The Puerto Rican crow (''Corvus pumilis'') is an extinct

crow A crow is a bird of the genus ''Corvus'', or more broadly a synonym for all of ''Corvus''. Crows are generally black in colour. The word "crow" is used as part of the common name of many species. The related term "raven" is not pinned scientifical ...

species in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Corvidae Corvidae is a cosmopolitan family of oscine passerine birds that contains the crows, ravens, rooks, jackdaws, jays, magpies, treepies, choughs, and nutcrackers. In colloquial English, they are known as the crow family or corvids. Currently, 13 ...

that was

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

Puerto Rico Puerto Rico (; abbreviated PR; tnq, Boriken, ''Borinquen''), officially the Commonwealth of Puerto Rico ( es, link=yes, Estado Libre Asociado de Puerto Rico, lit=Free Associated State of Puerto Rico), is a Caribbean island and Unincorporated ...

and the

United States Virgin Islands The United States Virgin Islands,. Also called the ''American Virgin Islands'' and the ''U.S. Virgin Islands''. officially the Virgin Islands of the United States, are a group of Caribbean islands and an unincorporated and organized territory ...

. Little is known about its habitat, but it possibly died out after the colonization of humans on these islands.

Description

The holotype specimen is a

subfossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...

ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...

. It was long, and lies in size between the formerly

sympatric In biology, two related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter one another. An initially interbreeding population that splits into two or more distinct species sh ...

'' C. leucognaphalus'' with and the

Hispaniola Hispaniola (, also ; es, La Española; Latin and french: Hispaniola; ht, Ispayola; tnq, Ayiti or Quisqueya) is an island in the Caribbean that is part of the Greater Antilles. Hispaniola is the most populous island in the West Indies, and th ...

n '' C. palmarum'' with .

Habitat

Residues of the crow were found on

, and on the island of

St. Croix Saint Croix; nl, Sint-Kruis; french: link=no, Sainte-Croix; Danish and no, Sankt Croix, Taino: ''Ay Ay'' ( ) is an island in the Caribbean Sea, and a county and constituent district of the United States Virgin Islands (USVI), an unincor ...

, which belongs to the

. Little is known about its habitat. As it existed together with ''C. leucognaphalus'' on Puerto Rico, it possibly occupied a different

ecological niche In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of resources and competitors (for ...

as the latter, and was perhaps rather common in the island's lowlands.

Classification and taxonomy

The earliest residues of the crow were found in 1916, in the

karst cave Karst is a topography formed from the dissolution of soluble rocks such as limestone, dolomite, and gypsum. It is characterized by underground drainage systems with sinkholes and caves. It has also been documented for more weathering-resistant ro ...

Cueva San Miguel near

Morovis, Puerto Rico Morovis (, ) is a town and municipality of Puerto Rico located in the central region of the island, north of Orocovis, south of Manatí, Vega Baja and Vega Alta; east of Ciales, and west of Corozal. Morovis is spread over 13 barrios and Morov ...

. It was a right ulna (

AMNH The American Museum of Natural History (abbreviated as AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. In Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 26 i ...

4925), which

Alexander Wetmore Frank Alexander Wetmore (June 18, 1886 – December 7, 1978) was an American ornithologist and avian paleontologist. He was the sixth Secretary of the Smithsonian Institution. Early life and education The son of a Country Physician, Frank Al ...

described in 1920 as a

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

for his first description of the species ''C. pumilis''. Wetmore did not comment on the

etymology Etymology ()The New Oxford Dictionary of English (1998) – p. 633 "Etymology /ˌɛtɪˈmɒlədʒi/ the study of the class in words and the way their meanings have changed throughout time". is the study of the history of the Phonological chan ...

of the

epithet An epithet (, ), also byname, is a descriptive term (word or phrase) known for accompanying or occurring in place of a name and having entered common usage. It has various shades of meaning when applied to seemingly real or fictitious people, di ...

''pumilis'', which means "dwarfish" in

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

. There are no insights on its relationships with other species of its genus within and beyond the

Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...

Extinction

''C. pumilis'' possibly became extinct before the colonization of the islands. In Puerto Rico, it is only known from ''

lagerstätte A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These for ...

n''; on St. Croix, it was found on a hearth from the

Pre-Columbian era In the history of the Americas, the pre-Columbian era spans from the Migration to the New World, original settlement of North and South America in the Upper Paleolithic period through European colonization of the Americas, European colonization, w ...

References

;Further reading * Pierce Brodkorb: ''Catalogue of Fossil Birds.'' In: ''Bulletin of the Florida State Museum. Biological sciences.'' 23 (3), 1978. pages 139–157.
full text
* Julian Pender Hume, Michael Walters: ''Extinct Birds.'' A & C Black, London 2012. . * Alexander Wetmore: ''Five New Species of Birds from Cave Deposits in Porto Rico.'' In: ''Proceedings of the Biological Society of Washington'' 33, 1920. pages 77–81.
full text
* Alexander Wetmore: ''Bird Remains from Cave Deposits on Great Exuma Island in the Bahamas.'' In: ''Bulletin of the Museum of Comparative Zoölogy at Harvard College'' 80, 1937. pages 427–441. {{Taxonbar, from=Q1548854 Corvus Extinct birds of the Caribbean Holocene extinctions Taxa named by Alexander Wetmore