''Pseudotsuga'' is a genus of
evergreen
In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
coniferous tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s in the family
Pinaceae
The Pinaceae, or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as cedars, firs, hemlocks, larches, pines and spruces. The family is included in the order Pinales, formerly know ...
(subfamily
Laricoideae
The Laricoideae are a subfamily of the Pinaceae, a Pinophyta division family. They take their name from the genus ''Larix'' ( larches), which contains inside most of the species of the group and is one of only two deciduous genera of the pine ...
).
Common names for species in the genus include Douglas fir, Douglas-fir, Douglas tree, Oregon pine and Bigcone spruce. ''
Pseudotsuga menziesii
The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three ...
'' (Douglas fir proper) is widespread in western North America and is an important source of timber. The number of species has long been debated, but two in western
North America and two to four in eastern
Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an ...
are commonly acknowledged.
Nineteenth-century botanists had problems in classifying Douglas firs, due to the species' similarity to various other conifers better known at the time; they have at times been classified in ''
Pinus
A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden a ...
'', ''
Picea
A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfam ...
'', ''
Abies
Firs (''Abies'') are a genus of 48–56 species of evergreen coniferous trees in the family Pinaceae. They are found on mountains throughout much of North and Central America, Europe, Asia, and North Africa. The genus is most closely related to ...
'', ''
Tsuga
''Tsuga'' (, from Japanese (), the name of ''Tsuga sieboldii'') is a genus of conifers in the subfamily Abietoideae of Pinaceae, the pine family. The common name hemlock is derived from a perceived similarity in the smell of its crushed foliage ...
'', and even ''
Sequoia''. Because of their distinctive cones, Douglas firs were finally placed in the new genus ''Pseudotsuga'' (meaning "false hemlock") by the French botanist
Carrière in 1867. The genus name has also been hyphenated as ''Pseudo-tsuga''.
Name
The tree takes its English name from
David Douglas, the
Scottish
Scottish usually refers to something of, from, or related to Scotland, including:
*Scottish Gaelic, a Celtic Goidelic language of the Indo-European language family native to Scotland
*Scottish English
*Scottish national identity, the Scottish ide ...
botanist who first introduced ''
Pseudotsuga menziesii
The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three ...
'' into cultivation at
Scone Palace
Scone Palace is a Category A- listed historic house near the village of Scone and the city of Perth, Scotland. Built in red sandstone with a castellated roof, it is an example of the Gothic Revival style in Scotland.
Scone was originally the ...
in 1827.
[Trust Walks: "Dunkeld and The Hermitage]
," a podcast by the National Trust for Scotland; 27 June 2009 Douglas is known for introducing many native American tree species to Europe. The hyphenated form "Douglas-fir" is used by some to indicate that ''Pseudotsuga'' species are not true
fir
Firs (''Abies'') are a genus of 48–56 species of evergreen coniferous trees in the family (biology), family Pinaceae. They are found on mountains throughout much of North America, North and Central America, Europe, Asia, and North Africa. The ...
s, which belong to the genus ''Abies''.
Description
Douglas-firs are medium-size to extremely large
evergreen
In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s, tall (although only coast Douglas-firs reach such great height). The
leaves are flat, soft, linear, long, generally resembling those of the firs, occurring singly rather than in
fascicles; they completely encircle the branches, which can be useful in recognizing the species. The female
cones are pendulous, with persistent scales (unlike true firs), and are distinctive in having a long tridentine (three-pointed)
bract that protrudes prominently above each scale (it resembles the back half of a mouse, with two feet and a tail).
''Pseudotsuga menziesii'' var. ''menziesii'' has attained heights of 393 feet (120* m). That was the estimated height of the tallest conifer ever well-documented, the Mineral Tree (
Mineral, Washington), measured in 1924 by Dr. Richard E. McArdle, former chief of the U.S. Forest Service. The volume of that tree was . The tallest living individual is the Brummitt (Doerner) Fir in
Coos County, Oregon
Coos County ( ) is one of the 36 counties in the U.S. state of Oregon. As of the 2020 census, the population was 64,929. The county seat is Coquille. The county was formed from the western parts of Umpqua and Jackson counties. It is named ...
, tall.
[Gymnosperm Database: ''Pseudotsuga menziesii'' var. ''menziesii'' (2006)]
/ref> Only coast redwood and '' Eucalyptus regnans'' reach greater heights based on current knowledge of living trees: 379 and 331 feet (116 and 101* m), respectively.
At Quinault, Washington, is found a collection of the largest Douglas-firs in one area. Quinault Rain Forest hosts the most of the top ten known largest Douglas-firs.
, the largest known Douglas-firs in the world are, by volume:
# Red Creek Tree (Red Creek, SW British Columbia)
# Queets Fir (Queets River Valley-Olympic National Park)
# Tichipawa (Quinault Lake Rain Forest-Olympic National Park)
# Rex (Quinault Lake Rain Forest-Olympic National Park)
# Ol' Jed (Jedediah Smith Redwoods State Park)
Species and varieties
By far the best-known is the very widespread and abundant North American species ''Pseudotsuga menziesii
The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three ...
'', a taxonomically complex species divided into two major varieties (treated as distinct species
In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
or subspecies by some botanists): coast Douglas-fir or "green Douglas-fir", on the Pacific
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
coast; and Rocky Mountain Douglas-fir or "interior Douglas-fir", in the interior west of the continent. According to some botanists, Rocky Mountain Douglas-fir extends south into Mexico to include all Mexican Douglas-fir populations, whereas others have proposed multiple separate species in Mexico and multiple varieties in the United States. Morphological and genetic evidence suggest that Mexican Douglas-fir should probably be considered a distinct variety within ''P. menziesii''.
All of the other species are of restricted range and little-known outside of their respective native environments, where they are often rare and of scattered occurrence in mixed forests; all those have unfavorable conservation status
The conservation status of a group of organisms (for instance, a species) indicates whether the group still exists and how likely the group is to become extinct in the near future. Many factors are taken into account when assessing conservatio ...
. The taxonomy of the Asian Douglas-firs continues to be disputed, but the most recent taxonomic treatment accepts four species: three Chinese and one Japanese.[ The three Chinese species have been variously considered varieties of ''P. sinensis''] or broken down into additional species and varieties. In the current treatment, the Chinese species ''P. sinensis'' is further subdivided into two varieties: var. ''sinensis'' and var. ''wilsoniana''.
North America
* '' Pseudotsuga macrocarpa'' ( Vasey) Mayr
Mayr is a German surname. Notable people with the surname include:
* Andrea Mayr (born 1979), Austrian female long-distance runner
* Ernst Mayr (1904–2005), German American evolutionary biologist
* Georg Mayr (1564–1623), Bavarian Jesuit prie ...
– bigcone Douglas-fir - southern California
* ''Pseudotsuga menziesii
The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three ...
'' ( Mirb.) Franco
Franco may refer to:
Name
* Franco (name)
* Francisco Franco (1892–1975), Spanish general and dictator of Spain from 1939 to 1975
* Franco Luambo (1938–1989), Congolese musician, the "Grand Maître"
Prefix
* Franco, a prefix used when ...
- western North America from Alaska
Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S ...
to Oaxaca
)
, population_note =
, population_rank = 10th
, timezone1 = CST
, utc_offset1 = −6
, timezone1_DST = CDT
, utc_offset1_DST = −5
, postal_code_type = Postal ...
** ''Pseudotsuga menziesii'' var. ''menziesii'' – coast Douglas-fir
** ''Pseudotsuga menziesii'' var. ''glauca'' ( Beissn.) Franco – Rocky Mountain Douglas-fir
** ''Pseudotsuga menziesii var. lindleyana
''Pseudotsuga menziesii var. lindleyana'', commonly known as the Mexican Douglas-fir, is a conifer in the genus ''Pseudotsuga'' that is endemic to Mexico. DNA sequence and morphological evidence suggests it is most closely related to Rocky Mo ...
'' (Roezl
Benedikt Roezl (13 August 1824, Horomeritz (Bohemia, Austrian Empire) – 14 October 1885, Prague) was a traveller, gardener and botanist (sometimes Benedict or Benito Roezl as called by the Indians). Probably the most famous collector of orchids ...
) Carrière – Mexican Douglas-fir
Asia
* ''Pseudotsuga brevifolia
''Pseudotsuga'' is a genus of evergreen coniferous trees in the family Pinaceae (subfamily Laricoideae).
Common names for species in the genus include Douglas fir, Douglas-fir, Douglas tree, Oregon pine and Bigcone spruce. ''Pseudotsuga menzie ...
'' W.C.Cheng & L.K.Fu – short-leaf Chinese Douglas-fir
* ''Pseudotsuga forrestii
''Pseudotsuga'' is a genus of evergreen coniferous trees in the family Pinaceae (subfamily Laricoideae).
Common names for species in the genus include Douglas fir, Douglas-fir, Douglas tree, Oregon pine and Bigcone spruce. '' Pseudotsuga m ...
'' Craib – Yunnan Douglas-fir
* '' Pseudotsuga japonica'' ( Shiras.) Beissn. – Japanese Douglas-fir
* ''Pseudotsuga sinensis
''Pseudotsuga sinensis'' (Chinese Douglas-fir; in Chinese 黃杉, pinyin romanization: huáng shān) is a species of conifer in the family Pinaceae. It is a tree up to 50 metres tall. It is found in China (in Anhui, Fujian, Guangxi, Guizhou, Hub ...
'' Dode – Chinese Douglas-fir
** ''Pseudotsuga sinensis'' var. ''sinensis''
** ''Pseudotsuga sinensis'' var. ''wilsoniana'' – Taiwan Douglas-fir
** ''Pseudotsuga sinensis'' var. ''gaussenii ''
Formerly placed in ''Pseudotsuga''
* '' Keteleeria davidiana'' (Bertrand) Beissn. (as ''P. davidiana'' Bertrand)
* ''Cathaya argyrophylla
''Cathaya'' is a genus in the pine family, Pinaceae, with one known living species, ''Cathaya argyrophylla''. ''Cathaya'' is a member of the subfamily Laricoideae, most closely related to ''Pseudotsuga'' and ''Larix''. A second species, ''C. nan ...
'' (as ''P. argyrophylla)''
* '' Keteleeria fortunei'' (as ''P. fortunei)''
* '' Abies magnifica'' (as ''P. magnifica)''
* '' Abies procera'' (as ''P. nobilis)''
Uses
Douglas-fir wood
Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of ligni ...
is used for structural applications that are required to withstand high loads. It is used extensively in the construction industry. Other examples include its use for homebuilt aircraft
An aircraft is a vehicle that is able to fly by gaining support from the air. It counters the force of gravity by using either static lift or by using the dynamic lift of an airfoil, or in a few cases the downward thrust from jet engines. ...
such as the RJ.03 IBIS canard. Very often, these aircraft were designed to utilize Sitka spruce
''Picea sitchensis'', the Sitka spruce, is a large, coniferous, evergreen tree growing to almost tall, with a trunk diameter at breast height that can exceed 5 m (16 ft). It is by far the largest species of spruce and the fifth-larg ...
, which is becoming increasingly difficult to source in aviation quality grades. Oregon pine is also used in boat building
Boat building is the design and construction of boats and their systems. This includes at a minimum a hull, with propulsion, mechanical, navigation, safety and other systems as a craft requires.
Construction materials and methods
Wood
W ...
when it is available in long, fairly knot
A knot is an intentional complication in Rope, cordage which may be practical or decorative, or both. Practical knots are classified by function, including List of hitch knots, hitches, List of bend knots, bends, List of loop knots, loop knots, ...
-free lengths. Most timber now comes from plantation forests in North America which are managed to produce faster growing timber with fewer knots. This timber is generally lighter but weaker. Traditionally, Oregon pine was used in mast
Mast, MAST or MASt may refer to:
Engineering
* Mast (sailing), a vertical spar on a sailing ship
* Flagmast, a pole for flying a flag
* Guyed mast, a structure supported by guy-wires
* Mooring mast, a structure for docking an airship
* Radio mast ...
building due to its ability to resist bending loads without fracturing. This was based on using older native forest wood with a high number of growth rings per inch. This sort of wood is seldom available new but can be sourced from merchants dealing in recycled timber. Native Oregon pine is considerably heavier than Sitka spruce
''Picea sitchensis'', the Sitka spruce, is a large, coniferous, evergreen tree growing to almost tall, with a trunk diameter at breast height that can exceed 5 m (16 ft). It is by far the largest species of spruce and the fifth-larg ...
, which is about the same weight as western red cedar
''Thuja plicata'' is an evergreen coniferous tree in the cypress family Cupressaceae, native to western North America. Its common name is western redcedar (western red cedar in the UK), and it is also called Pacific redcedar, giant arborvitae, w ...
, but with far better bending characteristics than cedar. Large-sized Oregon pine, as used in beams, is inclined to split as it dries, like oak, but this does not reduce its strength.
Douglas-fir is one of the most commonly marketed Christmas tree
A Christmas tree is a decorated tree, usually an evergreen conifer, such as a spruce, pine or fir, or an artificial tree of similar appearance, associated with the celebration of Christmas. The custom was further developed in early modern G ...
species in the United States, where they are sold alongside firs like noble fir and grand fir. Douglas-fir Christmas trees are usually trimmed to a near perfect cone instead of left to grow naturally like noble and grand firs.
Pests and diseases
Douglas-firs are used as food plants by the larva
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle.
Th ...
e of some Lepidoptera species, including autumnal moth
The autumnal moth (''Epirrita autumnata'') is a moth of the family Geometridae. The species was first described by Moritz Balthasar Borkhausen in 1794. It is found throughout the Palearctic region and the Near East and has a much wider distribut ...
, bordered white, engrailed moth, pine beauty and turnip moth. The gelechiids ''Chionodes abella
''Chionodes abella'' is a moth in the family Gelechiidae. It is found in North America, where it has been recorded from Idaho and south-western British Columbia to Colorado, Texas and California.
The wingspan is about 15 mm. The forewings a ...
'' and '' Chionodes periculella'' and the tortrix moth
The Tortricidae are a family of moths, commonly known as tortrix moths or leafroller moths, in the order Lepidoptera. This large family has over 11,000 species described, and is the sole member of the superfamily Tortricoidea, although the gen ...
''Cydia illutana
__NOTOC__
''Cydia illutana'' is a small moth of the family Tortricidae. It is found from western and central Europe (Great Britain, the Netherlands, Austria, Germany and France), north to Scandinavia (Denmark, Norway, Sweden and Finland) and east ...
'' have been specifically recorded on ''P. menziesii''.
Culture
A California Native American myth explains that each three-ended bract is the tail and two tiny legs of a mouse
A mouse ( : mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
that hid inside the scales of the tree's cones during forest fires, and the tree was kind enough to be its enduring sanctuary.
A Douglas-fir species, ''Pseudotsuga menziesii
The Douglas fir (''Pseudotsuga menziesii'') is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three ...
'', is the state tree of Oregon
Oregon () is a state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington, while the Snake River delineates much of its eastern boundary with Idah ...
.
References
External links
Gymnosperm Database - ''Pseudotsuga''
{{Taxonbar, from=Q158771
Conifer genera
Forestry in Canada
Forestry in the United States
Taxa named by Élie-Abel Carrière