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''Pungitius laevis'', commonly known as smoothtail ninespine stickleback, is a species of
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
of the family
Gasterosteidae Families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of socie ...
. It is distributed in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
benthopelagic The demersal zone is the part of the sea or ocean (or deep lake) consisting of the part of the water column near to (and significantly affected by) the seabed and the benthos. The demersal zone is just above the benthic zone and forms a layer of ...
waters of coastal western
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
.


Taxonomy

''Pungitius laevis'' was first formally described as ''Gasterosteus laevis'' in 1829 by the French
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuvier ...
with its type locality given as the basin of the
Seine ) , mouth_location = Le Havre/Honfleur , mouth_coordinates = , mouth_elevation = , progression = , river_system = Seine basin , basin_size = , tributaries_left = Yonne, Loing, Eure, Risle , tributarie ...
at
Bobigny Bobigny () is a commune, or town, in the northeastern suburbs of Paris, Île-de-France, France. It is located from the centre of Paris. Bobigny is the prefecture (capital city) of the Seine-Saint-Denis department, as well as the seat of the A ...
. This taxon has been considered to be
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organ ...
with the
ninespine stickleback The ninespine stickleback (''Pungitius pungitius''), also called the ten-spined stickleback, is a freshwater species of fish in the family Gasterosteidae that inhabits temperate waters. It is widely but locally distributed throughout Eurasia and ...
(''P. pungitius''), sometimes treated as a western European subspecies of that
holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical region ...
species. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''laevis'', means "smooth" and allusion to the lack of scutes on the
caudal peduncle Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
.


Description

''Pungitius laevis'' have a concave snout head shape,
truncate In mathematics and computer science, truncation is limiting the number of digits right of the decimal point. Truncation and floor function Truncation of positive real numbers can be done using the floor function. Given a number x \in \mathbb ...
caudal fin, and elongated body shape. It has a total of nine dorsal spines and 10-11 pectoral fin rays. Adults typically have a coloration of bright green with darker green tints which are covered by black blotches. This species has a maximum published
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
of .


Distribution and habitat

''Pungitius laevis'' is found in western Europe in Ireland, southern
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...
and from the Netherlands to the
Garonne The Garonne (, also , ; Occitan, Catalan, Basque, and es, Garona, ; la, Garumna or ) is a river of southwest France and northern Spain. It flows from the central Spanish Pyrenees to the Gironde estuary at the French port of Bordeaux – a ...
in France where it is found in densely vegetated shallow, still waters. It is a benthopelagic species which is found in fresh, brackish and marine waters.


Biology

Adult specimen prefer still, densely vegetated shallow water where they have access to
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
,
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
insects, and small crustaceans.


Reproduction

Breeding season lasts from April to June, with males being responsible for nesting and guarding the
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
before they hatch. Nests are usually built by vegetation, and eggs usually take around 10 to 20 days to hatch. Males die at the end of breeding season.


References

{{Taxonbar, from=Q5594727 Laevis Freshwater fish of the Arctic Freshwater fish of Europe Fish described in 1829 Taxa named by Georges Cuvier Holarctic fauna