The Puerto Rican emerald (''Riccordia maugaeus''), or zumbadorcito de Puerto Rico in Spanish, is

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics aro ...

in the "emeralds", tribe Trochilini of subfamily Trochilinae. It is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

Puerto Rico Puerto Rico (; abbreviated PR; tnq, Boriken, ''Borinquen''), officially the Commonwealth of Puerto Rico ( es, link=yes, Estado Libre Asociado de Puerto Rico, lit=Free Associated State of Puerto Rico), is a Caribbean island and Unincorporated ...

.Bündgen, R. and P. F. D. Boesman (2021). Puerto Rican Emerald (''Riccordia maugaeus''), version 1.1. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.pureme1.01.1 retrieved July 29, 2022

Taxonomy and systematics

The Puerto Rican emerald was formerly placed in the genus ''

Chlorostilbon ''Chlorostilbon'' is a genus of hummingbird in the family Trochilidae, known as emeralds (as are some hummingbirds in the genera ''Amazilia'' and '' Elvira''). A single species, the blue-chinned sapphire is variously placed in the monotypic gen ...

''. Based on a

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

study published in 2014 and a 2017 publication, the North American Classification Committee of the

American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its m ...

, the

International Ornithological Committee The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...

(IOC), and the

Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 2022 ...

moved it to the resurrected genus ''

Riccordia ''Riccordia'' is a genus of birds in the hummingbird family Trochilidae. They are endemic to the Caribbean. Species The species now placed in this genus were formerly assigned to the genus ''Chlorostilbon''. A molecular phylogenetic study publish ...

''.Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 25, 2021 However, as of 2020

BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...

Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. T ...

(HBW) retained it in ''Chlorostilbon''.

Description

Male Puerto Rican emeralds are long and females . The species weighs between . Both sexes have a small white spot behind the eye. Males have a short straight bill with black-tipped red

mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...

and a black

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...

. Its forehead and crown are iridescent green; the rest of its upperparts are dark shining metallic green. Its

gorget A gorget , from the French ' meaning throat, was a band of linen wrapped around a woman's neck and head in the medieval period or the lower part of a simple chaperon hood. The term later described a steel or leather collar to protect the thro ...

is iridescent bluish green and the rest of its underparts are shining green. The tail is forked and shining steely blue. The female's bill is all black. Its forehead and crown are dull dark green and the rest of the upperparts shining grass green. Its throat is light gray that darkens on the belly and undertail coverts. The tail is less forked than the male's. Its outermost pair of feathers have pale white bases, brown centers, and white ends. The next pair inward is steel blue with greenish white bases and a white spot at the end. The next two pairs are green on their body half and dull brown beyond. The central pair is shining green.

Distribution and habitat

The Puerto Rican emerald is found throughout its namesake island. It inhabits almost all the forested landscapes from coastal mangroves to montane forest on the summits including dry and moist forests, plantations, and

secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has re-grown after a timber harvest or clearing for agriculture, until a long enough period has passed so that the effects of the disturbance are no longer evident. ...

. It also occurs in gardens and urban areas.

Behavior

Movement

The Puerto Rican emerald is sedentary.

Feeding

The Puerto Rican emerald forages for nectar from wide variety flowering plants and trees. It mostly feeds at low to medium levels of the forest. In addition to nectar, it feeds on small

arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...

s by gleaning from leaves and branches; its diet includes flies,

Homoptera Homoptera is a suborder of order Hemiptera that is considered by some taxonomists to be paraphyly, paraphyletic, and therefore Deprecation, deprecated (obsolete). It was therefore split into the suborders Sternorrhyncha, Auchenorrhyncha, and Cole ...

Hymenoptera Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Par ...

, and spiders.

Breeding

The Puerto Rican emerald's peak breeding season spans from February to May but some nesting occurs at any time of year. The nest is a small cup of dry plant fibers lined with softer material and covered with lichen. It is typically built in a low to medium-height shrub or tree. The female incubates the clutch of two eggs for 14 to 16 days; fledging occurs 20 to 22 days after hatch.

Vocalization

What is believed to be the Puerto Rican emerald's song is "a repeated twittering phrase of high-pitched descending notes followed by a series of ‘lisping’ notes on constant pitch 'tseereetseetseetsee-tslew-tslew-tslew-tslew-tslew'". It also makes "high-pitched 'tsik'" and "si..si..sik-sik...tsik.." calls.

Status

The

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

has assessed the Puerto Rican emerald as being of Least Concern, though its population size and trend are not known. It is considered generally common throughout its large island and accepts human-made landscapes like plantations, gardens, and parks.

References

External links

Images of Puerto Rican emerald