The puku (''Kobus vardonii'') is a medium-sized

antelope The term antelope is used to refer to many species of even-toed ruminant that are indigenous to various regions in Africa and Eurasia. Antelope comprise a wastebasket taxon defined as any of numerous Old World grazing and browsing hoofed mammals ...

found in wet

grasslands A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natural ...

in southern

Democratic Republic of Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...

Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...

Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...

Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most cent ...

and more concentrated in the Okavango Delta in

Botswana Botswana (, ), officially the Republic of Botswana ( tn, Lefatshe la Botswana, label=Setswana, ), is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory being the Kalahar ...

. Nearly one-third of all puku are found in

protected area Protected areas or conservation areas are locations which receive protection because of their recognized natural, ecological or cultural values. There are several kinds of protected areas, which vary by level of protection depending on the ena ...

zoo A zoo (short for zoological garden; also called an animal park or menagerie) is a facility in which animals are kept within enclosures for public exhibition and often bred for Conservation biology, conservation purposes. The term ''zoological g ...

s, and

national park A national park is a nature park, natural park in use for conservation (ethic), conservation purposes, created and protected by national governments. Often it is a reserve of natural, semi-natural, or developed land that a sovereign state dec ...

s due to their diminishing

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

Description

Puku stand about at the shoulder and weigh from . The puku is sandy brown in colour, with the underbelly a slightly lighter brown. The coat is rougher than that of the similar-sized

southern reedbuck The southern reedbuck, rietbok or common reedbuck (''Redunca arundinum'') is a diurnal antelope typically found in southern Africa. It was first described by Pieter Boddaert, a Dutch physician and naturalist, in 1785. It is placed in the genus ...

lechwe The lechwe, red lechwe, or southern lechwe (''Kobus leche'') is an antelope found in wetlands of south-central Africa. Range The lechwe is native to Botswana, Zambia, southeastern Democratic Republic of the Congo, northeastern Namibia, and easte ...

impala The impala or rooibok (''Aepyceros melampus'') is a medium-sized antelope found in eastern and southern Africa. The only extant member of the genus '' Aepyceros'' and tribe Aepycerotini, it was first described to European audiences by Germa ...

, or the smaller

oribi The oribi (; ''Ourebia ourebi'') is a small antelope found in eastern, southern and western Africa. The sole member of its genus, it was described by the German zoologist Eberhard August Wilhelm von Zimmermann in 1783. While this is the only m ...

. Males have 50-cm-long, ridge-structured horns which are

lyre The lyre () is a stringed musical instrument that is classified by Hornbostel–Sachs as a member of the lute-family of instruments. In organology, a lyre is considered a yoke lute, since it is a lute in which the strings are attached to a yoke ...

-shaped.

Subspecies

There are two subspecies, the Senga Puku (''Kobus vardonii senganus'') and the southern puku (''Kobus vardonii vardonii'').

Ecology

Puku are found almost exclusively in

marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...

y grassland and

dambos A dambo is a class of complex shallow wetlands in central, southern and eastern Africa, particularly in Zambia, Malawi and Zimbabwe. They are generally found in higher rainfall flat plateau areas and have river-like branching forms which in themsel ...

, where they eat

grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...

es. The puku diet is flexible in regards to type of grasses consumed. There is little dietary competition with other bovids. This species is

crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of daylig ...

, active in the early morning and late afternoon. When scared, puku repeat a shrill whistle sound. Females gather in

herd A herd is a social group of certain animals of the same species, either wild or domestic. The form of collective animal behavior associated with this is called ''herding''. These animals are known as gregarious animals. The term ''herd'' is ...

s of up to 20 individuals. During the rainy season, herds will come together for added safety, typically reaching around 50 females. Males hold

territories A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or an ...

and attempt to persuade herds of females to stay within their territories for as long as possible. In the wet season, due to large floods in their habitat they migrate to a higher elevation and in the dry season remain near water. File:Puku male and female (Zambia).jpg, Male and female ''K. v. senganus'',

Kafue National Park Kafue National Park is the largest national park in Zambia, covering an area of about 22,400 km² (similar in size to Wales or Massachusetts). It is one of the largest parks in Africa and is home to 152 different species of mammals. The park ...

File:Puku female, South Luangwa.jpg, Female ''K. v. senganus'' and fawn, in South Luangwa NP,

File:Southern puku (Kobus vardonii vardonii) female.jpg, Female ''K. v. vardonii'', in

Chobe National Park Chobe National Park is Botswana's first national park, and also the most biologically diverse. Located in the north of the country, it is Botswana's third largest park, after Central Kalahari Game Reserve and Gemsbok National Park, and has one of ...

File:Southern puku (Kobus vardonii vardonii) female head.jpg, Female ''K. v. vardonii'', in

File:Southern puku (Kobus vardonii vardonii) fawn.jpg, ''K. v. vardonii'' fawn, in

References

External links

1881 drawing of puku
{{Taxonbar, from=Q273577 Marsh antelopes Mammals of the Democratic Republic of the Congo Mammals of Zambia Mammals of Tanzania Mammals described in 1857