Ptychococcus
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''Ptychococcus'' is a
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is conne ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
in the
palm Palm most commonly refers to: * Palm of the hand, the central region of the front of the hand * Palm plants, of family Arecaceae **List of Arecaceae genera * Several other plants known as "palm" Palm or Palms may also refer to: Music * Palm (ba ...
family from
New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...
and the
Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capita ...
. They are closely related to ''
Ptychosperma ''Ptychosperma'' is a genus of flowering plant in the family Arecaceae. Most are native to Australia and/or New Guinea, with a few in the Solomon Islands and in Maluku Province of eastern Indonesia. Some have been cultivated abroad as house or ...
'', only differentiated by the seed shape and endocarp type.Uhl, Natalie W. and Dransfield, John (1987) ''Genera Palmarum - A classification of palms based on the work of Harold E. Moore''. Lawrence, Kansas: Allen Press. / The name is a combination of the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
for "fold" and the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for "berry".


Description

The trunks grow to 15 m, usually no wider than 25 cm, and both are solitary, ringed, and
crownshaft An elongated circumferential leaf base formation present on some species of palm is called a crownshaft. The leaf bases of some pinnate leaved palms (most notable being ''Roystonea regia'' or the royal palm but also including the genera ''Areca' ...
ed. The
leaf A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...
is pinnately compounded, in long sheaths, usually covered in scales and hairs, as is the short petiole. The ridged
rachis In biology, a rachis (from the grc, ῥάχις [], "backbone, spine") is a main axis or "shaft". In zoology and microbiology In vertebrates, ''rachis'' can refer to the series of articulated vertebrae, which encase the spinal cord. In this c ...
is flattened on the bottom and also covered in hairy
tomentum Tomentum may refer to: * Plant trichomes, a covering of closely matted or fine hairs on plant leaves. * Tomentum (anatomy), short, soft pubescence or a covering of fine, soft hairs. {{disambig ...
. The unusual leaflets are once-folded and toothed, twisting upwards in their bottom half. Each leaflet also bears scales and a prominent
midrib This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary o ...
, with tomentose margins, and lacking visible veinlets. The
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
emerges below the
crownshaft An elongated circumferential leaf base formation present on some species of palm is called a crownshaft. The leaf bases of some pinnate leaved palms (most notable being ''Roystonea regia'' or the royal palm but also including the genera ''Areca' ...
, stiff and horizontal, and branched to three orders. Covered in scales, the peduncle is short and thick, the prophyll is tubular, beaked and tomentose, and the long rachis bears numerous short rachillae which are often Z-shaped. The rachillae bear short round bracts subtending triads of large
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s throughout. The
staminate The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
flowers are more or less asymmetrical and bear three distinct, hairy
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
s, and three ovate, scaly
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s. There may be as many as 100 stamens, with short filaments, and elongated, apically notched, deeply
bifid Bifid refers to something that is split or cleft into two parts. It may refer to: * Bifid, a variation in the P wave, R wave, or T wave in an echocardiogram in which a wave which usually has a single peak instead has two separate peaks * Bifid ci ...
anther The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
s. The
exine Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
is finely reticulate and tectate. The pistilode is bottle-shaped and has pointed tips. The pistillate flowers are smaller, ovoid, and occasionally hairy; both sepals and petals are imbricate, the latter bearing scales. There are three united staminodes forming a small cup, the
gynoecium Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
is ovoid and uniovulate; the pendulous stigma has three lobes. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
is egg-shaped with a wrinkly exterior, divided into lobed segments when dry, and mature at orange or red. The
epicarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggr ...
is fibrous, the
mesocarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggre ...
fleshy, covering a five-lobed
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
, resembling the dry fruit.


Distribution and habitat

In New Guinea and the Solomon Islands they are found in a variety of settings and at various elevations. Growing in rocky or mountainous
rain forest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainforest ...
, in rain forest low lands, and alongside rivers, they are not particular to soil type. ''P. lepidotus'' grows high, to 3000 m, not preferring truly tropical conditions.


Cultivation and uses

Only ''P. paradoxus'' is cultivated with any regularity, though it is relatively uncommon. They are tender to cold, need quickly draining soil and generous amounts of water. In the New Guinea highlands, the trunks of ''P. lepidotus'' are used in construction or cut into 2 m strips and carved into bows; smaller pieces are fashioned into arrows and arrow heads.PACSOA: ''Ptychococcus lepidotus''
/ref>


References


External links


''Ptychococcus'' on NPGS/GRIN

GBIF portal

Fairchild Guide to Palms: ''Ptychococcus''

Fairchild images


{{Taxonbar, from=Q143646 Ptychospermatinae Arecaceae genera Taxa named by Odoardo Beccari