northern half of the nation. Covering a total area of , Sierra ...
and
Liberia Liberia (), officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to its north, Ivory Coast to its east, and the Atlantic Ocean ...
, ''otie'' in
Ghana Ghana (; tw, Gaana, ee, Gana), officially the Republic of Ghana, is a country in West Africa. It abuts the Gulf of Guinea and the Atlantic Ocean to the south, sharing borders with Ivory Coast in the west, Burkina Faso in the north, and To ...
, ''eteng'' in
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
, ''lolako'' in
Zaire Zaire (, ), officially the Republic of Zaire (french: République du Zaïre, link=no, ), was a Congolese state from 1971 to 1997 in Central Africa that was previously and is now again known as the Democratic Republic of the Congo. Zaire was, ...
,''Pycnanthus angolensis''.
Wood Technical Fact Sheet. Forest Products Laboratory. Forest Service, USDA.
and ''adria'', ''effoi'', ''hétéré'', ''qualélé'', and ''walélé'' in
Ivory Coast Ivory Coast, also known as Côte d'Ivoire, officially the Republic of Côte d'Ivoire, is a country on the southern coast of West Africa. Its capital is Yamoussoukro, in the centre of the country, while its largest city and economic centre is ...
.


References

{{Taxonbar, from=Q6093520 Myristicaceae Trees of Africa Plants used in traditional African medicine Wood