Pterostylis Dolichochila
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''Pterostylis dolichochila'', commonly known as the long-tongued shell orchid, is a species of
orchid Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowering ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to southern
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
. As with similar orchids, the flowering plants differ from those which are not flowering. The non-flowering plants have a rosette of leaves but the flowering plants have a single flower with leaves on the flowering spike. This greenhood has a green and white flower with reddish-brown stripes and a sharply pointed
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal co ...
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
.


Description

''Pterostylis dolichochila'' is a terrestrial,
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
,
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
,
herb In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal ...
with an underground
tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing ...
and when not flowering, a rosette of four to ten egg-shaped leaves. Each leaf is 4–12 mm long and 3–8 mm wide. Flowering plants have a single flower 20–25 mm long and 7–9 mm wide borne on a spike 50–150 mm high with three or four stem leaves wrapped around it. The flowers are green and white with reddish-brown stripes. The dorsal sepal and
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s are fused, forming a hood or "galea" over the
column A column or pillar in architecture and structural engineering is a structural element that transmits, through compression, the weight of the structure above to other structural elements below. In other words, a column is a compression member. ...
. The dorsal sepal curves forward with sharp point or a thread-like tip 1–2 mm long. The
lateral Lateral is a geometric term of location which may refer to: Healthcare *Lateral (anatomy), an anatomical direction *Lateral cricoarytenoid muscle *Lateral release (surgery), a surgical procedure on the side of a kneecap Phonetics *Lateral cons ...
sepals are held closely against the galea, have an erect, thread-like tip 15–20 mm long and a broad, flat
sinus Sinus may refer to: Anatomy * Sinus (anatomy), a sac or cavity in any organ or tissue ** Paranasal sinuses, air cavities in the cranial bones, especially those near the nose, including: *** Maxillary sinus, is the largest of the paranasal sinuses, ...
between their bases. The labellum is 13–15 mm long, about 4 mm wide, brown, blunt, and curved and protrudes above the sinus. Flowering occurs from April to August.


Taxonomy and naming

''Pterostylis dolichochila'' was first formally described in 1985 by David Jones and Mark Clements and the description was published in the fourth edition of ''Flora of South Australia''. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
(''dolichochila'') is derived from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
words ''dolichos'' meaning "long" and ''cheilos'' meaning "lip".


Distribution and habitat

The long-tongued shell orchid grows on
calcareous Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an adje ...
sand and limestone, sometimes forming large colonies, in mallee in the south-east of
South Australia South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...
and the
Little Desert Little is a synonym for small size and may refer to: Arts and entertainment * ''Little'' (album), 1990 debut album of Vic Chesnutt * ''Little'' (film), 2019 American comedy film *The Littles, a series of children's novels by American author John P ...
and
Big Desert Big or BIG may refer to: * Big, of great size or degree Film and television * ''Big'' (film), a 1988 fantasy-comedy film starring Tom Hanks * '' Big!'', a Discovery Channel television show * ''Richard Hammond's Big'', a television show present ...
areas of
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
.


References

{{Taxonbar, from=Q15500470 dolichochila Endemic orchids of Australia Orchids of South Australia Orchids of Victoria (state) Plants described in 1986