The ''Prolecithophora'' are an

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...

consisting of an estimated 300 species of small (typically 0.2 – 12 mm, one species up to 50 mm), active, aquatic

flatworm The flatworms, flat worms, Platyhelminthes, or platyhelminths (from the Greek πλατύ, ''platy'', meaning "flat" and ἕλμινς (root: ἑλμινθ-), ''helminth-'', meaning "worm") are a phylum of relatively simple bilaterian, unsegment ...

s. The order lacks a common English name. Most species are shaped like an elongated, stylized droplet, and are opaque white or yellow; they frequently have contrasting bands or spots in colors, such as purple, yellow, red, or brown. They have no to three (normally two) pairs of pigment-cup eyes, and well-developed

tactile Tactile may refer to: * Tactile, related to the sense of touch * Haptics (disambiguation) * Tactile (device), a text-to-braille translation device See also * Tangibility, in law * Somatosensory system, where sensations are processed * CD96 CD ...

and

chemoreceptor A chemoreceptor, also known as chemosensor, is a specialized sensory receptor which transduces a chemical substance (endogenous or induced) to generate a biological signal. This signal may be in the form of an action potential, if the chemorecept ...

senses. With few exceptions, species are protandric hermaphrodites with

internal fertilization Internal fertilization is the union of an egg and sperm cell during sexual reproduction inside the female body. Internal fertilization, unlike its counterpart, external fertilization, brings more control to the female with reproduction. For inter ...

. Egg capsules are, according to species, glued to various hard surfaces; the young hatch as miniature copies of their parents.

Ecology

All prolecithophorans are aquatic, with most living in the

ocean The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the wo ...

s. Some species, especially those living in freshwater, are

predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...

s and

scavengers Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding be ...

, but many marine species are associated with colonial animals such as bryozoans or live as

symbionts Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasit ...

on larger animals such as urchins; a few species harbor symbiotic algae. Although most are accomplished swimmers, they normally rarely venture far from the bottom; young specimens are sometimes found in

plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...

. Many species display positive or negative

phototaxis Phototaxis is a kind of taxis, or locomotory movement, that occurs when a whole organism moves towards or away from a stimulus of light. This is advantageous for phototrophic organisms as they can orient themselves most efficiently to receive li ...

Distribution

The majority of prolecithophorans live in the oceans. There are also

brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...

and

freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...

species; in

Lake Baikal Lake Baikal (, russian: Oзеро Байкал, Ozero Baykal ); mn, Байгал нуур, Baigal nuur) is a rift lake in Russia. It is situated in southern Siberia, between the federal subjects of Irkutsk Oblast to the northwest and the Repu ...

, an

adaptive radiation In evolutionary biology, adaptive radiation is a process in which organisms diversify rapidly from an ancestral species into a multitude of new forms, particularly when a change in the environment makes new resources available, alters biotic int ...

has resulted in a number of

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

species. The order has a

cosmopolitan distribution In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all or most of the world in appropriate habitats. Such a taxon, usually a species, is said to exhibit cosmopolitanism or cosmopolitism. The ext ...

; most described species are from

temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...

waters, while little is known about

tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...

or deep-sea species. This is probably due to a sampling artefact, as prolecithophorans are known to be common in the tropics; when Norén & Jondelius sampled the shore adjacent to Phuket Marine Biological Center,

Phuket Phuket (; th, ภูเก็ต, , ms, Bukit or ''Tongkah''; Hokkien:普吉; ) is one of the southern provinces (''changwat'') of Thailand. It consists of the island of Phuket, the country's largest island, and another 32 smaller islands of ...

, Thailand, they found 14 species of prolecithophorans, all of which were new to science.

Families

The following families are recognised in the order Prolecithophora: * Multipeniatidae Karling, 1940 * Plagiostomidae - mainly marine, a few freshwater species (e.g.

Vänern Vänern ( , also , ) is the largest lake in Sweden, the largest lake in the European Union and the third-largest lake of all Europe after Ladoga and Onega in Russia. It is located in the provinces of Västergötland, Dalsland, and Värmland in ...

Lake Biwa is the largest freshwater lake in Japan, located entirely within Shiga Prefecture (west-central Honshu), northeast of the former capital city of Kyoto. Lake Biwa is an ancient lake, over 4 million years old. It is estimated to be the 13th ol ...

and

Lake Tanganyika Lake Tanganyika () is an African Great Lake. It is the second-oldest freshwater lake in the world, the second-largest by volume, and the second-deepest, in all cases after Lake Baikal in Siberia. It is the world's longest freshwater lake. ...

) * Protomonotresidae Reisinger, 1924 - exclusively freshwater species from Lake Baikal, with one exception * Pseudostomidae Graff, 1904-08 - exclusively marine species * Scleraulophoridae Marcus, 1950 * Urostomatidae

References

{{Taxonbar, from=Q3028843 Turbellaria