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''Psephophorus'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus of sea turtle that lived from the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
to the Pliocene. Its remains have been found in Europe, Africa,
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, and New Zealand. It was first named by Hermann von Meyer in 1847, and contains seven species, ''P. polygonus'', ''P. calvertensis'', ''P. eocaenus'', ''P. oregonesis'', ''P. californiensis'', ''P. rupeliensis'',''Proceedings''; page 8. By the Zoological Society of London; published 1891. Retrieved on June 28th, 2008. ''P. scaldii'', and a species discovered in 1995, ''P. terrypratchetti''. ''Psephophorus'' is the only Miocene dermochelyid turtle found in Europe. One species of ''Psephophorus'' could measure up to ten feet in length.


Discovery and identification

Von Meyer originally named ''Psephophorus'' in 1846. At first he was unable to identify the creature beyond its
dermal plate The dermis or corium is a layer of skin between the epidermis (with which it makes up the cutis) and subcutaneous tissues, that primarily consists of dense irregular connective tissue and cushions the body from stress and strain. It is divided in ...
s, but when he later received a drawing he was able to describe the specimen, which was then in
Pressburg Bratislava (, also ; ; german: Preßburg/Pressburg ; hu, Pozsony) is the capital and largest city of Slovakia. Officially, the population of the city is about 475,000; however, it is estimated to be more than 660,000 — approximately 140% of ...
, as a fragment of a
carapace A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tor ...
, which contained seventy bones. In 1879,
H. G. Seeley Harry Govier Seeley (18 February 1839 – 8 January 1909) was a British paleontologist. Early life Seeley was born in London on 18 February 1839, the second son of Richard Hovill Seeley, a goldsmith, and his second wife Mary Govier. When his fat ...
was asked to study the ''Psephophorus'' specimen by Franz Ritter von Hauer, the Director of the Austro-Hungarian
Imperial and Royal Geological Survey Imperial is that which relates to an empire, emperor, or imperialism. Imperial or The Imperial may also refer to: Places United States * Imperial, California * Imperial, Missouri * Imperial, Nebraska * Imperial, Pennsylvania * Imperial, Te ...
. Up until then, the specimen's identity had been undeterminable, with even Seeley describing it at first to seem like "the dermal covering of an
Edentate Xenarthra (; from Ancient Greek ξένος, xénos, "foreign, alien" + ἄρθρον, árthron, "joint") is a major clade of placental mammals native to the Americas. There are 31 living species: the anteaters, tree sloths, and armadillos. Ex ...
closely allied to the
Armadillo Armadillos (meaning "little armored ones" in Spanish) are New World placental mammals in the order Cingulata. The Chlamyphoridae and Dasypodidae are the only surviving families in the order, which is part of the superorder Xenarthra, along wi ...
es." Seeley examined some bone fragments and concluded the specimen was that of a reptilian creature, furthermore a chelonid. It also proved to be more closely related to '' Sphargis'' than any other type in the Chelonian order.


Relation to modern Leatherback sea turtles

For a long time, modern Leatherback sea turtles (''Dermochelys'') were believed to be descended directly from ''Psephophorus'', specifically the species ''P. polygonus''. However, a 1996 analysis by Wood et al. proved that most of the taxa in the two
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
were not connected, meaning ''Psephophorus'' could not be a direct ancestor of the modern leatherbacks. The
platelets Platelets, also called thrombocytes (from Greek θρόμβος, "clot" and κύτος, "cell"), are a component of blood whose function (along with the coagulation factors) is to react to bleeding from blood vessel injury by clumping, thereby ini ...
on ''Psephophorus'' are quite similar to those on ''
Dermochelys The leatherback sea turtle (''Dermochelys coriacea''), sometimes called the lute turtle or leathery turtle or simply the luth, is the largest of all living turtles and the heaviest non-crocodilian reptile, reaching lengths of up to and weights ...
'', despite differences in outer morphology and size. The platelet comprises an external compact layer and an internal zone of cancellous bone.


Species

*''Psephophorus polygonus'' is the type species, and was discovered by von Meyer in 1846.
Fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
of the species have shown bony, carapacial ridges, adjacent, small and polygonal bony ossicles which measure about 22 × 25 mm. When the ossicles extend further along the longitudinal axis, their size increases up to an average of 33 × 41 mm. *''Psephophorus calvertensis'' was first named by Palmer in 1909. It was so named for being found in the Calvert Formation. ''P. calvertensis'' is a rarely found specimen. *''Psephophorus eocaenus'' was first named by Andrews in 1901. *''Psephophorus californiensis'' was first named by Gilmore in 1937. *''Psephophorus terrypratchetti'' was discovered in the 1990s by
Richard Köhler Richard is a male given name. It originates, via Old French, from Old Frankish and is a compound of the words descending from Proto-Germanic ''*rīk-'' 'ruler, leader, king' and ''*hardu-'' 'strong, brave, hardy', and it therefore means 'stron ...
in New Zealand and named in 1995 after the author Terry Pratchett, who wrote a series of fantasy books set on a world carried on the back of a giant turtle. This species reached in body length which is more than 80% bigger than ''P. eocaenus''.


See also

* List of organisms named after famous people (born 1900–1949) * ''
Archelon ''Archelon'' is an extinct marine turtle from the Late Cretaceous, and is the largest turtle ever to have been documented, with the biggest specimen measuring from head to tail and in body mass. It is known only from the Dakota Pierre Shal ...
'' * ''
Gigantatypus ''Gigantatypus'' is an extinct late Maastrichtian sea turtle that lived in the southern regions of the Tethys Ocean about off the north eastern margins of Cretaceous Africa immediately before the Cretaceous–Paleogene extinction events . F ...
'' * Largest prehistoric animals


References


Further reading

* * * *


External links

* * * * * * {{Taxonbar, from=Q7254102 Dermochelyidae Fossil taxa described in 1995 Terry Pratchett Oligocene turtles Miocene turtles Pliocene turtles Cenozoic reptiles of Europe Cenozoic reptiles of North America Oligocene genus first appearances Pliocene genus extinctions Prehistoric turtle genera Taxa named by Christian Erich Hermann von Meyer Extinct turtles