''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on

strawberries The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid species of the genus '' Fragaria'', collectively known as the strawberries, which are cultivated worldwide for their fruit. The fruit is widely ap ...

. They can be found under rocks and prefer slightly moist, sandy soil although preferences differ between species.

Species

''Pterostichus'' contains the following species: * '' Pterostichus aanistschenkoi'' O. Berlov & E. Berlov, 1999 * '' Pterostichus aapsorum'' Belousov, 1991 * '' Pterostichus abaciformis'' Straneo, 1955 * '' Pterostichus abagonensis'' Reitter, 1896 * '' Pterostichus abasarukini'' O. Berlov & E. Berlov, 1996 * '' Pterostichus abasinus'' Belousov, 1991 * '' Pterostichus abaxoides'' Dejean, 1828 * '' Pterostichus abensis'' Morita, 2010 * ''

Pterostichus abishirensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Belousov, 1991 * '' Pterostichus abnormis'' (J.Sahlberg, 1880) * '' Pterostichus abrogatus'' Scudder, 1890 * ''

Pterostichus abudarini ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefe ...

'' O. Berlov & E. Berlov, 1997 * ''

Pterostichus acrogonus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Chaudoir, 1843) * ''

Pterostichus acuspina ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' ( Tschitscherine, 1901) * ''

Pterostichus acutidens ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Fairmaire, 1889) * ''

Pterostichus acutipes ''Pterostichus acutipes'' is a species of woodland ground beetle in the family Carabidae. It is found in North America. Subspecies These two subspecies belong to the species ''Pterostichus acutipes'': * ''Pterostichus acutipes acutipes'' Barr, ...

'' Barr, 1971 * ''

Pterostichus adatarasanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Sasakawa, 2005 * ''

Pterostichus ademidovae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefe ...

'' O.Berlov, 1996 * ''

Pterostichus adoxus ''Pterostichus adoxus'' is a species of woodland ground beetle in the family Carabidae. It is found in North America. Larvae are predators. References Further reading * Pterostichus Articles created by Qbugbot Beetles described in 1 ...

'' (Say, 1823) * ''

Pterostichus adstrictus ''Pterostichus adstrictus'' is a species of ground beetle native to Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. ...

'' Eschscholtz, 1823 * ''

Pterostichus adulterinus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefe ...

'' Wrase & J.Schmidt, 2006 * ''

Pterostichus aemiliae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefe ...

'' Facchini & Sciaky, 2000 * ''

Pterostichus aeneocupreus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefe ...

'' (Fairmaire, 1887) * ''

Pterostichus aenescens ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefe ...

'' Chaudoir, 1850 * ''

Pterostichus aeneus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer ...

'' Fischer von Waldheim, 1829 * '' Pterostichus aereipennis'' (Solsky, 1871) * '' Pterostichus aethiops'' (Panzer, 1796) * '' Pterostichus agilis'' Allegro & Sciaky, 2010 * '' Pterostichus agonus'' G.horn, 1880 * ''

Pterostichus aibgensis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Starck, 1890 * '' Pterostichus aimaki'' Jedlicka, 1968 * '' Pterostichus ainus'' Plutenko, 2005 * '' Pterostichus akitai'' Morita, 2004 * '' Pterostichus akkusianus'' Kirschenhofer, 1981 * '' Pterostichus akozyrevi'' O. Berlov & E. Berlov, 1999 * '' Pterostichus aksekianus'' (Straneo, 1988) * ''

Pterostichus alacer ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' A.Morawitz, 1862 * '' Pterostichus alamedae'' (Casey, 1918) * '' Pterostichus alexandrovi'' Lafer, 1979 * '' Pterostichus alexeji'' Zamotajlov & Kryzhanovskij, 1992 * '' Pterostichus algidus'' Leconte, 1852 * '' Pterostichus altaianus'' Jedlicka, 1958 * ''

Pterostichus altaicus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Germar, 1824) * '' Pterostichus altaiensis'' Poppius, 1906 * '' Pterostichus amadeus'' Bousquet, 2012 * ''

Pterostichus amagisanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Tanaka & Ishida, 1972 * ''

Pterostichus amanoi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Nakane, 1968 * ''

Pterostichus amaroides ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Dejean, 1828 * '' Pterostichus ambigenus'' Bates, 1883 * ''

Pterostichus ambiguus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Fairmaire, 1858) * '' Pterostichus amblypterus'' Chaudoir, 1868 * '' Pterostichus amethystinus'' Mannerheim, 1843 * '' Pterostichus amoenus'' Dejean, 1828 * '' Pterostichus ampliatus'' Bates, 1882 * '' Pterostichus amurensis'' Poppius, 1906 * ''

Pterostichus analis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Jedlicka, 1963 * '' Pterostichus anatolicus'' Jedlicka, 1963 * ''

Pterostichus andreae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Tschitscherine, 1897) * '' Pterostichus andreinii'' Dodero, 1922 * '' Pterostichus andrewesi'' Jedlicka, 1931 * '' Pterostichus angoarnigi'' Morvan, 1994 * '' Pterostichus angustus'' (Dejean, 1828) * '' Pterostichus annamita'' (Straneo, 1939) * '' Pterostichus annosus'' Casey, 1913 * '' Pterostichus anomalus'' Chaudoir, 1878 * '' Pterostichus anomostriatus'' Sciaky, 1995 * '' Pterostichus antepunctatus'' Straneo, 1949 * '' Pterostichus anthracinus'' (Illiger, 1798) * ''

Pterostichus apenninus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Dejean, 1831 * '' Pterostichus apfelbecki'' Csiki, 1908 * '' Pterostichus apiculatiphallus'' Nemoto, 1988 * '' Pterostichus aralarensis'' Espanol & Mateu, 1945 * '' Pterostichus araraticus'' Kirschenhofer, 1987 * ''

Pterostichus arator ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Faldermann, 1836) * '' Pterostichus arcanoides'' Lorens, 1998 * ''

Pterostichus arcanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Casey, 1913 * '' Pterostichus archangaicus'' Shilenkov, 2000 * '' Pterostichus arcticola'' (Chaudoir, 1868) * '' Pterostichus arcuaticarinatus'' Kasahara, 1986 * '' Pterostichus argutoriformis'' Poppius, 1906 * '' Pterostichus aristochroides'' Deuve, 2006 * ''

Pterostichus armenus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Faldermann, 1836) * '' Pterostichus arnoldii'' Erjiomin, 1996 * '' Pterostichus arribasi'' Ortuño, 1991 * ''

Pterostichus arrowi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Jedlicka, 1936 * '' Pterostichus arrowianus'' Jedlicka, 1938 * '' Pterostichus arsenjevi'' Lafer, 1979 * '' Pterostichus arvernus'' Jeannel, 1937 * '' Pterostichus asahinus'' Habu & Baba, 1960 * '' Pterostichus asiaticus'' Poppius, 1906 * '' Pterostichus asturicus'' (Jeanne, 1965) * ''

Pterostichus astutus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Tschitscherine, 1903) * '' Pterostichus asymmetricus'' Bates, 1883 * '' Pterostichus aterrimus'' (Herbst, 1784) * ''

Pterostichus atratus ''Pterostichus atratus'' is a species of woodland ground beetle in the family Carabidae. It is sometimes considered a species of the genus Abacidus References Further reading * atratus Articles created by Qbugbot Beetles described i ...

'' Gistel, 1857 * ''

Pterostichus atrox ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Andrewes, 1937) * ''

Pterostichus aubryi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Coiffait, 1952 * ''

Pterostichus audax ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Tschitscherine, 1895) * '' Pterostichus auratus'' Heer, 1837 * '' Pterostichus auriga'' Ball, 1962 * '' Pterostichus avaricus'' Kryzhanovskij & Abdurakhmanov, 1983 * ''

Pterostichus aztecus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Tschitscherine, 1897 * '' Pterostichus baenningeri'' Jedlicka, 1931 * '' Pterostichus balachowskyi'' Morvan, 1972 * '' Pterostichus baldwini'' (Casey, 1924) * '' Pterostichus balikliensis'' Hovorka & Skoupy, 2007 * '' Pterostichus balthasari'' Jedlicka, 1937 * '' Pterostichus balu'' Schmidt, 2009 * '' Pterostichus bandotaro'' Tanaka, 1958 * '' Pterostichus bannaticus'' Dejean, 1828 * '' Pterostichus barbarae'' Sciaky & Facchini, 2003 * '' Pterostichus barbarinus'' Casey, 1913 * '' Pterostichus barbatus'' Fischer von Waldheim, 1829 * '' Pterostichus bargusinicus'' Shilenkov, 2000 * '' Pterostichus barlensis'' Straneo, 1935 * '' Pterostichus barri'' Bousquet, 2006 * '' Pterostichus barryorum'' Ball, 1962 * '' Pterostichus basilobatus'' Sasakawa, 2005 * '' Pterostichus basipunctatus'' Straneo, 1955 * ''

Pterostichus batesi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Tschitscherine, 1894) * '' Pterostichus belizini'' Lutshnik, 1933 * '' Pterostichus bellatrix'' (Tschitscherine, 1895) * '' Pterostichus belousovi'' Kryzhanovskij, 1989 * '' Pterostichus beneshi'' Sciaky, 1996 * '' Pterostichus berezowskii'' (Tschitscherine, 1898) * '' Pterostichus besucheti'' (Straneo, 1988) * '' Pterostichus beyeri'' Vandyke, 1925 * '' Pterostichus bhadravati'' Jedlicka, 1965 * '' Pterostichus bhunetansis'' Davies, 2004 * '' Pterostichus bhutanensis'' Morvan, 1980 * '' Pterostichus bicolor'' Fischer von Waldheim, 1829 * '' Pterostichus bielzii'' (Fuss, 1858) * '' Pterostichus biexcisus'' Straneo, 1955 * '' Pterostichus bifidiphallus'' Nemoto, 1988 * '' Pterostichus bifoveolatus'' Park; Kwon & Lafer, 1996 * '' Pterostichus binotatus'' Fischer von Waldheim, 1829 * '' Pterostichus biocryus'' Ball, 1962 * '' Pterostichus bischoffianus'' Jedlicka, 1936 * '' Pterostichus bisetosus'' (Straneo, 1938) * '' Pterostichus bisulcitarsis'' N.Ito, 2012 * '' Pterostichus bituberculatus'' Tschitscherine, 1899 * '' Pterostichus blandulus'' L.Miller, 1859 * '' Pterostichus blodgettensis'' Will, 2007 * '' Pterostichus blumenthali'' Heinz, 1965 * '' Pterostichus bogatshevi'' (Kryzhanovskij, 1983) * '' Pterostichus borcka'' Jedlicka, 1963 * '' Pterostichus bousqueti'' Bergdahl, 2011 * ''

Pterostichus bowanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Sciaky, 1997 * '' Pterostichus brachylobus'' Kavanaugh and LaBonte, 2006 * '' Pterostichus brachypterus'' Chaudoir, 1878 * '' Pterostichus brancuccii'' Straneo, 1982 * '' Pterostichus breuili'' Jeannel, 1919 * '' Pterostichus brevicornis'' Kirby, 1837 * '' Pterostichus brevilama'' Straneo, 1989 * '' Pterostichus brevipennis'' (Chevrolat, 1840) * ''

Pterostichus brevis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Duftschmid, 1812) * '' Pterostichus breviusculus'' R.F. Sahlberg, 1844 * '' Pterostichus brittoni'' Habu, 1958 * '' Pterostichus bruckii'' Schaum, 1859 * '' Pterostichus brunneipennis'' Straneo, 1955 * '' Pterostichus bryanti'' (Vandyke, 1951) * '' Pterostichus bryantoides'' Ball, 1962 * '' Pterostichus bucolicus'' Casey, 1913 * '' Pterostichus buglaniensis'' Kirschenhofer, 1981 * '' Pterostichus bullatus'' Allegro & Sciaky, 2010 * '' Pterostichus bungei'' (Tschitscherine, 1894) * '' Pterostichus bureli'' Morvan, 1994 * '' Pterostichus burjaticus'' Poppius, 1906 * '' Pterostichus burkhan'' O.Berlov & Anistschenko, 2005 * '' Pterostichus burmeisteri'' Heer, 1838 * '' Pterostichus bythiniensis'' (Maran, 1944) * '' Pterostichus cacumensis'' Ball, 1966 * '' Pterostichus californicus'' (Dejean, 1828) * '' Pterostichus caligans'' G.Horn, 1891 * '' Pterostichus calvitarsis'' Breit, 1912 * '' Pterostichus campania'' Andrewes, 1937 * '' Pterostichus campbelli'' Bousquet, 1985 * ''

Pterostichus canallatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Casey, 1913 * '' Pterostichus cancellatus'' (Motschulsky, 1860) * '' Pterostichus cantabricus'' Schaufuss, 1862 * '' Pterostichus cantalicus'' (Chaudoir, 1868) * '' Pterostichus capitatus'' Chaudoir, 1850 * ''

Pterostichus capito ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Tschitscherine, 1900) * '' Pterostichus capitolinus'' Kurnakov, 1962 * '' Pterostichus caribou'' Ball, 1962 * '' Pterostichus carinatus'' Sciaky & Facchini, 2003 * '' Pterostichus carinthiacus'' Meschnigg, 1946 * '' Pterostichus carolinus'' Darlington, 1932 * '' Pterostichus carradei'' Gautier des Cottes, 1866 * ''

Pterostichus carri ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Angus; Galian; Wrase & Chaladze, 2009 * '' Pterostichus carsticus'' Kasahara & Y.ito, 1989 * ''

Pterostichus casalei ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Straneo, 1984 * '' Pterostichus casaleianus'' Kirschenhofer, 1991 * '' Pterostichus caspius'' (Ménétriés, 1832) * '' Pterostichus castaneus'' (Dejean, 1828) * '' Pterostichus castanipes'' (Ménétriés, 1843) * '' Pterostichus castor'' Goulet and Bousquet, 1983 * '' Pterostichus catalonicus'' J.Daniel, 1906 * '' Pterostichus catei'' Sciaky & Wrase, 1997 * '' Pterostichus cathaicus'' Sciaky, 1994 * ''

Pterostichus caucasicola ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Tschitscherine, 1893) * '' Pterostichus caucasicus'' (Ménétriés, 1832) * '' Pterostichus caudicalis'' (Say, 1823) * '' Pterostichus cavazzutii'' Allegro & Sciaky, 2010 * '' Pterostichus cavicollis'' Straneo, 1984 * '' Pterostichus cecchiniae'' Jakobson, 1907 * '' Pterostichus cervenkai'' Sciaky, 1994 * '' Pterostichus chainapaani'' J.Schmidt, 2005 * '' Pterostichus chameleon'' Motschulsky, 1866 * '' Pterostichus champenoisi'' Croissandeau, 1893 * '' Pterostichus championi'' Andrewes, 1926 * '' Pterostichus chasautianus'' Vysoky, 1984 * '' Pterostichus chechcirensis'' Lafer, 1979 * '' Pterostichus chefsuricus'' Reitter, 1896 * '' Pterostichus chenpengi'' Li, 1992 * '' Pterostichus chinkiangensis'' Kirschenhofer, 1991 * '' Pterostichus chipewyan'' Ball, 1962 * '' Pterostichus chogyesanus'' Park; Kwon & Lafer, 1996 * '' Pterostichus chokaisanus'' Sasakawa, 2009 * '' Pterostichus chotanensis'' (Tschitscherine, 1893) * ''

Pterostichus chujoi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Habu, 1959 * '' Pterostichus chujoiellus'' Jedlicka, 1962 * '' Pterostichus chungkingi'' Jedlicka, 1932 * ''

Pterostichus chydaeus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Tschitscherine, 1897) * '' Pterostichus circulosus'' Lindroth, 1966 * '' Pterostichus clepsydra'' Sciaky & Wrase, 1997 * '' Pterostichus cochlearis'' Hacker, 1968 * ''

Pterostichus cognatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Dejean, 1831) * '' Pterostichus coimbatoris'' Jedlicka, 1969 * '' Pterostichus colasi'' Jeannel, 1937 * '' Pterostichus colchicus'' Chaudoir, 1850 * '' Pterostichus collaris'' Motschulsky, 1844 * '' Pterostichus colonus'' (Bates, 1883 * '' Pterostichus comatus'' Shi & Sciaky, 2013 * '' Pterostichus cometes'' Andrewes, 1936 * '' Pterostichus commixtiformis'' Roubal, 1920 * '' Pterostichus commutabilis'' (Motschulsky, 1866) * '' Pterostichus comorus'' Jedlicka, 1932 * '' Pterostichus compressus'' Rost, 1892 * '' Pterostichus comptorius'' Gistel, 1857 * '' Pterostichus conaensis'' J.Schmidt & Tian, 2011 * '' Pterostichus confucius'' Sciaky & Wrase, 1997 * '' Pterostichus congestus'' (Menetries, 1843) * '' Pterostichus consanguineus'' (Chaudoir, 1878) * '' Pterostichus consors'' (Tschitscherine, 1893) * '' Pterostichus convexus'' Gebler, 1847 * ''

Pterostichus coracinus ''Pterostichus coracinus'' is a species of woodland ground beetle in the family Carabidae. It is found in North America. References Further reading * coracinus Articles created by Qbugbot Beetles described in 1838 {{Pterostichi ...

'' (Newman, 1838) * '' Pterostichus cordaticollis'' Heyden, 1884 * ''

Pterostichus cordatissimus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Straneo, 1937) * '' Pterostichus cordifer'' Reitter, 1896 * '' Pterostichus coreicus'' Jedlicka, 1962 * '' Pterostichus corrusculus'' Leconte, 1873 * '' Pterostichus coruscus'' Tschitscherine, 1895 * '' Pterostichus corvinus'' (Dejean, 1828) * '' Pterostichus costatus'' (Ménétriés, 1851) * '' Pterostichus crassiapex'' Shi & Sciaky, 2013 * '' Pterostichus crassiusculus'' (Chaudoir, 1868) * '' Pterostichus craterensis'' (Hatch, 1949) * '' Pterostichus cratocephalus'' Tschitscherine, 1897 * '' Pterostichus crebrpunctatus'' Stranao, 1937 * '' Pterostichus crenicollis'' Leconte, 1873 * '' Pterostichus crenulatopunctatus'' (R.F.Sahlberg, 1844) * ''

Pterostichus creper ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Tschitscherine, 1902 * '' Pterostichus cribratus'' (Dejean, 1828) * '' Pterostichus cristatoides'' Straneo, 1955 * ''

Pterostichus cristatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (L.Dufour, 1820) * ''

Pterostichus cristicaudis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Kurnakov, 1962 * '' Pterostichus cryobioides'' Chaudoir, 1868 * '' Pterostichus cubensis'' Darlington, 1937 * '' Pterostichus cucujinus'' Reitter, 1892 * '' Pterostichus cuneatulus'' Casey, 1913 * ''

Pterostichus cunibakus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Davies, 2004 * ''

Pterostichus cursor ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Dejean, 1828 * '' Pterostichus curtatus'' Fairmaire, 1886 * '' Pterostichus curvatus'' Sciaky & Facchini, 2003 * '' Pterostichus cylindricus'' (Herbst, 1784) * '' Pterostichus cylindriformis'' Reitter, 1887 * '' Pterostichus daghestanus'' Reitter, 1896 * '' Pterostichus daihizanus'' Ishida, 1968 * '' Pterostichus daisenicus'' Ishida, 1958 * '' Pterostichus dandongensis'' Kirschenhofer, 1997 * '' Pterostichus dandonis'' Kasahara, 1989 * '' Pterostichus darvasicus'' Kryzhanovskij, 1968 * '' Pterostichus darvazicus'' Kryzhanovskij, 1968 * '' Pterostichus datshenkoae'' Sundukov, 2013 * '' Pterostichus dauricus'' (Gebler, 1832) * '' Pterostichus davidi'' Tschitscherine, 1897 * '' Pterostichus davshensis'' Shilenkov, 2000 * '' Pterostichus decastriensis'' Lafer, 1979 * '' Pterostichus deceptrix'' Tschitscherine, 1898 * '' Pterostichus defossus'' Bates, 1883 * ''

Pterostichus deino ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Will, 2007 * '' Pterostichus dentellus'' Facchini & Sciaky, 2003 * '' Pterostichus denticaudis'' Kurnakov, 1962 * '' Pterostichus dentifer'' Allegro & Sciaky, 2010 * '' Pterostichus depilatus'' (Bates, 1889) * '' Pterostichus depletus'' Scudder, 1900 * '' Pterostichus depressidorsis'' Reitter, 1896 * '' Pterostichus destitutus'' Scudder, 1890 * ''

Pterostichus destructus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Scudder, 1890 * ''

Pterostichus deuvei ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Lassalle, 1985 * '' Pterostichus deuvesianus'' Morvan, 1995 * '' Pterostichus devillei'' Puel, 1924 * '' Pterostichus dhorpatanicus'' Straneo, 1977 * '' Pterostichus diabolus'' Casey, 1913 * ''

Pterostichus diana ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' LaBonte, 2013 * '' Pterostichus diligendus'' (Chaudoir, 1868) * '' Pterostichus diligens'' (Sturm, 1824) * '' Pterostichus dilutipes'' (Motschulsky, 1844) * ''

Pterostichus dipojranus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Straneo, 1988) * '' Pterostichus discrepans'' A.Morawitz, 1862 * '' Pterostichus dissimilis'' (A. & G.B.Villa, 1833) * '' Pterostichus distinctissimus'' Jedlicka, 1940 * '' Pterostichus diversus'' (Fairmaire, 1886) * '' Pterostichus dolens'' (Tschitscherine, 1900) * '' Pterostichus doris'' Jedlicka, 1962 * '' Pterostichus dorjulensis'' Morvan, 1978 * '' Pterostichus dormitans'' Scudder, 1890 * ''

Pterostichus dostali ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Kirschenhofer, 1981 * '' Pterostichus drescheri'' (Fischer, 1817) * ''

Pterostichus drescoi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Nègre, 1957 * '' Pterostichus dudkoi'' Sundukov, 2013 * '' Pterostichus dufourii'' (Dejean, 1828) * ''

Pterostichus dulcis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Bates, 1883 * '' Pterostichus dundai'' Sciaky, 1994 * ''

Pterostichus duratii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' A. & G.B.Villa, 1835 * ''

Pterostichus dux ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' L.Schaufuss, 1862 * '' Pterostichus ebeninus'' (Dejean, 1828) * ''

Pterostichus ebenus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Quensel, 1806) * '' Pterostichus eboshiyamanus'' Sasakawa, 2009 * '' Pterostichus ecarinatus'' Hatch, 1936 * '' Pterostichus ehlersi'' Heyden, 1881 * '' Pterostichus elmbergi'' Poppius, 1908 * '' Pterostichus elongatus'' (Duftschmid, 1812) * ''

Pterostichus emei ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Sciaky, 1994 * '' Pterostichus empetricola'' (Dejean, 1828) * '' Pterostichus enasanus'' Morita, 2007 * ''

Pterostichus enodis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Bousquet, 1992 * ''

Pterostichus enyo ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Will, 2007 * '' Pterostichus eobius'' (Tschitscherine, 1899) * '' Pterostichus eoyoritomus'' Sasakawa, 2009 * '' Pterostichus epiroticus'' Csiki, 1930 * '' Pterostichus eriwanicus'' Tschitscherine, 1897 * ''

Pterostichus esakii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Ishida, 1959 * '' Pterostichus eschscholtzii'' (Germar, 1824) * '' Pterostichus espanoli'' J. Vives & E. Vives, 1977 * '' Pterostichus esuriens'' Casey, 1913 * '' Pterostichus ethnicus'' Gistel, 1857 * '' Pterostichus euxinus'' Straneo, 1935 * '' Pterostichus evanescens'' Casey, 1918 * '' Pterostichus exceptus'' J.R. Sahlberg, 1887 * ''

Pterostichus exilis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Mateu, 2001 * ''

Pterostichus eximius ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' A.Morawitz, 1862 * '' Pterostichus exochus'' Andrewes, 1930 * '' Pterostichus expeditus'' Tschitscherine, 1898 * ''

Pterostichus externepunctatus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Dejean, 1828) * '' Pterostichus falcispinus'' Sasakawa, 2005 * ''

Pterostichus faldermanni ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Schatzmayr, 1929 * '' Pterostichus fallax'' (Dejean, 1828) * '' Pterostichus fallettii'' Sciaky, 1996 * '' Pterostichus falli'' Vandyke, 1925 * '' Pterostichus farkaci'' Sciaky, 1997 * '' Pterostichus fasciatopunctatus'' (Creutzer, 1799) * ''

Pterostichus faunus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Kurnakov, 1962 * '' Pterostichus femoralis'' (Kirby, 1837) * '' Pterostichus fencli'' Dvorak, 1995 * '' Pterostichus fenestratus'' Kasahara & Ohtani, 1989 * '' Pterostichus fenyesi'' Csiki, 1930 * '' Pterostichus ferghanicus'' Kabak, 2001 * '' Pterostichus fernquisti'' Wickham, 1931 * ''

Pterostichus ferreri ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Español & Mateu, 1942) * '' Pterostichus ferreroi'' Straneo, 1989 * '' Pterostichus filipjevi'' (Lutshnik, 1928) * ''

Pterostichus filum ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Tschitscherine, 1897 * '' Pterostichus findelii'' (Dejean, 1828) * '' Pterostichus firmus'' Lutshnik, 1933 * '' Pterostichus flavofemoratus'' (Dejean, 1828) * '' Pterostichus focarilei'' Casale & Giachino, 1985 * '' Pterostichus fornicatus'' Kolenati, 1845 * '' Pterostichus foveolatus'' (Duftschmid, 1812) * '' Pterostichus fractus'' Scudder, 1890 * ''

Pterostichus franzi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Nègre, 1955 * '' Pterostichus fritzhiekei'' J.Schmidt, 1994 * '' Pterostichus fuchsi'' Schaeffer, 1910 * '' Pterostichus fujimurai'' Habu, 1958 * '' Pterostichus fujisanus'' Tanaka & Suga, 1972 * ''

Pterostichus fujitai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Tanaka & Ishida, 1972 * ''

Pterostichus fukube ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Sugimura, 2005 * '' Pterostichus fuligineus'' A.Morawitz, 1862 * '' Pterostichus fulvescens'' (Motschulsky, 1844) * '' Pterostichus funakoshii'' Morita, 2007 * '' Pterostichus funestes'' Csiki, 1930 * '' Pterostichus fuscicornis'' (Reiche & Saulcy, 1855) * '' Pterostichus gagates'' (Hope, 1831) * '' Pterostichus galae'' Farkac & Plutenko, 1996 * '' Pterostichus galaecianus'' Lauffer, 1909 * '' Pterostichus galinae'' Kabak, 1992 * ''

Pterostichus gallega ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Fairmaire, 1859) * '' Pterostichus gallopavo'' Sciaky & Wrase, 1997 * ''

Pterostichus ganesh ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' J.Schmidt, 2005 * '' Pterostichus ganja'' J.Schmidt, 1995 * '' Pterostichus gaoligongensis'' Wrase & J.Schmidt, 2006 * '' Pterostichus gassanus'' Sasakawa, 2009 * '' Pterostichus geberti'' Sciaky &, Wrase, 1997 * '' Pterostichus geojensis'' Sasakawa; Kim & Kubota, 2006 * '' Pterostichus georgi'' Straneo, 1963 * ''

Pterostichus gerdi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Schmidt, 2009 * '' Pterostichus gerstlensis'' Ball, 1962 * '' Pterostichus ghilarovi'' Kryzhanovskij, 1988 * '' Pterostichus ghilianii'' Putzeys, 1846 * '' Pterostichus giacomazzai'' Sciaky, 1994 * '' Pterostichus giacomazzoi'' Sciaky, 1994 * '' Pterostichus gibbicollis'' (Motschulsky, 1844) * '' Pterostichus giresuni'' Jedlicka, 1965 * '' Pterostichus glabricollis'' Jedlicka, 1962 * '' Pterostichus glabripennis'' Jedlicka, 1962 * '' Pterostichus glacialis'' Brisout de Barneville, 1863 * ''

Pterostichus glaferi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' O.Berlov & E.Berlov, 1996 * '' Pterostichus gliscans'' Casey, 1913 * '' Pterostichus glukhomanka'' Sundukov, 2013 * '' Pterostichus gobettii'' Straneo, 1983 * '' Pterostichus gompanus'' Straneo, 1983 * ''

Pterostichus gongga ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Sciaky, 1997 * '' Pterostichus goriensis'' (Tschitscherine, 1897) * '' Pterostichus goschi'' Jedlicka, 1930 * '' Pterostichus gotoensis'' Kasahara & Matsumoto, 1990 * '' Pterostichus gracilior'' Leconte, 1873 * '' Pterostichus gracilis'' (Dejean, 1828) * ''

Pterostichus grajus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Dejean, 1828) * '' Pterostichus gravis'' Jedlicka, 1939 * '' Pterostichus gregalis'' Casey, 1913 * '' Pterostichus gromykoi'' Sundukov, 2005 * '' Pterostichus guizhouensis'' Sciaky, 1997 * '' Pterostichus gwervaenus'' Morvan, 1995 * ''

Pterostichus gyrosus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Motschulsky, 1866) * '' Pterostichus haesitatus'' Fairmaire, 1889 * '' Pterostichus haftorni'' Lindroth, 1969 * '' Pterostichus hagenbachii'' Sturm, 1824 * '' Pterostichus hakusanus'' Kasahara, 1989 * ''

Pterostichus hamiltoni ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' G. Horn, 1880 * '' Pterostichus haptoderoides'' (Tschitscherine, 1889) * ''

Pterostichus harmandi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Tschitscherine, 1900) * '' Pterostichus harponifer'' Tanaka, 1987 * '' Pterostichus hartmanni'' J.Schmidt, 1995 * '' Pterostichus hatchi'' Hacker, 1968 * '' Pterostichus hedkeinekus'' Morvan, 1995 * '' Pterostichus heilongjiangensis'' Li, 1992 * ''

Pterostichus heinzi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Jedlicka, 1965 * ''

Pterostichus heinzianus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Sciaky, 1996 * '' Pterostichus hejkali'' B.Gueorguiev & Skoupy, 2011 * '' Pterostichus heptapotamicus'' (Lutshnik, 1927) * '' Pterostichus herculaneus'' Mannerheim, 1843 * '' Pterostichus herzi'' Poppius, 1906 * ''

Pterostichus hidanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Morita, 2010 * '' Pterostichus hikosanus'' Kasahara, 1994 * '' Pterostichus himifuriho'' Morita, 2007 * '' Pterostichus hiramatsui'' Morita, 2010 * '' Pterostichus hirasawai'' Morita, 2010 * '' Pterostichus hirobane'' Habu & Baba, 1958 * '' Pterostichus hisamatsui'' Ishida & Shibata, 1961 * '' Pterostichus hoelli'' Schmidt, 2009 * '' Pterostichus homalonotus'' Tschitscherine, 1894 * '' Pterostichus honnoratii'' (Dejean, 1828) * ''

Pterostichus hoplites ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Bates, 1883) * ''

Pterostichus hornii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Leconte, 1873 * '' Pterostichus horstweiperti'' J.Schmidt & Guorguiev, 2013 * '' Pterostichus horvatovichi'' Kirschenhofer, 1991 * '' Pterostichus hozumii'' Ishida, 1961 * '' Pterostichus huashanus'' Sciaky, 1994 * '' Pterostichus hubeicus'' Facchini & Sciaky, 2003 * '' Pterostichus hudsonicus'' Leconte, 1863 * '' Pterostichus humboldti'' Casey, 1913 * '' Pterostichus humidulus'' (Vandyke, 1943) * '' Pterostichus humilis'' Casey, 1913 * ''

Pterostichus hummeli ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Jedlicka, 1935 * '' Pterostichus hungaricus'' (Dejean, 1828) * ''

Pterostichus hypogeus ''Pterostichus hypogeus'' is a species of woodland ground beetle in the family Carabidae. It is found in North America. References Further reading * hypogeus Articles created by Qbugbot Beetles described in 1971 {{Pterostichina ...

'' Barr, 1971 * '' Pterostichus iberlovi'' O.Berlov, 1996 * '' Pterostichus idahoae'' Csiki, 1930 * '' Pterostichus ienishteai'' Nitzu, 1988 * '' Pterostichus ignavus'' (Tschitscherine, 1897) * '' Pterostichus ikizderensis'' Hovorka & Skoupy, 2007 * ''

Pterostichus ikukoae ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Morita, 2007 * '' Pterostichus illigeri'' (Panzer, 1803) * '' Pterostichus illustris'' Leconte, 1851 * ''

Pterostichus imasakai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Kasahara & Ohtani, 1988 * '' Pterostichus imitatus'' Morvan, 1978 * '' Pterostichus immarginatus'' Straneo, 1977 * ''

Pterostichus impressicollis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Fairmaire & Laboulbène, 1854) * '' Pterostichus impressus'' (Fairmaire & Laboulbène, 1854) * '' Pterostichus imurai'' Morita, 2007 * '' Pterostichus inanis'' G.Horn, 1891 * '' Pterostichus incommodus'' Schaum, 1858 * '' Pterostichus inermis'' Fall, 1901 * '' Pterostichus infaustus'' Gistel, 1857 * '' Pterostichus infernalis'' Hatch, 1936 * ''

Pterostichus infimus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Chaudoir, 1868 * ''

Pterostichus ingens ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Casey, 1918) * '' Pterostichus ingusha'' (Lutshnik, 1928) * '' Pterostichus ingushus'' (Lutshnik, 1928) * '' Pterostichus innshanensis'' Jedlicka, 1960 * '' Pterostichus inopinatus'' Lutshnik, 1933 * ''

Pterostichus inopinus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Casey, 1918) * '' Pterostichus insidatrix'' (Piochard de la Brûlerie, 1872) * '' Pterostichus insidiatrix'' (Brulerie, 1872) * '' Pterostichus insidiosus'' Fairmaire, 1866 * '' Pterostichus insignicollis'' (Tschitscherine, 1893) * '' Pterostichus intectus'' (Casey, 1918) * ''

Pterostichus interruptus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Dejean, 1828) * ''

Pterostichus irideus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Sciaky, 1994 * '' Pterostichus iripennis'' Chaudoir, 1868 * '' Pterostichus isabellae'' Leconte, 1851 * '' Pterostichus ishiharai'' Ishida & Shibata, 1961 * ''

Pterostichus ishii ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Morita; Kurosa & Mori, 2009 * '' Pterostichus ishikawai'' Nemoto, 1988 * '' Pterostichus ishikawaioides'' Sasakawa; Kim & Kubota, 2008 * '' Pterostichus ishizuchiensis'' Kasahara, 1985 * '' Pterostichus ishizukai'' Kasahara, 1995 * '' Pterostichus isolatus'' Sasakawa; Kim & Kubota, 2008 * '' Pterostichus isumiensis'' Kasahara & Saito, 1997 * '' Pterostichus itoi'' Kasahara, 1986 * '' Pterostichus iwakiensis'' Sasakawa, 2009 * '' Pterostichus ixion'' (Tschitscherine, 1902) * '' Pterostichus jacobinus'' Casey, 1913 * '' Pterostichus jacobsoni'' Poppius, 1906 * '' Pterostichus jaechi'' Kirschenhofer, 1997 * ''

Pterostichus jaechianus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Sciaky & Facchini, 2003 * '' Pterostichus jakobsonianus'' Lutshnik, 1928 * '' Pterostichus jaljaleensis'' J.Schmidt, 2009 * '' Pterostichus janatai'' Sciaky &, Wrase, 1997 * '' Pterostichus janbritoi'' Morvan, 1994 * '' Pterostichus jani'' Sciaky & Facchini, 2003 * '' Pterostichus jankowskyi'' (Tschitscherine, 1897) * ''

Pterostichus janoi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Jedlicka, 1952 * '' Pterostichus japonicus'' (Motschulsky, 1860) * '' Pterostichus jarkendis'' Jedlicka, 1965 * '' Pterostichus jelepus'' Andrewes, 1932 * '' Pterostichus jenisseicus'' Csiki, 1930 * '' Pterostichus jiricola'' Sasakawa; Kim & Kubota, 2008 * '' Pterostichus jogaesanensis'' Lafer; Paik & Park, 1996 * ''

Pterostichus johnsoni ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Ulke, 1889 * '' Pterostichus josephi'' Csiki, 1930 * ''

Pterostichus juga ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' J.schmidt, 2005 * '' Pterostichus jugicola'' (Lutshnik, 1916) * '' Pterostichus jugivagus'' (Tschitscherine, 1898) * '' Pterostichus jungens'' (Tschitscherine, 1893) * '' Pterostichus jureceki'' (Jedlicka, 1936) * '' Pterostichus jurecekianus'' Maran, 1940 * '' Pterostichus jurinei'' (Panzer, 1803) * '' Pterostichus justusii'' W.Redtenbacher, 1842 * '' Pterostichus kadjaranze'' Jedlicka, 1947 * ''

Pterostichus kadleci ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Dvorak, 1995 * '' Pterostichus kadoudali'' Morvan, 1981 * '' Pterostichus kajimurai'' Habu & Tanaka, 1957 * '' Pterostichus kalabi'' Sciaky, 1994 * '' Pterostichus kalhys'' (Motschulsky, 1866) * ''

Pterostichus kalo ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' J.Schmidt, 2012 * '' Pterostichus kambaiti'' (Andrewes, 1947) * '' Pterostichus kamtschaticus'' Motschulsky, 1860 * ''

Pterostichus kangchenjunga ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' J.Schmidt, 2012 * ''

Pterostichus kaniei ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Morita, 2005 * '' Pterostichus kaninensis'' Poppius, 1906 * ''

Pterostichus kansanus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Casey, 1918) * '' Pterostichus karasawai'' Tanaka, 1958 * '' Pterostichus karateghinicus'' Mikhailov, 1972 * ''

Pterostichus kataevi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Kryzhanovskij, 1989 * '' Pterostichus katashinensis'' Habu, 1958 * '' Pterostichus kavanaughi'' Shilenkov, 2000 * '' Pterostichus kaynashensis'' Schweiger, 1967 * '' Pterostichus keltiekus'' Morvan, 1981 * '' Pterostichus kerzhneri'' Lafer, 1983 * ''

Pterostichus kiangsu ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Jedlicka, 1965 * '' Pterostichus kiiensis'' Morita & Ohkura, 1988 * ''

Pterostichus kimurai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Morita, 1994 * '' Pterostichus kiritshenkoi'' (Lutshnik, 1928) * '' Pterostichus kirschenblatti'' Kryzhanovskij, 1988 * '' Pterostichus kitakamisanus'' Sasakawa, 2005 * '' Pterostichus klapperichi'' Jedlicka, 1956 * '' Pterostichus kleinfeldi'' Straneo, 1982 * '' Pterostichus kleinfeldianus'' Sciaky & Wrase, 1997 * '' Pterostichus koenigi'' (Reitter, 1887) * '' Pterostichus koenigianus'' (Tschitscherine, 1890) * '' Pterostichus koheii'' Nakane, 1963 * '' Pterostichus kokeilii'' L.Miller, 1850 * '' Pterostichus kolosovi'' Lutshnik, 1928 * '' Pterostichus kolymensis'' Erjiomin, 1998 * '' Pterostichus komalus'' (Jedlicka, 1936) * '' Pterostichus komiyai'' Morita, 2007 * '' Pterostichus kongosanus'' Nakane, 1963 * ''

Pterostichus kopetzi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' J.Schmidt, 2005 * '' Pterostichus korgei'' Jedlicka, 1964 * '' Pterostichus korolkowi'' (Tschitscherine, 1901) * '' Pterostichus kosakai'' Morita, 1998 * '' Pterostichus kotzebuei'' Ball, 1962 * ''

Pterostichus krali ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Sciaky, 1997 * '' Pterostichus krasnopolensis'' Kirschenhofer, 1987 * '' Pterostichus krulikovskyi'' (Lutshnik, 1933) * '' Pterostichus kubanicus'' Kirschenhofer, 1987 * '' Pterostichus kucerai'' Sciaky, 1997 * '' Pterostichus kultianus'' Jedlicka, 1946 * '' Pterostichus kuraicus'' Shilenkov, 2000 * '' Pterostichus kurentzovi'' Lafer, 1979 * '' Pterostichus kurikomasanus'' Sasakawa, 2005 * '' Pterostichus kurosai'' Tanaka, 1958 * '' Pterostichus kurosawai'' Tanaka, 1958 * '' Pterostichus kuwolsanensis'' Li & Zhang, 2014 * '' Pterostichus kvirensis'' Belousov, 1991 * '' Pterostichus kyushuensis'' Habu, 1955 * '' Pterostichus laborans'' Casey, 1913 * '' Pterostichus labzuki'' Lafer, 1979 * '' Pterostichus lacertus'' Casey, 1913 * '' Pterostichus lachrymosus'' (Newman, 1838) * '' Pterostichus lacunosus'' (Chaudoir, 1844) * '' Pterostichus laevigatus'' Scudder, 1890 * '' Pterostichus laevipunctatus'' Tschitscherine, 1889 * '' Pterostichus laferi'' O. Berlov & E. Berlov, 1996 * '' Pterostichus lama'' (Ménétriés, 1843) * '' Pterostichus lamuticus'' Poppius, 1906 * ''

Pterostichus lanei ''Pterostichus lanei'' is a species of woodland ground beetle in the family Carabidae. It is found in North America. References Further reading * * lanei Articles created by Qbugbot Beetles described in 1926 {{Pterostichinae-s ...

'' Vandyke, 1925 * ''

Pterostichus lanista ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Tschitscherine, 1898) * '' Pterostichus larisae'' Sundukov, 2013 * '' Pterostichus lasserrei'' (Dejean, 1828) * '' Pterostichus lassulus'' (Casey, 1920) * '' Pterostichus latemarginatus'' (Straneo, 1936) * '' Pterostichus laticollis'' (Motschulsky, 1844) * '' Pterostichus latifianus'' Apfelbeck, 1906 * '' Pterostichus latistylis'' Tanaka, 1958 * '' Pterostichus latitemporis'' Sciaky & Wrase, 1997 * '' Pterostichus latiusculus'' Chaudoir, 1868 * '' Pterostichus lattini'' Labonte, 2006 * '' Pterostichus ledenikensis'' (Knirsch, 1925) * ''

Pterostichus leica ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' J.Schmidt, 2012 * '' Pterostichus leonisi'' Apfelbeck, 1904 * ''

Pterostichus leptis ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Bates, 1873 * '' Pterostichus lesticoides'' (Straneo, 1939) * '' Pterostichus letensis'' Habu, 1973 * '' Pterostichus levadensis'' Lafer, 1979 * '' Pterostichus leviculus'' Andrewes, 1933 * '' Pterostichus licenti'' Jedlicka, 1939 * '' Pterostichus liciniformis'' Csiki, 1930 * '' Pterostichus lidara'' Andrewes, 1937 * '' Pterostichus lidarus'' (Andrewes, 1937) * '' Pterostichus lineatopunctatus'' L.Miller, 1850 * '' Pterostichus lingshanus'' Sciaky &, Wrase, 1997 * '' Pterostichus liodactylus'' (Tschitscherine, 1898) * '' Pterostichus lobatus'' Hacker, 1968 * ''

Pterostichus lodosi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Heinz, 1977 * ''

Pterostichus lolo ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Bergdahl, 2011 * '' Pterostichus lombardus'' K.Daniel, 1901 * '' Pterostichus longinquus'' Bates, 1873 * '' Pterostichus longipennis'' Straneo, 1942 * '' Pterostichus longipes'' Poppius, 1906 * '' Pterostichus longissimus'' Bates, 1882 * '' Pterostichus longiusculus'' (J.Sahlberg, 1880) * '' Pterostichus lubricus'' LeConte, 1853 * '' Pterostichus lucidus'' Motschulsky, 1844 * '' Pterostichus luctuosus'' (Dejean, 1828) * ''

Pterostichus lugi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Morvan, 1982 * '' Pterostichus lukjanovitshi'' Lutshnik, 1927 * '' Pterostichus lumensis'' Apfelbeck, 1905 * '' Pterostichus luscus'' (Casey, 1918) * '' Pterostichus lustrans'' Leconte, 1851 * '' Pterostichus lutschniki'' Jedlicka, 1962 * '' Pterostichus lutshnikianus'' Bogachev & Kurnakov, 1958 * '' Pterostichus lyroderus'' (Chaudoir, 1846) * '' Pterostichus macedonicus'' Apfelbeck, 1918 * ''

Pterostichus macer ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Marsham, 1802) * '' Pterostichus machulkai'' Jedlicka, 1931 * '' Pterostichus macrocephalus'' Habu, 1955 * '' Pterostichus macrogenys'' Bates, 1883 * '' Pterostichus macrothorax'' Poppius, 1906 * '' Pterostichus maderi'' Jedlicka, 1938 * '' Pterostichus madidus'' (Fabricius, 1775) * '' Pterostichus maeklini'' (Poppius, 1906) * '' Pterostichus magoides'' (Straneo, 1937) * '' Pterostichus magus'' (Mannerheim, 1825) * '' Pterostichus maichensis'' Lafer, 1979 * ''

Pterostichus makalu ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' J.Schmidt, 2012 * '' Pterostichus makolskii'' J.Muller, 1933 * '' Pterostichus malaisei'' Jedlicka, 1965 * '' Pterostichus malissorum'' Apfelbeck, 1905 * '' Pterostichus malkini'' (Hatch, 1953) * '' Pterostichus mancus'' (Leconte, 1852) * '' Pterostichus mandibularoides'' Ball, 1966 * '' Pterostichus mandzhuricus'' Lutshnik, 1916 * '' Pterostichus mannerheimii'' (Dejean, 1831) * '' Pterostichus marani'' Jedlicka, 1967 * '' Pterostichus marginatus'' Matsumura, 1911 * '' Pterostichus marginellus'' Sasakawa; Kim & Kim, 2006 * '' Pterostichus mariae'' (Lutshnik, 1921) * '' Pterostichus marinensis'' Hacker, 1968 * '' Pterostichus masahiroi'' Kasahara, 1988 * '' Pterostichus masatakai'' Morita, 2007 * '' Pterostichus masidai'' Ishida, 1959 * '' Pterostichus masumotoi'' Tanaka; Morita & Suga, 1987 * '' Pterostichus matchai'' Jedlicka, 1930 * '' Pterostichus matsumurai'' Habu, 1973 * '' Pterostichus mattolensis'' Hacker, 1968 * '' Pterostichus matveichuki'' O.Berlov, 1996 * '' Pterostichus maurusiacus'' Mannerheim, 1825 * ''

Pterostichus maximus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Tschitscherine, 1889 * ''

Pterostichus mediocris ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Dvorak, 1995 * '' Pterostichus medvedevi'' Kryzhanovskij & Mikhailov, 1972 * '' Pterostichus megacephalus'' Sciaky & Allegro, 2013 * '' Pterostichus megaloderus'' Sciaky, 1994 * '' Pterostichus meisteri'' Reitter, 1885 * ''

Pterostichus melanarius ''Pterostichus melanarius'', the Rain-Beetle, is a species of ground beetle native to Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its hist ...

'' (Illiger, 1798) * '' Pterostichus melanodes'' (Chaudoir, 1878) * '' Pterostichus melas'' (Creutzer, 1799) * '' Pterostichus mellyi'' (Gebler, 1843) * '' Pterostichus menetriesii'' Leconte, 1873 * '' Pterostichus mercedianus'' (Casey, 1918) * '' Pterostichus merkli'' (J.Frivaldszky, 1879) * '' Pterostichus merklii'' (J.Frivaldszky, 1879) * '' Pterostichus meskheticus'' Belousov, 1991 * '' Pterostichus metallifer'' Gistel, 1857 * '' Pterostichus mewakholensis'' J.Schmidt, 2012 * ''

Pterostichus meyeri ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Jedlicka, 1934 * '' Pterostichus micans'' Heer, 1838 * '' Pterostichus michailovi'' Wrase, 1992 * ''

Pterostichus microbus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Sciaky & Facchini, 2003 * '' Pterostichus microcephalus'' (Motschulsky, 1860) * '' Pterostichus microphthalmus'' Delarouzée, 1857 * ''

Pterostichus microps ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Heyden, 1887 * '' Pterostichus middendorffi'' J.R. Sahlberg, 1875 * ''

Pterostichus miehei ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' J.Schmidt, 2005 * '' Pterostichus migliaccioi'' Straneo, 1982 * '' Pterostichus militaris'' (Tschitscherine, 1893) * '' Pterostichus mimus'' Casey, 1913 * ''

Pterostichus minax ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Oustalet, 1874) * ''

Pterostichus ming ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Sciaky &, Wrase, 1997 * ''

Pterostichus minor ''Pterostichus minor'' is a species of woodland ground beetle native to Europe.Carl H. Lindroth 1974 Coleoptera. Carabidae. ''Handbooks for the Identification of British Insects'' Vol 4 Part 2. Royal Entomological Society, Londopdf References ...

'' (Gyllenhal, 1827) * '' Pterostichus minutulus'' Heer, 1862 * '' Pterostichus mirabilis'' Jedlicka, 1939 * '' Pterostichus mirificus'' Bates, 1883 * ''

Pterostichus miroshnikovi ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Zamotajlov, 1991 * '' Pterostichus miroslavi'' Sciaky &, Wrase, 1997 * ''

Pterostichus mirus ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' (Tschitscherine, 1894) * '' Pterostichus miscellus'' Casey, 1913 * '' Pterostichus mitoyamanus'' Tanaka, 1971 * '' Pterostichus miyamai'' Kasahara & Y.Ito, 1987 * ''

Pterostichus mizunoyai ''Pterostichus'' is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and pr ...

'' Morita, 2007 * '' Pterostichus modicellus'' Tschitscherine, 1897 * '' Pterostichus moestus'' (Say, 1823) * ''Pterostichus molopsoides'' Jedlicka, 1934 * ''Pterostichus monostigma'' (Tschitscherine, 1898) * ''Pterostichus montanus'' (Motschulsky, 1844) * ''Pterostichus monticoloides'' Shilenkov, 1995 * ''Pterostichus montigena'' (Tschitscherine, 1898) * ''Pterostichus montisdeorum'' Schweiger, 1967 * ''Pterostichus morawitzianus'' (Lutshnik, 1922) * ''Pterostichus morio'' (Duftschmid, 1812) * ''Pterostichus morionides'' (Chaudoir, 1868) * ''Pterostichus mosaicus'' Sasakawa, 2005 * ''Pterostichus mucronatus'' Straneo, 1955 * ''Pterostichus muehlfeldii'' (Duftschmid, 1812) * ''Pterostichus muelleri'' (Straneo, 1936) * ''Pterostichus muellermotzfeldi'' Wrase & J.Schmidt, 2006 * ''Pterostichus mujahedeeni'' Savich, 1999 * ''Pterostichus mukdenensis'' Breit, 1933 * ''Pterostichus mulensis'' Sciaky, 1997 * ''Pterostichus multinodosus'' Sasakawa; Kim & Kim, 2006 * ''Pterostichus multipunctatus'' (Dejean, 1828) * ''Pterostichus multiseta'' (Straneo, 1984) * ''Pterostichus mundatus'' Jedlicka, 1940 * ''Pterostichus mundus'' Jedlicka, 1938 * ''Pterostichus musashiensis'' Kasahara, 1993 * ''Pterostichus mutoides'' Bousquet, 1992 * ''Pterostichus mutus'' (Say, 1823) * ''Pterostichus nadari'' (Vuillefroy, 1893) * ''Pterostichus nakanei'' Straneo, 1955 * ''Pterostichus namrum'' Jedlicka, 1963 * ''Pterostichus naokii'' Morita, 2003 * ''Pterostichus napaea'' Kasahara, 1988 * ''Pterostichus nasuensis'' Morita, 2007 * ''Pterostichus nasui'' Kasahara, 1993 * ''Pterostichus necessarius'' (Tschitscherine, 1894) * ''Pterostichus neglectus'' A.Morawitz, 1862 * ''Pterostichus negligens'' Sturm, 1824 * ''Pterostichus neilgaimani'' Chaladze, Kalatozishvili, Janiashvili, & Bakuradze 2017 * ''Pterostichus nemoralis'' Graells, 1851 * ''Pterostichus neobrunneus'' Lindroth, 1966 * ''Pterostichus nepalensis'' Straneo, 1977 * ''Pterostichus nicaeensis'' A. Villa & C.B. Villa, 1835 * ''Pterostichus nigellatus'' Kirschenhofer, 1991 * ''Pterostichus nigellus'' A.Morawitz, 1862 * ''Pterostichus niger'' (Schaller, 1783) * ''Pterostichus nigripalpis'' Poppius, 1906 * ''Pterostichus nigrita'' (Paykull, 1790) * ''Pterostichus nigrocaeruleus'' Vandyke, 1925 * ''Pterostichus nigrocoerulus'' * ''Pterostichus nikodymi'' Dvorak, 1995 * ''Pterostichus nimbatidius'' Chaudoir, 1878 * ''Pterostichus nishidai'' Nakane, 1989 * ''Pterostichus nishiyamai'' Kasahara, 1986 * ''Pterostichus nivalis'' R.F. Sahlberg, 1844 * ''Pterostichus nivicola'' (Ménétriés, 1832) * ''Pterostichus noburai'' Morita, 2008 * ''Pterostichus nodicornis'' (Fairmaire & Laboulbène, 1854) * ''Pterostichus noguchii'' Bates, 1873 * ''Pterostichus nordqvisti'' J.R. Sahlberg, 1887 * ''Pterostichus novotnyorum'' Vysoky, 1981 * ''Pterostichus novus'' Straneo, 1944 * ''Pterostichus nowitzkii'' (Tschitscherine, 1899) * ''Pterostichus obesulus'' Leconte, 1873 * ''Pterostichus oblongiusculus'' Motschulsky, 1850 * ''Pterostichus oblongoparallelus'' Maran, 1932 * ''Pterostichus oblongopunctatus'' (Fabricius, 1787) * ''Pterostichus obsidianus'' Casey, 1913 * ''Pterostichus occultus'' Casey, 1913 * ''Pterostichus ochoticus'' R.F. Sahlberg, 1844 * ''Pterostichus ochraceus'' Gautier des Cottes, 1868 * ''Pterostichus ochsi'' Fagniez, 1921 * ''Pterostichus odontocnemis'' G.Müller, 1932 * ''Pterostichus odvarkai'' Dvorak, 1995 * ''Pterostichus oenotrius'' Ravizza, 1975 * ''Pterostichus ogaensis'' Morita, 1995 * ''Pterostichus ohbayashii'' Ishida, 1968 * ''Pterostichus ohdaisanus'' Nakane, 1963 * ''Pterostichus ohkawai'' Morita, 2010 * ''Pterostichus ohkurai'' Morita, 1996 * ''Pterostichus ohsawacavus'' Sasakawa, 2005 * ''Pterostichus okiensis'' Nakane, 1989 * ''Pterostichus okutamae'' Tanaka, 1963 * ''Pterostichus olafi'' J.Schmidt, 2005 * ''Pterostichus olegi'' Belousov, 1991 * ''Pterostichus omogoensis'' Nakane, 1972 * ''Pterostichus ompoensis'' Jedlicka, 1932 * ''Pterostichus opacipennis'' Jedlicka, 1934 * ''Pterostichus ophryoderus'' (Chaudoir, 1878) * ''Pterostichus ordinarius'' Casey, 1913 * ''Pterostichus ordinatus'' (Fischer, 1823) * ''Pterostichus orduensis'' (Straneo, 1988) * ''Pterostichus oregonus'' Leconte, 1861 * ''Pterostichus oreophilus'' (Tschitscherine, 1898) * ''Pterostichus orestes'' (Jedlicka, 1936) * ''Pterostichus orientalis'' (Motschulsky, 1844) * ''Pterostichus orion'' (Tschitscherine, 1901) * ''Pterostichus orionis'' Jedlicka, 1962 * ''Pterostichus orman'' Jedlicka, 1963 * ''Pterostichus orobius'' Fairmaire, 1861 * ''Pterostichus oscitans'' Tschitscherine, 1900 * ''Pterostichus osseticus'' Kirschenhofer, 1987 * ''Pterostichus ottomanus'' Apfelbeck, 1908 * ''Pterostichus ovalipennis'' Casey, 1913 * ''Pterostichus ovaliphallus'' Sasakawa, 2005 * ''Pterostichus ovoideus'' (Sturm, 1824) * ''Pterostichus pachinus'' Bates, 1883 * ''Pterostichus palekhai'' E. Berlov & O. Berlov, 1996 * ''Pterostichus palgongsanus'' Nemoto, 1988 * ''Pterostichus palmi'' Schaeffer, 1910 * ''Pterostichus panticulatus'' Casey, 1913 * ''Pterostichus pantomus'' Jedlicka, 1958 * ''Pterostichus panzeri'' (Panzer, 1803) * ''Pterostichus paralleloides'' Kirschenhofer, 1987 * ''Pterostichus parasimilis'' G.E. Ball, 1962 * ''Pterostichus parkwon'' Davies, 2004 * ''Pterostichus parnassius'' Schaum, 1859 * ''Pterostichus parviceps'' Poppius, 1906 * ''Pterostichus parvicollis'' Sciaky &, Wrase, 1997 * ''Pterostichus pasanauricus'' Kirschenhofer, 1987 * ''Pterostichus patruelis'' (Dejean, 1831) * ''Pterostichus paulinoi'' (Vuillefroy, 1868) * ''Pterostichus pavani'' (Straneo, 1988) * ''Pterostichus pedemontanus'' Ganglbauer, 1891 * ''Pterostichus peilingi'' Jedlicka, 1937 * ''Pterostichus pemphredo'' Will, 2007 * ''Pterostichus peninsularis'' Park & Kwon, 1996 * ''Pterostichus pensylvanicus'' Leconte, 1873 * ''Pterostichus pentheri'' Apfelbeck, 1918 * ''Pterostichus percontator'' Reitter, 1887 * ''Pterostichus pergracilis'' (Casey, 1920) * ''Pterostichus perhoplites'' J.Schmidt & Tian, 2011 * ''Pterostichus perisi'' Novoa, 1979 * ''Pterostichus perlamatus'' Straneo, 1989 * ''Pterostichus perlutus'' Jedlicka, 1938 * ''Pterostichus permundus'' (Say, 1830) * ''Pterostichus pertinax'' (Tschitscherine, 1895) * ''Pterostichus petulans'' Jedlicka, 1938 * ''Pterostichus pfizenmayeri'' Poppius, 1906 * ''Pterostichus phaeopus'' (Sainte-Claire Deville, 1902) * ''Pterostichus phaeus'' (Lutshnik, 1922) * ''Pterostichus phungaraziensis'' Wrase & J.Schmidt, 2006 * ''Pterostichus piceolus'' (Chaudoir, 1850) * ''Pterostichus picipes'' Fischer von Waldheim, 1829 * ''Pterostichus pilifer'' (Bates, 1878) * ''Pterostichus piliferus'' (Bates, 1878) * ''Pterostichus pilosus'' (Host, 1789) * ''Pterostichus pinguedineus'' Eschscholtz, 1823 * ''Pterostichus piniphilus'' Gistel, 1857 * ''Pterostichus pithyusicus'' Lutshnik, 1933 * ''Pterostichus placerensis'' (Casey, 1918) * ''Pterostichus planaticollis'' Kirschenhofer, 1987 * ''Pterostichus planctus'' Leconte, 1852 * ''Pterostichus planicola'' (Tschitscherine, 1899) * ''Pterostichus planiusculus'' (Chaudoir, 1859) * ''Pterostichus planus'' J.R. Sahlberg, 1887 * ''Pterostichus platyderus'' Chaudoir, 1850 * ''Pterostichus platyops'' Sciaky, 1997 * ''Pterostichus plesiomorphus'' Nemoto, 1989 * ''Pterostichus plutonicus'' Casey, 1913 * ''Pterostichus poecilma'' Andrewes, 1937 * ''Pterostichus pohnerti'' Jedlicka, 1934 * ''Pterostichus polygenus'' Bates, 1883 * ''Pterostichus ponticus'' Kirschenhofer, 1987 * ''Pterostichus poppiusi'' (Semenov, 1906) * ''Pterostichus poppiusianus'' Jakobson, 1907 * ''Pterostichus porcellus'' Kurnakov, 1962 * ''Pterostichus potanini'' Tschitscherine, 1889 * ''Pterostichus praedo'' (Tschitscherine, 1901) * ''Pterostichus praetermissus'' (Chaudoir, 1868) * ''Pterostichus pratti'' Bates, 1890 * ''Pterostichus prattii'' Bates, 1890 * ''Pterostichus primus'' Darlington, 1932 * ''Pterostichus probus'' Park & Kwon, 1996 * ''Pterostichus procax'' A.Morawitz, 1862 * ''Pterostichus procephalus'' Bates, 1873 * ''Pterostichus procerulus'' Heyden, 1880 * ''Pterostichus properans'' Chaudoir, 1868 * ''Pterostichus protensiformis'' (Casey, 1924) * ''Pterostichus protensipennis'' (Casey, 1918) * ''Pterostichus protractus'' Leconte, 1860 * ''Pterostichus provincialis'' (Oustalet, 1874) * ''Pterostichus przewalskyi'' Tschitscherine, 1888 * ''Pterostichus pseodostuxbergi'' Poppius, 1906 * ''Pterostichus pseudoastutus'' Hovorka & Skoupy, 2007 * ''Pterostichus pseudobarbarae'' Sciaky & Facchini, 2003 * ''Pterostichus pseudodiversus'' Shi & Sciaky, 2013 * ''Pterostichus pseudoharmandi'' Morvan, 1981 * ''Pterostichus pseudojugivagus'' J.Schmidt & Tian, 2011 * ''Pterostichus pseudopachinus'' Nakane, 1963 * ''Pterostichus pseudopedius'' Reitter, 1887 * ''Pterostichus pseudopercus'' Reitter, 1889 * ''Pterostichus pseudopilifer'' (Straneo, 1957) * ''Pterostichus pseudoplatyderus'' Sciaky, 1994 * ''Pterostichus pseudoplatysma'' Straneo, 1982 * ''Pterostichus pseudorotundus'' Sciaky & Facchini, 2003 * ''Pterostichus pseudosinensis'' Sciaky & Facchini, 2003 * ''Pterostichus pseudostuxbergi'' (Poppius, 1906) * ''Pterostichus pulchellus'' Faldermann, 1836 * ''Pterostichus pulcher'' Sciaky & Allegro, 2013 * ''Pterostichus pumilio'' Dejean, 1828 * ''Pterostichus pumilus'' Casey, 1913 * ''Pterostichus pumpellyi'' Scudder, 1900 * ''Pterostichus punctatissimus'' (Randall, 1838) * ''Pterostichus punctatus'' L. Redtenbacher, 1843 * ''Pterostichus punctibasalis'' N.Ito, 2012 * ''Pterostichus punctiger'' J.R. Sahlberg, 1880 * ''Pterostichus punctisternus'' Straneo, 1989 * ''Pterostichus punctiventris'' (Chaudoir, 1878) * ''Pterostichus pusillus'' Dejean, 1828 * ''Pterostichus putus'' Casey, 1913 * ''Pterostichus quadraticollis'' Chaudoir, 1846 * ''Pterostichus quadrifoveolatus'' Letzner, 1852 * ''Pterostichus quadriimpressus'' Straneo, 1984 * ''Pterostichus quelpartensis'' Habu, 1985 * ''Pterostichus raptor'' (Tschitscherine, 1901) * ''Pterostichus raraensis'' Morvan, 1980 * ''Pterostichus rasilis'' Park & Kwon, 1996 * ''Pterostichus regeli'' (Tschitscherine, 1894) * ''Pterostichus reiseri'' Ganglbauer, 1889 * ''Pterostichus reissi'' G.Müller, 1931 * ''Pterostichus relictus'' (Newman, 1838) * ''Pterostichus rengensis'' Morita, 2010 * ''Pterostichus restrictus'' (Casey, 1918) * ''Pterostichus reuteri'' Wrase & Schmidt, 2006 * ''Pterostichus rhaeticus'' Heer, 1837 * ''Pterostichus rhanis'' (Tschitscherine, 1902) * ''Pterostichus rhilensis'' Rottenberg, 1874 * ''Pterostichus rifensis'' Antoine, 1933 * ''Pterostichus riffensis'' Antoine, 1933 * ''Pterostichus riparius'' (Dejean, 1828) * ''Pterostichus ripensis'' (Motschulsky, 1866) * ''Pterostichus robustistylis'' Sasakawa, 2005 * ''Pterostichus rodionovi'' Shilenkov 2000 RU (Chita Area) * ''Pterostichus rolex'' Morvan, 1995 * ''Pterostichus rostratus'' (Newman, 1838) * ''Pterostichus rothi'' (Hatch, 1951) * ''Pterostichus rotundangulus'' A.Morawitz, 1862 * ''Pterostichus rotundus'' Sciaky, 1994 * ''Pterostichus rousi'' Kirschenhofer, 1982 * ''Pterostichus rousianus'' Kirschenhofer, 1987 * ''Pterostichus rousselli'' Colas, 1963 * ''Pterostichus rubripalpis'' Csiki, 1930 * ''Pterostichus rudestriatus'' (Reitter, 1883) * ''Pterostichus ruffoi'' Sciaky, 1984 * ''Pterostichus rufipes'' (Dejean, 1828) * ''Pterostichus rufitarsis'' (Dejean, 1828) * ''Pterostichus rufonitens'' Fairmaire, 1866 * ''Pterostichus rufopiceus'' Heyden, 1890 * ''Pterostichus rugiceps'' (Straneo, 1984) * ''Pterostichus rugosiceps'' J.Schmidt, 2012 * ''Pterostichus rugosipennis'' Jedlicka, 1932 * ''Pterostichus rugosus'' (Gebler, 1823) * ''Pterostichus rutenus'' (Jeannel, 1942) * ''Pterostichus rutilans'' (Dejean, 1828) * ''Pterostichus ryomoensis'' Morita & Suda, 2007 * ''Pterostichus sacheri'' I.Frivaldszky von Frivald, 1865 * ''Pterostichus sachtlebeni'' Jedlicka, 1962 * ''Pterostichus safonovae'' Budarin, 1995 * ''Pterostichus sagarmatha'' J.Schmidt, 2012 * ''Pterostichus sahlbergi'' (Tschitscherine, 1894) * ''Pterostichus sakagamii'' Morita, 2001 * ''Pterostichus sakuraii'' Morita, 2008 * ''Pterostichus santostamangi'' J.Schmidt, 2006 * ''Pterostichus sasajii'' Morita, 2007 * ''Pterostichus satoi'' Ishida, 1961 * ''Pterostichus satsumanus'' Habu, 1958 * ''Pterostichus satunini'' Tschitscherine, 1903 * ''Pterostichus satyrus'' Kurnakov, 1962 * ''Pterostichus saueri'' Sciaky, 1994 * ''Pterostichus saxicola'' (Tschitscherine, 1899) * ''Pterostichus sayanus'' Csiki, 1930 * ''Pterostichus scalptus'' Sciaky & Wrase, 1997 * ''Pterostichus schaschli'' (Marseul, 1880) * ''Pterostichus schmidtii'' (Chaudoir, 1838) * ''Pterostichus schneideri'' Sciaky & Facchini, 2003 * ''Pterostichus schodaicus'' Kirschenhofer, 1987 * ''Pterostichus schoenherri'' Faldermann, 1836 * ''Pterostichus schoenmanni'' Kirschenhofer, 1991 * ''Pterostichus schrettenbrunneri'' J.Schmidt, 2012 * ''Pterostichus schuelkei'' Sciaky & Wrase, 1997 * ''Pterostichus scitus'' Maklin, 1878 * ''Pterostichus sculpturalis'' N.Ito, 2012 * ''Pterostichus sculptus'' LeConte, 1853 * ''Pterostichus scurra'' (Tschitscherine, 1901) * ''Pterostichus scutellaris'' Leconte, 1873 * ''Pterostichus scuticollis'' (Fairmaire, 1889) * ''Pterostichus sediorcus'' Sasakawa & Kim, 2006 * ''Pterostichus sejunctus'' Bates, 1883 * ''Pterostichus sejungendus'' (Chaudoir, 1868) * ''Pterostichus selmanni'' (Duftschmid, 1812) * ''Pterostichus semenovi'' Tschitscherine, 1888 * ''Pterostichus semirugosus'' (Andrewes, 1947) * ''Pterostichus septemtrionalis'' Sciaky & Facchini, 2003 * ''Pterostichus septentrionis'' (Chaudoir, 1868) * ''Pterostichus serbicus'' Apfelbeck, 1899 * ''Pterostichus seriatus'' (Chaudoir, 1850) * ''Pterostichus serratipes'' Chaudoir, 1878 * ''Pterostichus serripes'' (Leconte, 1875) * ''Pterostichus seticeps'' (Straneo, 1984) * ''Pterostichus setipes'' (Tschitscherine, 1898) * ''Pterostichus setosicus'' Davies, 2004 * ''Pterostichus setosus'' Hatch, 1951 * ''Pterostichus seunglaki'' Park & Kwon, 1996 * ''Pterostichus seungmoi'' Park & Kwon, 1996 * ''Pterostichus shaovuensis'' Jedlicka, 1956 * ''Pterostichus sharanus'' (Straneo, 1984) * ''Pterostichus shavrini'' O.Berlov & Anistschenko, 1999 * ''Pterostichus shennongjianus'' Facchini & Sciaky, 2003 * ''Pterostichus shibatai'' Ishida, 1961 * ''Pterostichus shiibanus'' Habu, 1958 * ''Pterostichus shilenkovi'' Erjiomin & Kabak, 1991 * ''Pterostichus shimizui'' Morita, 2010 * ''Pterostichus shinbodakensis'' Sasakawa & H.Ito, 2017 * ''Pterostichus shingarevi'' Lafer, 1979 * ''Pterostichus shirakamisan'' Sasakawa, 2009 * ''Pterostichus shirakamisanus'' Sasakawa, 2005 * ''Pterostichus shiranus'' (Straneo, 1984) * ''Pterostichus shirozui'' Habu, 1985 * ''Pterostichus shojii'' Sugimura, 2006 * ''Pterostichus shotaroi'' Morita, 1987 * ''Pterostichus shulli'' (Hatch, 1949) * ''Pterostichus sierranus'' Casey, 1913 * ''Pterostichus similatus'' Gautier des Cottes, 1870 * ''Pterostichus similis'' Mannerheim, 1852 * ''Pterostichus simillimus'' Fairmaire, 1886 * ''Pterostichus sincerus'' Park & Kwon, 1996 * ''Pterostichus sinensis'' Jedlicka, 1962 * ''Pterostichus singularis'' Tschitscherine, 1889 * ''Pterostichus sinicus'' (Fairmaire, 1897) * ''Pterostichus sintanus'' Andrewes, 1927 * ''Pterostichus skoupyi'' Hovorka & Skoupy, 2007 * ''Pterostichus smetanai'' Bousquet, 1985 * ''Pterostichus sodalicius'' Heyden, 1885 * ''Pterostichus sojaki'' Kirschenhofer, 1987 * ''Pterostichus sojot'' Shilenkov, 2000 * ''Pterostichus solskyi'' (Chaudoir, 1878) * ''Pterostichus songoricus'' (Motschulsky, 1845) * ''Pterostichus soperi'' Ball, 1966 * ''Pterostichus sotkaensis'' Jedlicka, 1958 * ''Pterostichus spathifer'' Bousquet, 1992 * ''Pterostichus sphodriformis'' Bates, 1873 * ''Pterostichus sphodrinus'' Leconte, 1863 * ''Pterostichus spiculifer'' Bates, 1883 * ''Pterostichus spinolae'' (Dejean, 1828) * ''Pterostichus sponsor'' Casey, 1913 * ''Pterostichus spornyi'' Sundukov, 2013 * ''Pterostichus spraguei'' Leconte, 1873 * ''Pterostichus ssemenovi'' Tschitscherine, 1888 * ''Pterostichus stagnatilis'' Gistel, 1857 * ''Pterostichus stantonensis'' Ball, 1966 * ''Pterostichus stapedius'' Hacker, 1968 * ''Pterostichus starcki'' Heyden, 1885 * ''Pterostichus starckianus'' Reitter, 1887 * ''Pterostichus staveni'' Hovorka & Skoupy, 2007 * ''Pterostichus sterbai'' Jedlicka, 1934 * ''Pterostichus stictopleurus'' (Fairmaire, 1889) * ''Pterostichus stoeckleini'' (Straneo, 1941) * ''Pterostichus stomoides'' Chaudoir, 1868 * ''Pterostichus straneellus'' Jedlicka, 1938 * ''Pterostichus strasseri'' Reitter, 1898 * ''Pterostichus strenuus'' (Panzer, 1796) * ''Pterostichus stricticollis'' (Solsky, 1874) * ''Pterostichus strigipennis'' N.Ito, 2012 * ''Pterostichus strigosus'' Sciaky & Wrase, 1997 * ''Pterostichus stygicus'' (Say, 1823) * ''Pterostichus styx'' Andrewes, 1937 * ''Pterostichus subabacipennis'' N.Ito, 2012 * ''Pterostichus subacutus'' (Casey, 1918) * ''Pterostichus subaeneus'' (Chaudoir, 1850) * ''Pterostichus subgibbus'' Motschulsky, 1860 * ''Pterostichus subiasi'' Ortuno & Zaballos, 1992 * ''Pterostichus subitus'' Csiki, 1930 * ''Pterostichus sublaevis'' (J.Sahlberg, 1880) * ''Pterostichus subovatus'' (Motschulsky, 1860) * ''Pterostichus subrugosus'' Straneo, 1955 * ''Pterostichus subsinuatus'' Dejean, 1828 * ''Pterostichus subtilissimus'' Sciaky, 1996 * ''Pterostichus sudai'' Morita, 1991 * ''Pterostichus sudrei'' Sainte-Claire Deville, 1922 * ''Pterostichus suffusus'' Casey, 1913 * ''Pterostichus sugimurai'' Morita, 2007 * ''Pterostichus sulcitarsis'' A.Morawitz, 1863 * ''Pterostichus sumondakensis'' Sasakawa, 2005 * ''Pterostichus sungariensis'' Lafer, 1979 * ''Pterostichus superciliosus'' (Say, 1823) * ''Pterostichus surgens'' Leconte, 1878 * ''Pterostichus suruganus'' Straneo, 1979 * ''Pterostichus sutschanensis'' Jedlicka, 1962 * ''Pterostichus svanicus'' (Lutshnik, 1922) * ''Pterostichus swaneticus'' (Reitter, 1883) * ''Pterostichus syleus'' Kirschenhofer, 1997 * ''Pterostichus symmetricus'' Straneo, 1955 * ''Pterostichus szekessyianus'' Sciaky, 1996 * ''Pterostichus szetschuanensis'' Tschitscherine, 1889 * ''Pterostichus tachongi'' Jedlicka, 1936 * ''Pterostichus taebaegsanus'' Nemoto, 1988 * ''Pterostichus taeniolus'' Bates, 1882 * ''Pterostichus tahirai'' Kasahara, 1992 * ''Pterostichus tahoensis'' Casey, 1913 * ''Pterostichus takadateyamanus'' Sasakawa, 2009 * ''Pterostichus takahashii'' Ishida, 1958 * ''Pterostichus takaosanus'' Habu, 1958 * ''Pterostichus taksonyis'' Csiki, 1930 * ''Pterostichus talibani'' Savich, 1999 * ''Pterostichus tamarae'' Wrase & Kirschenhofer, 1991 * ''Pterostichus tamsii'' (Dejean, 1831) * ''Pterostichus tanakai'' Ishida, 1964 * ''Pterostichus tanakaorum'' Morita & Ohkura, 1988 * ''Pterostichus tanjae'' Hovorka & Skoupy, 2007 * ''Pterostichus tantillus'' Fairmaire, 1888 * ''Pterostichus taoi'' Kasahara, 1993 * ''Pterostichus taradakensis'' Kasahara & Ohtani, 1988 * ''Pterostichus tareumiut'' G. E. Ball, 1962 * ''Pterostichus tarsalis'' Leconte, 1873 * ''Pterostichus taskylensis'' Shilenkov, 2000 * ''Pterostichus tateishiyamanus'' Sasakawa & H.Ito, 2017 * ''Pterostichus tatianae'' E.Berlov, 1996 * ''Pterostichus tatricus'' Kult, 1947 * ''Pterostichus tenenbaumianus'' Jedlicka, 1930 * ''Pterostichus tenuimarginatus'' Chaudoir, 1868 * ''Pterostichus tenuis'' (Casey, 1924) * ''Pterostichus teretis'' Park & Kwon, 1996 * ''Pterostichus testaceus'' (Vandyke, 1926) * ''Pterostichus thanglaensis'' J.Schmidt, 2012 * ''Pterostichus theeli'' Maklin, 1878 * ''Pterostichus thorectes'' Bates, 1873 * ''Pterostichus thorectoides'' Jedlicka, 1958 * ''Pterostichus thunbergi'' A.Morawitz, 1862 * ''Pterostichus tichomirovi'' Erjiomin, 1990 * ''Pterostichus tienmushanus'' Sciaky, 1997 * ''Pterostichus tiliaceoradix'' Ball, 1962 * ''Pterostichus todai'' Morita & Kanie, 1997 * ''Pterostichus tokejii'' Yoshida & Tanaka, 1960 * ''Pterostichus tokmakovae'' Sundukov, 2013 * ''Pterostichus tokui'' Morita, 2007 * ''Pterostichus toledanoi'' Facchini & Sciaky, 2003 * ''Pterostichus tomensis'' (Gebler, 1847) * ''Pterostichus tonkinensis'' Straneo, 1980 * ''Pterostichus tosanus'' Kasahara & Y.ito, 1997 * ''Pterostichus tottoriensis'' Morita, 2003 * ''Pterostichus toyodai'' Morita & Kurosa, 1998 * ''Pterostichus transversalis'' (Duftschmid, 1812) * ''Pterostichus triangularis'' Sciaky & Facchini, 2003 * ''Pterostichus trinarius'' (Casey, 1918) * ''Pterostichus trinitensis'' Hacker, 1968 * ''Pterostichus triseriatus'' (Gebler, 1847) * ''Pterostichus tristis'' (Dejean, 1828) * ''Pterostichus troglophilus'' Mateu, 1997 * ''Pterostichus tropicalis'' Bates, 1882 * ''Pterostichus trubilini'' Zamotajlov, 1997 * ''Pterostichus truncatus'' (Dejean, 1828) * ''Pterostichus tsagerkericus'' (Lutshnik, 1928) * ''Pterostichus tschitscherinianus'' (Jakobson, 1907) * ''Pterostichus tsukubasanus'' Kasahara, 1988 * ''Pterostichus tsurugaensis'' Ishida, 1959 * ''Pterostichus tsurugiyamanus'' Habu, 1959 * ''Pterostichus tuberculiger'' (Tschitscherine, 1897) * ''Pterostichus tuberculofemoratus'' Hatch, 1936 * ''Pterostichus tuberifer'' Sasakawa, 2006 * ''Pterostichus tulobalu'' Schmidt, 2009 * ''Pterostichus tumescens'' Leconte, 1863 * ''Pterostichus tundrae'' (Tschitscherine, 1894) * ''Pterostichus tunkinensis'' Shilenkov, 2000 * ''Pterostichus turanensis'' Jedlicka, 1959 * ''Pterostichus turcomannicus'' Motschulsky, 1850 * ''Pterostichus uchiyamai'' Morita, 1987 * ''Pterostichus udokanensis'' Shilenkov, 2000 * ''Pterostichus uedaorum'' Morita & Hirasawa, 1996 * ''Pterostichus uenoi'' Straneo, 1955 * ''Pterostichus ulubeyensis'' Kirschenhofer, 1981 * ''Pterostichus umbralis'' Gistel, 1857 * ''Pterostichus umbricola'' Gistel, 1857 * ''Pterostichus unctulatus'' Duftschmid, 1812 * ''Pterostichus uralensis'' Motschulsky, 1850 * ''Pterostichus urengaicus'' Jurecek, 1924 * ''Pterostichus ussuriensis'' (Tschitscherine, 1897) * ''Pterostichus vagepunctatus'' Heer, 1837 * ''Pterostichus validiceps'' Reitter, 1887 * ''Pterostichus validior'' Tschitscherine, 1889 * ''Pterostichus validus'' (Dejean, 1828) * ''Pterostichus vandykei'' Schaeffer, 1910 * ''Pterostichus variabilis'' (Ménétriés, 1832) * ''Pterostichus variicornis'' Sasakawa; Kim & Kim, 2009 * ''Pterostichus variolatus'' (Dejean, 1828) * ''Pterostichus variseta'' (Straneo, 1984) * ''Pterostichus vecors'' (Tschitscherine, 1897) * ''Pterostichus ventralis'' (Say, 1823) * ''Pterostichus ventricosus'' Eschscholtz, 1823 * ''Pterostichus vermiculosus'' (Ménétriés, 1851) * ''Pterostichus vernalis'' Panzer, 1796 * ''Pterostichus vexatus'' Bousquet, 1985 * ''Pterostichus vicinus'' Mannerheim, 1843 * ''Pterostichus violaceotinctus'' Straneo, 1953 * ''Pterostichus virescens'' (Gebler, 1833) * ''Pterostichus viridellus'' Straneo, 1984 * ''Pterostichus vladivostokensis'' Lafer, 1979 * ''Pterostichus vseteckai'' (Maran, 1944) * ''Pterostichus vulgaris'' (Linnaeus, 1758) * ''Pterostichus walcotti'' Scudder, 1900 * ''Pterostichus walteri'' Reitter, 1883 * ''Pterostichus watanabei'' Nakane, 1960 * ''Pterostichus weigeli'' J.Schmidt, 2005 * ''Pterostichus weiperti'' J.Schmidt, 2005 * ''Pterostichus wenxianensis'' Allegro & Sciaky, 2010 * ''Pterostichus wittmeri'' Morvan, 1978 * ''Pterostichus woodi'' Ball and Currie, 1997 * ''Pterostichus woongbii'' Park & Kwon, 1996 * ''Pterostichus woronowi'' (Lutshnik, 1916) * ''Pterostichus wrasei'' Sciaky & Facchini, 2003 * ''Pterostichus xatartii'' (Dejean, 1828) * ''Pterostichus xilingensis'' Allegro & Sciaky, 2010 * ''Pterostichus yahikosanus'' Sasakawa, 2009 * ''Pterostichus yakushimanus'' Nakane & Ishida, 1961 * ''Pterostichus yamajii'' Kasahara, 1993 * ''Pterostichus yamauchii'' Morita, 1992 * ''Pterostichus yamizosanus'' Sasakawa, 2005 * ''Pterostichus yanoi'' Jedlicka, 1953 * ''Pterostichus yatsuensis'' Straneo, 1955 * ''Pterostichus ybousqueti'' O.Berlov, 1999 * ''Pterostichus yokohamae'' Nakane & Straneo in Nakane, 1979 * ''Pterostichus yorikoae'' Sugimura, 2006 * ''Pterostichus yoritomus'' Bates, 1873 * ''Pterostichus yosemitensis'' Hacker, 1968 * ''Pterostichus yoshidai'' Kasahara, 1985 * ''Pterostichus yoshikawai'' Ishida, 1959 * ''Pterostichus yoshizawai'' Morita, 2010 * ''Pterostichus yulongshanensis'' Sciaky, 1997 * ''Pterostichus yunnanensis'' Jedlicka, 1934 * ''Pterostichus yvanii'' (Dejean, 1828) * ''Pterostichus zamotajlovi'' Belousov, 1991 * ''Pterostichus zhejiangensis'' Kirschenhofer, 1997 * ''Pterostichus ziegleri'' (Duftschmid, 1812) * ''Pterostichus zoiai'' Sciaky, 1996 * ''Pterostichus zolotarewi'' Reitter, 1911 * ''Pterostichus zunianus'' Casey, 1913 * ''Pterostichus zyzzovi'' O. Berlov & E. Berlov, 1996

References

{{Taxonbar, from=Q1540893 Pterostichus, Pterostichinae Taxa named by Franco Andrea Bonelli