''Pterostylis despectans'', commonly known as the lowly rustyhood, is a plant in the orchid family

Orchidaceae Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowering ...

and is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to south-eastern

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

. It has a rosette of leaves at its base and up to six flowers on long stalks, branching off a short flowering stem. The flowers have an insect-like labellum and often touch the ground.

Description

''Pterostylis despectans'', is a terrestrial,

perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...

herb In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal ...

with an underground tuber. It has a rosette of between six and ten leaves at the base of the flowering spike, each leaf long and wide. The leaves are often withered by the time of flowering. Up to six translucent flowers with green and brown markings and long, wide are borne on a flowering spike tall. Each flower is carried on the end of a long, thin stalk. Three or four papery

bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...

s are wrapped around the flowering spike. The

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal co ...

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...

and

petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...

s are joined to form a hood called the "galea" over the column with the dorsal sepal having a thread-like tip long. The

lateral Lateral is a geometric term of location which may refer to: Healthcare *Lateral (anatomy), an anatomical direction *Lateral cricoarytenoid muscle *Lateral release (surgery), a surgical procedure on the side of a kneecap Phonetics *Lateral cons ...

sepals are slightly wider than the galea and often touch the ground. They are shallowly dished, densely hairy on their outer edges and suddenly taper to a thread-like tip, . The labellum is brown to green, thin and insect-like, about long and wide. The "head" end has two long bristles and there are up to 10 to 16 shorter bristles on the side. Flowering occurs from November to December.

Taxonomy and naming

The lowly greenhood was first formally described in 1950 by William Nicholls who gave it the name ''Pterostylis rufa'' var. ''despectans'' and published the description in ''

The Victorian Naturalist ''The Victorian Naturalist'' is a bimonthly scientific journal covering natural history, especially of Australia. It is published by the Field Naturalists Club of Victoria and is received as part of the membership subscription of that club. From ...

''. The type specimen was collected near Maryborough in

Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...

. In 1989, Mark Clements and David Jones raised it to species status.

Distribution and habitat

''Pterostylis despectans'' has a restricted distribution in each of the three Australian states in which it occurs. In

New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...

there is only a single population growing in grassland near

Moama Moama ( or ) is a town in the Riverina district of southern New South Wales, Australia, in the Murray River Council local government area. The town is directly across the Murray River from the larger town of Echuca in the neighbouring state of Vi ...

in the

Riverina The Riverina is an agricultural region of south-western New South Wales, Australia. The Riverina is distinguished from other Australian regions by the combination of flat plains, warm to hot climate and an ample supply of water for irrigation ...

district. In

it grows in forest near Maryborough, grassland near

Bendigo Bendigo ( ) is a city in Victoria, Australia, located in the Bendigo Valley near the geographical centre of the state and approximately north-west of Melbourne, the state capital. As of 2019, Bendigo had an urban population of 100,991, makin ...

and woodland near

Horsham Horsham is a market town on the upper reaches of the River Arun on the fringe of the Weald in West Sussex, England. The town is south south-west of London, north-west of Brighton and north-east of the county town of Chichester. Nearby to ...

and in

South Australia South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...

in parts of the Northern Lofty flora region.

Conservation

''Pterostylis despectans'' is classified as "endangered" under the Victorian Government ''

Flora and Fauna Guarantee Act 1988 The ''Flora and Fauna Guarantee Act 1988'', also known as the ''FFG Act'', is an act of the Victorian Government designed to protect species, genetic material and habitats, to prevent extinction and allow maximum genetic diversity within the Au ...

'', "critically endangered" in New South Wales, "endangered" in South Australia and as "endangered" (EN) under the Australian Government

Environment Protection and Biodiversity Conservation Act 1999 The ''Environment Protection and Biodiversity Conservation Act 1999'' (Cth) is an Act of the Parliament of Australia that provides a framework for protection of the Australian environment, including its biodiversity and its natural and cultu ...

(EPBC Act). The main threats to this species are habitat loss and fragmentation, invasive species and low reproduction rates.

References

{{Taxonbar, from=Q15499987 despectans Endemic orchids of Australia Orchids of New South Wales Orchids of South Australia Orchids of Victoria (Australia) Plants described in 1950