Ptychopariida is a large, heterogeneous order of

trilobite Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...

containing some of the most primitive species known. The earliest species occurred in the second half of the Lower

Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...

, and the last species did not survive the

Ordovician–Silurian extinction event The Late Ordovician mass extinction (LOME), sometimes known as the end-Ordovician mass extinction or the Ordovician-Silurian extinction, is the first of the "big five" major mass extinction events in Earth's history, occurring roughly 443 Mya. It ...

. Trilobites have facial sutures that run along the margin of the

glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ...

and/or fixigena to the shoulder point where the

cephalon Cephalon, Inc. was an American biopharmaceutical company co-founded in 1987 by pharmacologist Frank Baldino, Jr., neuroscientist Michael Lewis, and organic chemist James C. Kauer—all three former scientists with the DuPont Company. Baldino s ...

meets the

thorax The thorax or chest is a part of the anatomy of humans, mammals, and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main divisions of the cre ...

. These sutures outline the cranidium, or the main, central part of the head that does not include the librigena (free cheeks). The eyes are medial along the glabella on the suture line (and some species have no eyes). The

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s of the moults of trilobites can often be told from the fossils of the actual animals by whether the librigena are present. (The librigena, or cheek spines, detach during moulting.) In ptychopariids, short bladelike genal spines are often present on the tips of the librigena. The thorax is large and is typically made up of eight or more segments. The thorax is usually much longer than the

pygidium The pygidium (plural pygidia) is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is compo ...

, which is usually small. In some species the pygidium is outlined with a flat border. The Subclass Librostoma was recently erected to encompass several related orders, including Ptychopariida,

Asaphida Asaphida is a large, morphologically diverse order of trilobites found in marine strata dated from the Middle Cambrian until their extinction during the Silurian. Asaphida contains six superfamilies (Anomocaroidea, Asaphoidea, Cyclopygoidea, ...

Proetida Proetida is an order of trilobite that lived from the Ordovician to the Permian. It was the last order of trilobite to go extinct, finally dying out in the Permian-Triassic extinction event. Description These typically small trilobites resem ...

Harpetida Harpetida is one of the eleven orders of the extinct arthropod class Trilobita. The first harpetid trilobites appear in the Upper Cambrian, and the last species die out at the end of the Devonian period. Harpetid trilobites are characterize ...

, and possibly

Phacopida Phacopida ("lens-face") is an order of trilobites that lived from the Late Cambrian to the Late Devonian. It is made up of a morphologically diverse assemblage of taxa in three related suborders. Characteristics Phacopida had 8 to 19 thorac ...

. These are now known as the "Librostome Orders". Trilobites of the orders Proetida, Harpetida, and of the family

Damesellidae Damesellidae is a family of odontopleurid trilobites found in late Middle to Late Cambrian marine strata, primarily of China. Damesellids are closely related to the odontopleurids of Odontopleuridae, but are not nearly as spinose, nor possess ...

were originally placed in Ptychopariida.

Taxonomy

Suborder

Olenina ''Olenina'' is an extinct suborder of the trilobite order Ptychopariida. Subdivisions *Superfamily Olenoidea **Family Ellipsocephaloididae **Family Olenidae Olenidae is a family of ptychopariid trilobites. Some genera, '' Balnibarbi'' and ...

*Superfamily Olenoidea **Family Ellipsocephaloididae **Family

Olenidae Olenidae is a family of ptychopariid trilobites. Some genera, ''Balnibarbi'' and ''Cloacaspis'', are thought to have evolved a symbiotic relationship with sulfur-eating bacteria from which they derived nutrition. Genera * Acerocare * Aceroca ...

*Superfamily ''Incertae sedis'' **Genus ''

Triarthrus ''Triarthrus'' is a genus of Upper Ordovician ptychopariid trilobite found in New York, Ohio, Kentucky, and Indiana, eastern and northern Canada, China and Scandinavia. It is the last of the Olenid trilobites, a group which flourished in the Camb ...

'' Suborder

Ptychopariina Ptychopariina is an extinct suborder of trilobites of the order Ptychopariida.Swinnerton HH, 1915. Suggestions for a Revised Classification of trilobites. ''Geological Magazine''. Also known as the ''primitive Ptychopariida'', they are a notably ...

*Superfamily Ellipsocephaloidea **Family Agraulidae **Family Aldonaiidae **Family Bigotinidae **Family Chengkouiidae **Family Ellipsocephalidae **Family Estaingiidae **Family Palaeolenidae **Family

Yunnanocephalidae ''Yunnanocephalus'' is a genus of ptychopariid trilobite. It lived during the late Atdabanian and Botomian stages, in what are currently Antarctica, Australia and China. It was a "moderately common" member of the Chengjiang Fauna. ''Yunnanocepha ...

*Superfamily

Ptychoparioidea Ptychoparioidea is a superfamily of the Ptychopariida order of trilobites.Pour M. G., Popov L. E. (2009) Silicified Middle Cambrian trilobites from Kyrgyzstan, Palaeontology, 1039-1056 Taxonomy *Family Acrocephalitidae *Family Alokistocaridae ...

**Family Acrocephalitidae **Family

Alokistocaridae Alokistocaridae is a family of ptychopariid trilobites that lived from the Botomian epoch of the Early Cambrian until the Late Cambrian. Alokistocarids were particle feeders and left small furrows which are occasionally preserved.Coppold, Murray ...

**Family Antagmidae **Family Asaphiscidae **Family Atopidae **Family Bolaspididae **Family Cedariidae **Family Changshaniidae **Family Conocoryphidae **Family Conokephalinidae **Family Crepicephalidae **Family

Diceratocephalidae Diceratocephalidae is an family of trilobites belonging to the order Ptychopariida Ptychopariida is a large, heterogeneous order of trilobite containing some of the most primitive species known. The earliest species occurred in the second half ...

**Family Elviniidae **Family Eulomidae **Family Holocephalinidae **Family Ignotogregatidae **Family Inouyiidae **Family Isocolidae **Family Kingstoniidae **Family Liostracinidae **Family Llanoaspididae **Family Lonchocephalidae **Family Lorenzellidae **Family Mapaniidae **Family Marjumiidae **Family Menomoniidae **Family

Nepeidae The Nepeidae are a family of trilobites, that lived during the late Middle Cambrian - earliest Upper Cambrian in what are today Australia, Antarctica, China and New Zealand. The Nepeidae can be recognized by a vaulted area between the front of th ...

**Family Norwoodiidae **Family Papyriaspididae **Family Phylacteridae **Family Proasaphiscidae **Family

Ptychopariidae Ptychopariidae is a family of trilobites, containing the following genera: *'' Achlysopsis'' *'' Altikolia'' *'' Altitudella'' *'' Amecephalites'' *'' Asthenopsis'' *'' Austinvillia'' *'' Balangcunaspis'' *''Bashania'' *'' Bathyholcus'' *'' Bathy ...

**Family Shumardiidae **Family

Solenopleuridae Solenopleuridae is a family (biology), family of trilobites, containing the following genera: *''Abakanopleura'' *''Acanthometopus'' *''Acrocephalaspis'' *''Aiaiaspis'' *''Aidarella'' *''Aikhaliella'' *''Albansia'' *''Aldanaspis'' *''Badulesia'' ...

**Family Tricrepicephalidae **Family Utiidae **Family Wuaniidae *Superfamily ''Incertae sedis'' **Genus '' Tonopahella'' Suborder ''incertae sedis'' *Family Avoninidae *Family Catillicephalidae *Family Ityophoridae *Family Plethopeltidae

References

Order Ptychopariida

By Sam Gon III Trilobite orders Cambrian first appearances Late Ordovician extinctions {{Ptychopariida-stub

Taxonomy

See also

References