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''Pseudorhombus arsius'', the largetooth flounder, is a species of left-eyed
flatfish A flatfish is a member of the Ray-finned fish, ray-finned demersal fish order (biology), order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the ...
, that is the dark side with the eyes on the adult fish is the left side of the fish's body, from the family
Paralichthyidae Large-tooth flounders or sand flounders are a family, Paralichthyidae, of flounders. The family contains 14 genera with a total of about 110 species. They lie on the sea bed on their right side; both eyes are always on the left side of the head, ...
. As ''Rhombus polyspilos'' it was named as the type species of the genus '' Pseudorhombus''. It is an
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
species and is fished for by both recreational and commercial fisheries.


Description

''Pseudorhombus arsius'' has an oval-shaped body which has a depth which is 1.8 to 2.3 times its
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
. The head has a slight notch in front of upper eye. The eyes are on the left side and have a diameter which is a fifth of the length of the head and the snout has a length equal to or slightly greater than the diameter of the eye. The dorsal fin starts in front of its upper eye and of the nostrils on blind side. The mouth forms a deep cleft with the maxilla normally almost half the length of the head. The jaws and teeth are nearly equally developed on both jaws. The upper jaw reaches to below the posterior edge of the lower eye. There are several pairs of moderately large caniform teeth in the anterior parts of both jaws, which are of unequal size and arranged in a single row. These number 5 to 8 in the lower jaw on the eyed side, and 6 to 13 lateral teeth in lower jaw of blind side which are more robust and more widely spaced than those of upper jaw. The gill rakers are pointed, longer than they are broad. It has simple fin-rays which number 71–84 in the dorsal fin, 53–62 in the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
and 11–13 in the pectoral fin. The
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
curves above the pectoral fin has 69-81 scales and there are 36 vertebrae. In the dorsal fin the longest rays are found in the posterior third and only the rearmost three or four rays are branched, as at the last five anal fin rays. Two dark spots are situated on the central part of its dorsal surface, one just to the posterior of the pectoral fin and the second is midway between that the forward spot and the tail. The body is greenish to pale brownish in colour and is normally marked with variously sized rings, and there is frequently two dark blotches on the straight and the curved sections of the lateral line with a smaller blotch which is half-way to the caudal-fin peduncle. It can vary the colour of the body so that it closely matches the surface the fish rests on. They grow to 50 cm, but 30 cm in length is the more usual size


Distribution

''Pseudorhombus arsius'' is found in the tropical and temperate waters from the Persian Gulf and the eastern coast of Africa as far south as
Algoa Bay Algoa Bay is a maritime bay in the Eastern Cape, South Africa. It is located in the east coast, east of the Cape of Good Hope. Algoa Bay is bounded in the west by Cape Recife and in the east by Cape Padrone. The bay is up to deep. The harbour c ...
and possibly even Knysna to
Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...
in the western Pacific, to southern Japan in the north and as far south as the northern coast of Australia. It may extend as far south as the
Bass Strait Bass Strait () is a strait separating the island state of Tasmania from the Australian mainland (more specifically the coast of Victoria, with the exception of the land border across Boundary Islet). The strait provides the most direct waterwa ...
.


Habitat and biology

''Pseudorhombus arsius'' occurs in shallow waters and in estuaries where the substrate consists of mud and sand bottoms, to depths of 200 m. The juveniles are common in brackish water. When they are spawning they are found in shallow water on sandbanks and close to shore. They move to deeper waters in the winter. They are predators which prey mainly on
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
animals. The spawning season runs for 3 to 4 months, from April to July and peaks in April and May, during which they move into the sandy shallows close to the shore. The spawn is laid in a single batch. They mature when they attain a total length of between 16 and 17 cm in total length. The sex ratio is always biased towards females and the fecundity of the fish is dependent on its total length and body weight.


Fisheries

''Pseudorhombus arsius'' is subjected to a minor commercial fishery in West Bengal in the estuary of the
Hooghly River The Bhagirathi Hooghly River (Anglicized alternatively spelled ''Hoogli'' or ''Hugli'') or the 'Bhāgirathi-Hooghly', called the Ganga or the Kati-Ganga in mythological texts, is the eastern distributary of the Ganges River in West Bengal, Indi ...
. Otherwise it is a quarry for recreational angling.


References

{{Taxonbar, from=Q127620 Paralichthyidae Fish of the Indian Ocean Fish of the Pacific Ocean Fish described in 1822 Taxa named by Francis Buchanan-Hamilton