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The purple-throated mountaingem (''Lampornis calolaemus'') is a species of hummingbird in tribe
Lampornithini Lampornithini is one of the three tribes that make up the subfamily Trochilinae in the hummingbird family Trochilidae. The other two tribes in the subfamily are Mellisugini (bees) and Trochilini (emeralds). The informal name "mountain gems" has ...
of subfamily
Trochilinae Trochilinae is one of the six subfamilies that make up the hummingbird family Trochilidae. The subfamily is divided into three tribes: Lampornithini (mountain gems) containing 18 species, Mellisugini (bees) containing 37 species and Trochili ...
. It is found in Costa Rica,
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the countr ...
, and
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Co ...
.HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved 27 May 2021


Taxonomy and systematics

The purple-throated mountaingem's taxonomy is somewhat unsettled. As late as 1999 various authors have treated it, the
white-throated mountaingem The white-throated mountaingem or white-throated mountain-gem (''Lampornis castaneoventris'') is a species of hummingbird in tribe Lampornithini of subfamily Trochilinae. It is endemic to Panama.HBW and BirdLife International (2020) ''Handbook ...
(''L. castaneoventris'') and
gray-tailed mountaingem The grey-tailed mountaingem (''Lampornis cinereicauda''), also variously spelled gray-tailed mountaingem, grey-tailed mountain-gem, or gray-tailed mountain-gem, is a species of hummingbird in tribe Lampornithini of subfamily Trochilinae. It is ...
(''L. cinereicauda'') as a single species. Others have kept the purple-throated separate but lumped the other two as a single species.Peters, J. (2020). Purple-throated Mountain-gem (''Lampornis calolaemus''), version 1.0. In Birds of the World (T. S. Schulenberg, Editor). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.ptmgem.01 retrieved 18 May 2022 These three species form a closely related group that evolved some 3.5 million years ago and has diversified since. The purple-throated mountaingem currently (2022) is its own species with three subspecies, the nominate ''L. c. calolaemus'', ''L. c. pectoralis'', and ''L. c. homogenes''.


Description

The purple-throated mountaingem is long and weighs . Both sexes of all subspecies have a medium-length straight black bill, a white to pale buff stripe behind the eye, and a fairly long tail. Adult males of the nominate subspecies ''L. c. calolaemus'' have a glittering emerald green to bluish green forehead and crown. The rest of their upperparts are metallic bronze-green that shades to bluish or grass green on the uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are s ...
. Much of the face is dusky bronze green and the
gorget A gorget , from the French ' meaning throat, was a band of linen wrapped around a woman's neck and head in the medieval period or the lower part of a simple chaperon hood. The term later described a steel or leather collar to protect the th ...
is metallic violet or purple. The breast is bright metallic green with duller flanks and belly that tend to bronzy green or gray. The undertail coverts are deep bronzy gray with paler gray margins. The tail is dull blue-black. Adult females of the nominate subspecies have bright metallic green upperparts that are somewhat bluish on the uppertail coverts and a bit bronzish elsewhere. The face is mostly blackish. The throat, breast, and belly are tawny yellow and the undertail coverts dull white to tawny buff. The central tail feathers and the upper half of the others are dull metallic green or bronze green. The lower half of the outer tail feathers are mostly black with pale gray tips. Immature males of the nominate subspecies are much like the adult female, but with a dusky chin and throat and brownish gray undertail coverts with dull white margins. Males of subspecies ''L. c. pectoralis'' have a deeper purple gorget than the nominate. The green of their neck and upperatail coverts is significantly darker than the nominates, and their belly and undertail coverts are also darker, with a smaller area of metallic green on the breast. Males of subspecies ''L. c. homogenes'' are similar to those of the nominate but have a darker gray breast and belly. Females' upperparts are more bluish green than the nominate's, their central tail feathers darker bronze-green, and their underparts darker and more reddish.


Distribution and habitat

Subspecies ''L. c. pectoralis'' of purple-throated mountaingem is found from far southwestern Nicaragua to the
Cordillera de Guanacaste The Cordillera de Guanacaste, also called Guanacaste Cordillera, are a volcanic mountain range in northern Costa Rica near the border with Nicaragua. The mountain range stretches 110 km from northwest to the southeast and contains mostly com ...
in northwestern Costa Rica. The nominate ''L. c. calolaemus'' is found in the Coreillera Central and northern part of the
Cordillera de Talamanca The Cordillera de Talamanca is a mountain range that lies in the southeast half of Costa Rica and the far west of Panama. Much of the range and the area around it is included in La Amistad International Park, which also is shared between the two ...
of northern and central Costa Rica. ''L. c. homogenes'' is found in western Panama from
Chiriquí Province Chiriquí () is a province of Panama located on the western coast; it is the second most developed province in the country, after the Panamá Province. Its capital is the city of David. It has a total area of 6,490.9 km², with a population ...
to
Coclé Province Coclé () is a province of central Panama on the nation's southern coast. The administrative capital is the city of Penonomé. This province was created by the Act of September 12, 1855 with the title of Department of Coclé during the presidenc ...
and probably also in southern Costa Rica. The species inhabits humid montane
evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, Live oak, acacias, magnolia, and banksia in more temperate zo ...
and
cloudforest A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest (TMCF), is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud ...
, where it favors steep slopes and broken terrain. In central Costa Rica it ranges in elevation from and is found as low as in northern Costa Rica.


Behavior


Movement

In some areas the purple-throated mountaingem moves to as low as after the breeding season.


Feeding

The purple-throated mountaingem feeds mostly on nectar. Males defend patches of flowers from other hummingbird species, from males of its own species, and also from females after courtship. Females are less territorial and often feed by
trap-lining In ethology and behavioral ecology, trap-lining or traplining is a feeding strategy in which an individual visits food sources on a regular, repeatable sequence, much as trappers check their lines of traps. Traplining is usually seen in species ...
. The species takes nectar from a wide variety of flowers. It is the primary pollinator of ''
Psychotria elata ''Palicourea elata'', formerly ''Psychotria elata'', commonly known as girlfriend kiss and labios de puta, is a tropical plant that ranges from Central to South American rain forests in countries such as Mexico, Costa Rica, Ecuador, Panama, and ...
'' and '' Palicourea lasiorrachis'' (
Rubiaceae The Rubiaceae are a family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with interpetiolar stipules a ...
). Females have slightly longer bills than males. There is some degree of
niche differentiation In ecology, niche differentiation (also known as niche segregation, niche separation and niche partitioning) refers to the process by which competing species use the environment differently in a way that helps them to coexist. The competitive excl ...
between the sexes. Though both prefer flowers with a corolla long by wide, females far more often than males utilize plants with longer and thinner corollas. Like most hummingbirds, the purple-throated mountaingem also feeds on insectes. Males capture them by
hawking Hawking may refer to: People * Stephen Hawking (1942–2018), English theoretical physicist and cosmologist * Hawking (surname), a family name (including a list of other persons with the name) Film * ''Hawking'' (2004 film), about Stephen Ha ...
from a perch. Females hover-glean, often beating their wings against foliage which apparently flushes prey.


Breeding

The purple-throated mountain-gem breeds between October and April, the rainy season. The female is entirely responsible for nest building, incubation, and nestling care. The nest is a thick-walled open cup of plant fibers with moss and lichen on the outside. It is placed in the understory, on bamboo or in a small tree, and typically about above the ground. The female incubates the clutch of two white eggs for 17 to 18 days and fledging occurs 22 to 23 days after hatch.


Vocalization

The purple-throated mountaingem's song is "high, thin, and dry, a complex medley of sputtering and warbling notes." A frequently heard call is variously described as "''trrrt''" or "a sharp, penetrating, buzzy ''zeet'' or ''zeep''." It also gives "higher-pitched, scratchy, chattering notes" when interacting with other hummingbirds.


Status

The IUCN has assessed the purple-throated mountaingem as being of Least Concern. Its population is estimated to be between 50,000 and 500,000 mature individuals but is thought to be decreasing. No specific threats have been identified. Though some of its habitat has been altered by humans, much remains untouched, and "studies attest to the strong resiliency of Purple-throated Mountain-gem populations in the face of habitat alteration."


Gallery

Purple-throated mountaingem (Lampornis calolaema) male composite.jpg, Composite showing the effect of light reflection on colors of male's gorget, Mount Totumas cloud forest,
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Co ...
Lampornis_calolaemus-7.jpg, Male at feeder


Notes


References


Further reading

* *


External links

* {{Taxonbar, from=Q838353 purple-throated mountaingem Birds of Nicaragua Birds of the Talamancan montane forests purple-throated mountaingem purple-throated mountaingem