''Pterostylis hamiltonii'', commonly known as the red-veined shell orchid, is a species of

orchid Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowering ...

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to the

south-west The points of the compass are a set of horizontal, radially arrayed compass directions (or azimuths) used in navigation and cartography. A compass rose is primarily composed of four cardinal directions—north, east, south, and west—each se ...

Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...

. As with similar orchids, the flowering plants differ from those which are not flowering. The non-flowering plants have a rosette of leaves but the flowering plants lack a rosette and have a single flower with leaves on the flowering spike. This greenhood has a green and white, striped flower with reddish-brown markings and forms colonies, sometimes of thousands of plants.

Description

''Pterostylis hamiltonii'' is a terrestrial,

perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...

herb In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal ...

with an underground

tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing ...

and when not flowering, a rosette of bluish-green leaves, each leaf 5–14 mm long and 5–12 mm wide. Flowering plants usually have a single flower 25–30 mm long and 12–15 mm wide borne on a spike 50–150 mm high with four to six stem leaves 20–40 mm long and 2–8 mm wide. The flowers are white with green or reddish-brown stripes and markings. The dorsal sepal and

petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...

s are fused, forming a hood or "galea" over the

column A column or pillar in architecture and structural engineering is a structural element that transmits, through compression, the weight of the structure above to other structural elements below. In other words, a column is a compression member. ...

. The dorsal sepal curves forward with a thread-like tip 1–3 mm long. The

lateral Lateral is a geometric term of location which may refer to: Healthcare *Lateral (anatomy), an anatomical direction *Lateral cricoarytenoid muscle *Lateral release (surgery), a surgical procedure on the side of a kneecap Phonetics *Lateral cons ...

sepals are erect and held closely against the galea. They have thread-like ends 20-25 long and between their bases there is a broad, flat

sinus Sinus may refer to: Anatomy * Sinus (anatomy), a sac or cavity in any organ or tissue ** Paranasal sinuses, air cavities in the cranial bones, especially those near the nose, including: *** Maxillary sinus, is the largest of the paranasal sinuses, ...

with a central notch. The labellum is 15–20 mm long, about 3 mm wide, dark red and curved and protrudes above the sinus. Flowering occurs from late May to early August.

Taxonomy and naming

''Pterostylis hamiltonii'' was first formally described in 1810 by

William Henry Nicholls William Henry Nicholls (23 July 1885 – 10 March 1951) was an Australian amateur botanist, authority on, and collector of Australian orchids. An accomplished photographer and watercolourist, he contributed almost 100 articles on orchids to ''The ...

and the description was published in ''

The Victorian Naturalist ''The Victorian Naturalist'' is a bimonthly scientific journal covering natural history, especially of Australia. It is published by the Field Naturalists Club of Victoria and is received as part of the membership subscription of that club. From ...

'' from a specimen collected near Boyup Brook. The

specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

(''hamiltonii'') honours Alex. G. Hamilton, (1852-1941) "a veteran in the field of botanical research".

Distribution and habitat

The red-veined shell orchid is found between Toodyay and the

Stirling Range The Stirling Range or Koikyennuruff is a range of mountains and hills in the Great Southern region of Western Australia, south-east of Perth. It is over wide from west to east, stretching from the highway between Mount Barker and Cranb ...

where it usually grows in thickets of ''

Allocasuarina huegeliana ''Allocasuarina huegeliana'', commonly known as rock sheoak or sighing sheoak, is a tree in the family Casuarinaceae. Endemic to Western Australia, it occurs mostly throughout the Wheatbelt region. It is now especially common on road verges, w ...

'' on or near granite outcrops in the

Avon Wheatbelt The Avon Wheatbelt is a bioregion in Western Australia. It has an area of . It is considered part of the larger Southwest Australia savanna ecoregion. Geography The Avon Wheatbelt bioregion is mostly a gently undulating landscape with low reli ...

Esperance Plains Esperance Plains, also known as Eyre Botanical District, is a biogeographic region in southern Western Australia on the south coast between the Avon Wheatbelt and Hampton bioregions, and bordered to the north by the Mallee region. It is a pl ...

Jarrah Forest Jarrah forest is tall open forest in which the dominant overstory tree is ''Eucalyptus marginata'' (jarrah). The ecosystem occurs only in the Southwest Botanical Province of Western Australia. It is most common in the biogeographic region named in ...

and

Swan Coastal Plain The Swan Coastal Plain in Western Australia is the geographic feature which contains the Swan River as it travels west to the Indian Ocean. The coastal plain continues well beyond the boundaries of the Swan River and its tributaries, as a geol ...

biogeographic regions A biogeographic realm or ecozone is the broadest biogeographic division of Earth's land surface, based on distributional patterns of terrestrial organisms. They are subdivided into bioregions, which are further subdivided into ecoregions. De ...

Conservation

''Pterostylis grandiflora'' is classified as "not threatened" by the Western Australian Government

Department of Parks and Wildlife The Department of Parks and Wildlife (DPaW) was the department of the Government of Western Australia responsible for managing lands described in the ''Conservation and Land Management Act 1984'' and implementing the state's conservation and e ...

References

{{Taxonbar, from=Q15494469 hamiltonii Endemic orchids of Australia Orchids of Western Australia Plants described in 1933