Ptyas Mucosus
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''Ptyas mucosa'', commonly known as the oriental ratsnake, Indian rat snake, ''darash'' or ''dhaman'', is a common non-venomous
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
colubrid Colubridae (, commonly known as colubrids , from la, coluber, 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest species of the family date back to the Oligocene epoch. Colubrid snakes are found on ever ...
snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...
found in parts of
South South is one of the cardinal directions or Points of the compass, compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Pro ...
and
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
. Dhamans are large snakes. Typical mature total length is around though some exceed . The record length for this species was , second only to their cousin ''
Ptyas carinata ''Ptyas carinata'', commonly known as the keeled rat snake, is a species of colubrid snake. It is found in Indonesia, Myanmar, Malaysia, Thailand, Philippines, Cambodia,India, Vietnam and Singapore. This little known species is probably the larg ...
'' among living colubrid snakes.Auliya, M. (2010). ''Conservation Status and Impact of Trade on the Oriental Rat Snake Ptyas mucosa in Java, Indonesia''. TRAFFIC Southeast Asia. Despite their large size, oriental ratsnakes are usually quite slender with even a specimen of commonly measuring only around in diameter. Furthermore, the average weight of ratsnakes caught in
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
was around , though larger males of over (which average mildly larger of the two sexes in the species) may easily weigh over . Their color varies from pale browns in dry regions to nearly black in moist forest areas. Rat snakes are diurnal, semi-
arboreal Arboreal locomotion is the Animal locomotion, locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. Th ...
, non-venomous, and fast-moving. Rat snakes eat a variety of prey and are frequently found in urban areas where rodents thrive.


Geographic range

Found in
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is bordere ...
,
Andaman Andaman may refer to: * Andaman Islands * Andaman Sea * ''Andaman'' (1998 film), a Kannada-language film * ''Andaman'' (2016 film), a Tamil-language film * ''Andaman'' (2021 film), a Hindi-language film See also * Andaman and Nicobar Islands ...
and Nicobar Islands,
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
,
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand t ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
(Zhejiang, Hubei, Jiangxi, Fujian, Guangdong, Hainan, Guangxi, Yunnan, Tibet, Hong Kong),
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
,
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
(Sumatra, Java, Bali),
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
, West
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
,
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...
,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
,
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
(Sindh area),
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
,
Turkmenistan Turkmenistan ( or ; tk, Türkmenistan / Түркменистан, ) is a country located in Central Asia, bordered by Kazakhstan to the northwest, Uzbekistan to the north, east and northeast, Afghanistan to the southeast, Iran to the sout ...
and
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
.


Predators

Adult rat snakes have no natural predators, although younger specimens are the natural prey of King cobras that overlap them in their range. Juveniles fear
birds of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators ...
, larger reptiles, and mid-sized
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s. They are wary, quick to react, and fast-moving. Rat snakes and related colubrids are aggressively hunted by humans in some areas of their range for skins and meat. Harvesting and trade regulations exist in
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
and
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
, but these regulations are often ignored.


Description

Description from Boulenger's ''Fauna of British India: Reptilia and Batrachia'' volume of 1890:
Snout obtuse, slightly projecting; eye large; rostral a little broader than deep, visible from above; suture between the internasals shorter than that between the prefrontals; frontal as long as its distance from the end of the snout, as long as the parietals or slightly shorter; usually three loreals; one large preocular, with a small subocular below; two postoculars; temporals 2+2; 8 Upper labials, fourth and fifth entering the eye; 5 Lower labials in contact with the anterior chin shields, which are shorter than the posterior; the latter in contact anteriorly.
Dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
in 17 rows at midbody, more or less strongly keeled on the posterior part of the body. Ventrals 190–208; anal divided; subcaudals 95–135, divided. Brown above, frequently with more or less distinct black crossbands on the posterior part of the body and on the tail; young usually with light crossbands on the front half of the body. Lower surface yellowish; the posterior ventral and the caudal shields may be edged with black.
It is the second largest snake in Sri Lanka, after the
Indian rock python The Indian python (''Python molurus'') is a large python species native to tropical and subtropical regions of the Indian subcontinent and Southeast Asia. It is also known by the common names black-tailed python, Indian rock python, and Asian ...
.


Behavior

Rat snakes, though harmless to humans, are fast-moving, excitable snakes. In captivity, they are territorial and may defend their turf aggressively, attempting to startle or strike at passing objects. Rat snakes are diurnal and semi-arboreal. They inhabit forest floors, wetlands, rice paddies, farmland, and suburban areas where they prey upon small reptiles, amphibians, birds, and mammals. Adults, unusually for a colubrid, prefer to subdue their prey by sitting on it rather than by constricting, using body weight to weaken prey. Rat snakes mate in late spring and early summer, though in tropical areas reproduction may take place year round. Males establish boundaries of territory using a ritualised test of strength in which they intertwine their bodies. The behaviour is sometime misread by observers as a "mating dance" between opposite-sex individuals. Females produce 6–15 eggs per clutch several weeks after mating. Adult members of this species emit a growling sound and inflate their necks when threatened. This adaptation may represent mimicry of the king cobra or
Indian cobra The Indian cobra (''Naja naja''), also known as the spectacled cobra, Asian cobra, or binocellate cobra, is a species of cobra found in India, Pakistan, Bangladesh, Sri Lanka, Nepal, and Bhutan, and a member of the "big four" species that are ...
which overlaps this species in range. The resemblance often backfires in human settlements, though, as the harmless animal may be mistaken for a venomous snake and killed.


Nomenclature

The International Code for Zoological Nomenclature (ICZN) directs that the
grammatical gender In linguistics, grammatical gender system is a specific form of noun class system, where nouns are assigned with gender categories that are often not related to their real-world qualities. In languages with grammatical gender, most or all nouns ...
of any given
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
name should follow logically from the gender of its associated
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
name. As ''Ptyas'' is a feminine word form (from πτυάς, a Greek word for a venom-spitting snake), the proper form of the species name is ''mucosa'' (a
Late Latin Late Latin ( la, Latinitas serior) is the scholarly name for the form of Literary Latin of late antiquity.Roberts (1996), p. 537. English dictionary definitions of Late Latin date this period from the , and continuing into the 7th century in t ...
word meaning "slimy"). Reference materials older than 2004 often show the masculine form, ''mucosus,'' and the
CITES CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
list continues to list the species this way.


Gallery

Image:AB084_Ptyas_mucosos.JPG, Head with open mouth Image:AB085_Ptyas_mucosos.JPG, Top view of head Image:AB086_Ptyas_mucosos.JPG, View of temporals Image:AB087_Ptyas_mucosos.JPG, Underside of head Image:AB088_Ptyas_mucosos.JPG, Belly of snake Image:AB089_Ptyas_mucosos.JPG, Body and scales Image:AB090_Ptyas_mucosos.JPG, Body and scales Image:AB091_Ptyas_mucosos.JPG, Body and scales Image:AB092_Ptyas_mucosos.JPG, Long tail of oriental rat snake Image:AB093 Ptyas mucosos Temporals.JPG, Temporals indicated


References


Further reading

* David, P., and I. Das. 2004. On the grammar of the gender of ''Ptyas'' Fitzinger, 1843 (Serpentes: Colubridae). Hamaddryad 28 (1 & 2): 113–116. * Günther, A. 1898. Notes on Indian Snakes in Captivity. Ann. Mag. Nat. Hist., Series 7, 1: 30–31. (''Zamenis mucosus'', p. 30.) * Jan, G., & F. Sordelli. 1867. ''Iconographie générale des Ophidiens: Vingt-quatrième livraison''. Baillière. Paris. Index + Plates I.–VI. ("''Coryphodon Blumenbachi'', Merr.", Plate III., Figures 2–4.) * Lazell, J.D. 1998. Morphology and the status of the snake genus ''Ptyas''. Herpetological Review 29 (3): 134. * Linnaeus, C. 1858. ''Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, diferentiis, synonymis, locis. Tomus I. Editio Decima, Reformata.'' L. Salvius. Stockholm. 824 pp. (''Coluber mucosus'', p. 226.) * Morris, P.A. 1948. ''Boy's Book of Snakes: How to Recognize and Understand Them''. A volume of the Humanizing Science Series, edited by
Jacques Cattell Jaques (Jack) Cattell (2 June 1904 in Garrison, New York – 19 December 1961) was an American publisher and founder of a company bearing his name, "Jaques Cattell Press, Inc.," based in Lancaster, Pennsylvania. Jaques Cattell Press, Inc. The Sc ...
. Ronald Press. New York. viii + 185 pp. ("The Indian Rat Snake", pp. 136–137, 181.) * Nixon, A.M.A., and S. Bhupathy. 2001. Notes on the occurrence of Dhaman (''Ptyas mucosus'') in the higher altitudes of Nilgiris, Western Ghats. Cobra (44): 30–31.


External links

* {{Taxonbar, from=Q2338867 Colubrids Reptiles described in 1758 Reptiles of Afghanistan Reptiles of Bangladesh Reptiles of Cambodia Reptiles of Central Asia Reptiles of China Reptiles of India Reptiles of Indonesia Reptiles of Iran Reptiles of Laos Reptiles of Myanmar Reptiles of Nepal Reptiles of Pakistan Reptiles of Sri Lanka Reptiles of Taiwan Reptiles of Thailand Reptiles of Vietnam Snakes of Asia Taxa named by Carl Linnaeus ceb:Ptyas korros sv:Ptyas korros